bioRxiv preprint doi: https://doi.org/10.1101/059311; this version posted June 16, 2016. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY-NC-ND 4.0 International license. 1 The genetic structure of the world’s first farmers 2 3 Iosif Lazaridis1,2,†, Dani Nadel3, Gary Rollefson4, Deborah C. Merrett5, Nadin Rohland1, 4 Swapan Mallick1,2,6, Daniel Fernandes7,8, Mario Novak7,9, Beatriz Gamarra7, Kendra Sirak7,10, 5 Sarah Connell7, Kristin Stewardson1,6, Eadaoin Harney1,6,11, Qiaomei Fu1,12,13, Gloria 6 Gonzalez-Fortes14, Songül Alpaslan Roodenberg15, György Lengyel16, Fanny Bocquentin17, 7 Boris Gasparian18, Janet M. Monge19, Michael Gregg19, Vered Eshed20, Ahuva-Sivan 8 Mizrahi20, Christopher Meiklejohn21, Fokke Gerritsen22, Luminita Bejenaru23, Matthias 9 Blüher24, Archie Campbell25, Gianpiero Cavalleri26, David Comas27, Philippe Froguel28,29, 10 Edmund Gilbert26, Shona M. Kerr25, Peter Kovacs30, Johannes Krause31,32,33, Darren 11 McGettigan34, Michael Merrigan35, D. Andrew Merriwether36, Seamus O'Reilly35, Martin B. 12 Richards37, Ornella Semino38, Michel Shamoon-Pour36, Gheorghe Stefanescu39, Michael 13 Stumvoll24, Anke Tönjes24, Antonio Torroni38, James F. Wilson40,41, Loic Yengo28,29, Nelli A. 14 Hovhannisyan42, Nick Patterson2, Ron Pinhasi7,*,† and David Reich1,2,6,*,† 15 16 *Co-senior authors; 17 †Correspondence and requests for materials should be addressed to: 18 I. L. (
[email protected]), R. P. (
[email protected]), or D. R. 19 (
[email protected]) 20 21 1 Department of Genetics, Harvard Medical School, Boston, Massachusetts 02115, USA 22 2 Broad Institute of MIT and Harvard, Cambridge, Massachusetts 02142, USA 23 3 The Zinman Institute of Archaeology, University of Haifa, Haifa 3498838, Israel 24 4 Dept.