Gambel Oak Management Fact Sheet No
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Clonality and Dynamics of Leaf Abscission of Gambel Oaks at Small Spatial Scales in Utah
For. Sci. 61(●):000–000 FUNDAMENTAL RESEARCH http://dx.doi.org/10.5849/forsci.14-204 Copyright © 2015 Society of American Foresters forest ecology Clonality and Dynamics of Leaf Abscission of Gambel Oaks at Small Spatial Scales in Utah Jacob W. Chalmers, Karen E. Mock, Kevin D. Kohl, Katelin J. Madsen, and Nalini M. Nadkarni The Intermountain West of the United States supports a mosaic of Gambel oak (Quercus gambelii Nutt.) clumps interspersed with grassland. Both the tree and shrub forms of Gambel oak are typically found in distinct clumps that have often been presumed to be single clones, although quantitative evidence to support this is minimal. We examined the patterns of clonality within and between oak clumps and explored the variance in phenology of leaf abscission within and among clones and clumps in a study site in the Wasatch Mountains, Utah, USA. Using amplified fragment length polymorphism analysis, we genotyped multiple stems within 10 clumps of Gambel oaks in Red Butte Canyon. Nine of the 10 clumps were monoclonal (each a single distinct genet). The 10th clump was composed of 2 genets. To characterize the genetic influence on leaf abscission phenology, we compared the timing of abscission of individually marked leaves among genets, ramets, and branches. Of these three levels of organization, genet membership had the greatest influence on timing of leaf abscission, and branch membership had the smallest influence. Our results suggest that clonal diversity may be an important metric of phenotypic and ecological diversity in Gambel oak landscapes. Keywords: genet, ramet, abscission, Gambel oak lant communities often comprise genetic mosaics at multiple trait variances are partitioned among genets (clones) versus ramets spatial scales because species in the communities reproduce (stems). -
The Collection of Oak Trees of Mexico and Central America in Iturraran Botanical Gardens
The Collection of Oak Trees of Mexico and Central America in Iturraran Botanical Gardens Francisco Garin Garcia Iturraran Botanical Gardens, northern Spain [email protected] Overview Iturraran Botanical Gardens occupy 25 hectares of the northern area of Spain’s Pagoeta Natural Park. They extend along the slopes of the Iturraran hill upon the former hay meadows belonging to the farmhouse of the same name, currently the Reception Centre of the Park. The minimum altitude is 130 m above sea level, and the maximum is 220 m. Within its bounds there are indigenous wooded copses of Quercus robur and other non-coniferous species. Annual precipitation ranges from 140 to 160 cm/year. The maximum temperatures can reach 30º C on some days of summer and even during periods of southern winds on isolated days from October to March; the winter minimums fall to -3º C or -5 º C, occasionally registering as low as -7º C. Frosty days are few and they do not last long. It may snow several days each year. Soils are fairly shallow, with a calcareous substratum, but acidified by the abundant rainfall. In general, the pH is neutral due to their action. Collections The first plantations date back to late 1987. There are currently approximately 5,000 different taxa, the majority being trees and shrubs. There are around 3,000 species, including around 300 species from the genus Quercus; 100 of them are from Mexico and Central America. Quercus costaricensis photo©Francisco Garcia 48 International Oak Journal No. 22 Spring 2011 Oaks from Mexico and Oaks from Mexico -
F.3 References for Appendix F
F-1 APPENDIX F: ECOREGIONS OF THE 11 WESTERN STATES AND DISTRIBUTION BY ECOREGION OF WIND ENERGY RESOURCES ON BLM-ADMINISTERED LANDS WITHIN EACH STATE F-2 F-3 APPENDIX F: ECOREGIONS OF THE 11 WESTERN STATES AND DISTRIBUTION BY ECOREGION OF WIND ENERGY RESOURCES ON BLM-ADMINISTERED LANDS WITHIN EACH STATE F.1 DESCRIPTIONS OF THE ECOREGIONS Ecoregions delineate areas that have a general similarity in their ecosystems and in the types, qualities, and quantities of their environmental resources. They are based on unique combinations of geology, physiography, vegetation, climate, soils, land use, wildlife, and hydrology (EPA 2004). Ecoregions are defined as areas having relative homogeneity in their ecological systems and their components. Factors associated with spatial differences in the quality and quantity of ecosystem components (including soils, vegetation, climate, geology, and physiography) are relatively homogeneous within an ecoregion. A number of individuals and organizations have characterized North America on the basis of ecoregions (e.g., Omernik 1987; CEC 1997; Bailey 1997). The intent of such ecoregion classifications has been to provide a spatial framework for the research, assessment, management, and monitoring of ecosystems and ecosystem components. The ecoregion discussions presented in this programmatic environmental impact statement (PEIS) follow the Level III ecoregion classification based on Omernik (1987) and refined through collaborations among U.S. Environmental Protection Agency (EPA) regional offices, state resource management agencies, and other federal agencies (EPA 2004). The following sections provide brief descriptions of each of the Level III ecoregions that have been identified for the 11 western states in which potential wind energy development may occur on BLM-administered lands. -
Ships of 8 Tree Species in Navajo National Monument, Arizona
Population Dynamics and Age Relation- ships of 8 Tree Species in Navajo National Monument, Arizona J.D. BROTHERSON, S.R. RUSHFORTH, W.E. EVENSON, J.R. JOHANSEN, AND C. MORDEN The presence of 3 major archeological ruins dating from the 1 lth cent slickrock areas and the plateau behind the canyon rim to 13th centuries (Woodbury 1963) provided the primary motiva- (Brotherson et al. 1978). The pigmy woodland community covers tion for including Navajo National Monument in the National more area than any other type in northeastern Arizona. Parks System. Also included in the monument are some unique Gambel oak is the most extensive type in the Monument aside ecosystems, especially a small relict of “mountain vegetation” from tne pinyon-juniper community. It is found inall 3 segments of found in Betatakin Canyon. As visitor pressures mount annually, the monument but reaches its greatest development at Keet Steel proper management of these unique ecosystems becomes highly and Betatakin. Pnrnus emarginata (bittercherry) also grows at important. Since trees are the dominant features of these ecosys- Keet Steel but is uncommon. tems, and are central to management considerations, the present The streamside community of the Inscription House segment of work has examined populations of 8 major tree species in the the monument contains Popufus fremontii (Fremont poplar), monument. Populus angustifolia (narrowleaf cottonwood), Salk goodingii The objectives of this study were, first, to develop age prediction (Gooding willow), Tamarix ramosissima (salt cedar), and Elaeg- equations for tree species growing in Navajo National Monument, nus angustifolia (Russian olive). Fremont poplar is the dominant and second, to assess the present age profiles, reproductive recruit- tree within this community with salt cedar attaining local impor- ment, and density relationships of these tree populations. -
Oaks for Nebraska Justin R
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Nebraska Statewide Arboretum Publications Nebraska Statewide Arboretum 2013 Oaks for Nebraska Justin R. Evertson University of Nebraska - Lincoln, [email protected] Follow this and additional works at: http://digitalcommons.unl.edu/arboretumpubs Part of the Botany Commons, and the Forest Biology Commons Evertson, Justin R., "Oaks for Nebraska" (2013). Nebraska Statewide Arboretum Publications. 1. http://digitalcommons.unl.edu/arboretumpubs/1 This Article is brought to you for free and open access by the Nebraska Statewide Arboretum at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Nebraska Statewide Arboretum Publications by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Oaks for Nebraska Justin Evertson, Nebraska Statewide Arboretum arboretum.unl.edu or retreenebraska.unl.edu R = belongs to red oak group—acorns mature over two seasons & leaves typically have pointed lobes. W = belongs to white oak group— acorns mature in one season & leaves typically have rounded lobes. Estimated size range is height x spread for trees growing in eastern Nebraska. A few places to see oaks: Indian Dwarf chinkapin oak, Quercus Cave State Park; Krumme Arboretum Blackjack oak, Quercus prinoides (W) in Falls City; Peru State College; marilandica (R) Variable habit from shrubby to Fontenelle Nature Center in Bellevue; Shorter and slower growing than tree form; prolific acorn producer; Elmwood Park in Omaha; Wayne most oaks with distinctive tri- can have nice yellow fall color; Park in Waverly; University of lobed leaves; can take on a very national champion grows near Nebraska Lincoln; Lincoln Regional natural look with age; tough and Salem Nebraska; 10-25’x 10-20’. -
Cercocarpus Montanus Cercocarpus Montanus ‘USU-CEMO-001’ ‘USU-CEMO- Was Collected As a Suspected Dwarf Plant in TM Moffat County, CO, on 20 June 2014
HORTSCIENCE 55(11):1871–1875. 2020. https://doi.org/10.21273/HORTSCI15343-20 Origin Cercocarpus montanus Cercocarpus montanus ‘USU-CEMO-001’ ‘USU-CEMO- was collected as a suspected dwarf plant in TM Moffat County, CO, on 20 June 2014. This 001’: A New Sego Supreme Plant unique, procumbent specimen was discovered laying over a rock on a windy ridge at an Asmita Paudel, Youping Sun, and Larry A. Rupp elevation of 2708 m (Fig. 2A). The appearance Center for Water Efficient Landscaping, Department of Plants, Soils, and looked different from the typical C. montanus Climate, Utah State University, 4820 Old Main Hill, Logan, UT 84322 nearby (Fig. 2B). The leaves are smaller and narrower with less serrations compared with Richard Anderson the typical plants (Fig. 2C). Healthy cuttings USU Botanical Center, 725 Sego Lily Drive, Kaysville, UT 84037 were collected, wrapped in moist newspapers and placed on ice until transferred to a cooler at Additional index words. alder-leaf mountain mahogany, cutting propagation, landscape plant, 4 °C. On 21 June, the terminal cuttings were native plant, true mountain mahogany rinsed in 1% ZeroTol (27.1% hydrogen diox- ide, 2.0% peroxyacetic acid, 70.9% inert in- gredient; BioSafe Systems, Hartford, CT), Sego SupremeTM is a plant introduction mose style with elongated achenes wounded by scraping 1 cm of bark from the program developed by the Utah State Uni- (Fig. 1B) (Shaw et al., 2004). It possesses base of the cutting on one side, treated with versity (USU) Botanical Center (Anderson an extensive root system and adapts to 2000 mg·L–1 indole-3-butyric acid (IBA)/1000 et al., 2014) to introduce native and adaptable medium to coarse textured soil. -
C P the Important Distinctions
C P The Important Distinctions S Common M Pin Cherry Black Cherry Chokecherry Canada Plum Prunus pennsylvanica Prunus serotina Prunus nigra U Prunus virginiana L P BARK D Smooth with a pungent, Young trunks: prominent N Nearly smooth. Large disagreeable odor. white lenticals. TEXTURE horizontal lenticels show Lenticels less prominent Lenticels yellowish A Older trunks: fissured orange when rubbed. than on other and ridged. Prunus species. S E Grayish-brown, with I COLOR Reddish-brown Young trunks are black Dull reddish-brown to black light-colored fissures R LEAVES R E Elliptic/oblong, widest in H Long and tapering from the center, thick leathery Obovate, widest in the C base to tip. Widest in the and shiny. Underside of terminal 1⁄3, sharply saw- Ovate or obovate tapering GENERAL lower 1⁄3; thin and firm midrib near stalk end cov- toothed and without hairs, abruptly into a long thin DESCRIP- textured with round teeth. ered with rusty, brown medium leathery in tex- point. Teeth rounded. TION Glands on stalk, and no hairs. Glands on stalk ture, glands on stalk and Glands on stalk. hairs on midribs. near blade. Margin has no brown hairs on midrib. rounded teeth. TWIGS Thorns common on SHAPE Very fine Waxy Medium slender older twigs Red-brown with a lighter Current growth gray, older COLOR Red and reddish-brown Gray or purplish-brown or greenish margin growth darkening to black Sharp, pungent smell Strong, pungent bitter- ODOR Slight cherry odor None when broken almond odor BUDS Cone shaped, slender – Football-shaped with a SHAPE Ovate, -
TREES for WESTERN NEBRASKA Justin Evertson & Bob Henrickson
THE NEBRASKA STATEWIDE ARBORETUM PRESENTS TREES FOR WESTERN NEBRASKA Justin Evertson & Bob Henrickson. For more plant information, visit plantnebraska.org or retreenbraska.unl.edu The following species are recommended for areas in the western half of Nebraska and/or typically receive less than 20” of moisture per year. Size Range: The size range indicated for each plant is the expected average mature height x spread for Nebraska. Large Deciduous Trees (typically over 40 feet tall at maturity) NOTE ON ASH SPECIES: Native American ash trees are being decimated by Emerald Ash Borer (EAB) and the insect is now in Nebraska. NSA recommends that native ash species no longer be planted in Nebraska. 1. Ash, Manchurian - Fraxinus mandshurica (from Asia; upright growth; drought tolerant; may be resistant to EAB; 40’x 30’) 2. Catalpa, Northern - Catalpa speciosa (native; tough tree; large, heart-shaped leaves, showy flowers and long seed pods; 50’x 35’) 3. Coffeetree, Kentucky - Gymnocladus dioicus (native; amazingly adaptable; beautiful winter form; 50’x 40’) 4. Cottonwood, Eastern - Populus deltoides (majestic native; not for extremely dry sites; avoid most cultivars; 80’x 60’) 5. Cottonwood, Lanceleaf - Populus acuminata (native; naturally occurring hybrid; narrow leaves; for west. G.P.; 50’x 35’) 6. Elm, American - Ulums americana (disease resistant varieties include ‘Valley Forge’ and ‘New Harmony’; 50’x50’) 7. Elm, Japanese - Ulmus davidiana var. japonica (cold tolerant; rounded; glossy green; ‘Discovery’ is a cultivar from Manitoba Canada; 45’x 45’) 8. Elm, Rock - Ulmus thomasii (distinctive corky stems; upright habit; DED resistance in west; 50-60’x 30-40’) New Elm, Hybrids - many disease resistant hybrid elms have been developed and show promise, including: 9. -
Chokecherry Prunus Virginiana L
chokecherry Prunus virginiana L. Synonyms: Prunus virginiana ssp. demissa (Nutt.) Roy L. Taylor & MacBryde, P. demissa (Nutt.) Walp. Other common names: black chokecherry, bitter-berry, cabinet cherry, California chokecherry, caupulin, chuckleyplum, common chokecherry, eastern chokecherry, jamcherry, red chokecherry, rum chokecherry, sloe tree, Virginia chokecherry, western chokecherry, whiskey chokecherry, wild blackcherry, wild cherry Family: Rosaceae Invasiveness Rank: 74 The invasiveness rank is calculated based on a species’ ecological impacts, biological attributes, distribution, and response to control measures. The ranks are scaled from 0 to 100, with 0 representing a plant that poses no threat to native ecosystems and 100 representing a plant that poses a major threat to native ecosystems. Description Similar species: Several non-native Prunus species can Chokecherry is a deciduous, thicket-forming, erect shrub be confused with chokecherry in Alaska. Unlike or small tree that grows 1 to 6 m tall from an extensive chokecherry, which has glabrous inner surfaces of the network of lateral roots. Roots can extend more than 10 basal sections of the flowers, European bird cherry m horizontally and 2 m vertically. Young twigs are often (Prunus padus) has hairy inner surfaces of the basal hairy. Stems are numerous, slender, and branched. Bark sections of the flowers. Chokecherry can also be is smooth to fine-scaly and red-brown to grey-brown. differentiated from European bird cherry by its foliage, Leaves are alternate, elliptic to ovate, and 3 to 10 cm which turns red in late summer and fall; the leaves of long with pointed tips and toothed margins. Upper European bird cherry remain green throughout the surfaces are green and glabrous, and lower surfaces are summer. -
25. Utah Serviceberry-Saskatoon Serviceberry Ecological Series
G. NON-RIPARIAN TALL SHRUBLANDS (SA) 25. Utah Serviceberry-Saskatoon Serviceberry Ecological Series Table 25-1. Full and short names for the ecological types in the Utah-Saskatoon Serviceberry Ecological Series. Ecological Type Plant Association Code Name Code Short Name Utah serviceberry/dryland sedge-sun sedge–Deep Utah serviceberry/sedge– SA1 Argiborolls–Leeward upper backslopes and shoulders, AMUT/CAGE-CAPEH Dark clay soils–Leeward < 9,100 ft Utah serviceberry-mountain-mahogany/dryland sedge-sun Utah serviceberry-mountain- AMUT-CEMO2/ SA2 sedge–Argiborolls–Protected colluvial backslopes and mahogany/sedge–Dark clay CAGE-CAPEH shoulders, < 8,700 ft soils–Protected Serviceberry-Gambel oak/Sedge–Deep Argiborolls, little AMELA-QUGA/ Serviceberry-oak–Dark clay SA3 coarse on surface–Lees or other protected slopes, CAREX soils–Protected 7,600 - 8,600 ft Serviceberry/green needlegrass-spike-fescue–Deep Serviceberry/green SA4 Argiborolls, sometimes Pachic–Lee sides of ridges and AMELA/NAVI4-LEKI2 needlegrass–Deep clay soils– mesas, 8,000-9,300 ft Lees Saskatoon serviceberry/elk sedge–Deep Argiborolls and Argic Saskatoon serviceberry/elk SA5 Cryoborolls, often Pachic–Lee slopes on ridges and mesas, AMAL2/CAGE2 sedge–Deep dark soils–Lees 8,500-9,900 ft Saskatoon serviceberry/Thurber fescue–Deep Argic Serviceberry/Thurber fescue– SA6 Cryoborolls, loam surface–subalpine slopes, AMAL2/FETH Deep dark cold loamy soils– 8,700-10,000 ft Subalpine The Amelanchier utahensis-Amelanchier includes part of the Mountain Mahogany-Mixed alnifolia series is described as new here, based on Shrub series of Dick-Peddie (1993), but it is not the the Amelanchier alnifolia series of Komárková same as the eastern slope Cercocarpus montanus (1986-1988) and on the Quercus gambelii series of series of Hess (1981). -
Native Shrubs
Native Shrubs Downy Serviceberry (Amelanchier arborea) Grows from 10 to 25 feet high with a spread of 12 feet. A deciduous tree that prefers rich loamy soil but will grow well in clay or any soil that has moderate moisture. White showy flowers bloom in early to mid spring and are followed by dark red to purple edible berries. Zones 4 - 9. Shadblow Serviceberry (Amelanchier canadensis) Grows from 25 to 30 feet high with a spread of 15 to 20 feet. Grows best in medium wet, well-drained soil but will tolerate a wide range. Prefers partial shade to full sun. Clusters of white flowers are followed by edible red/purple berries in late summer. Zones 4-8 Allegheny Serviceberry (Amelancheir laevis) Grows to approximately 25 feet high with a spread of 20 feet. Grows in shade and partial shade and prefers moist soils. A hardy serviceberry species that will tolerate more moisture and light then some other varieties. White flowers and purple/black edible berries are typical. Zones 4-8. Bog Rosemary (Andromeda polifolia) Grows from 6 to 30 inches high with a spread of 3 feet. Leaves are narrow, evergreen and leathery with a blue-green color. Some resemblance to the culinary herb. Typically found in northern bogs and marshes. Flowers are small, pink, and bell- shaped. Grows best in very moist, acidic soil in cooler climates. Zones 2- 6. Black Chokeberry (Aronia melancarpa) Can grow up to 8 feet high with a spread of 8 feet. Grows best in moist, well-drained, acidic soils but will tolerate drier sandy soils or wet clayey ones. -
Bird Communities of Gambel Oak: a Descriptive Analysis
United States Department of Agriculture Bird Communities Forest Service Rocky Mountain of Gambel Oak: A Research Station General Technical Report RMRS-GTR-48 Descriptive Analysis March 2000 Andreas Leidolf Michael L. Wolfe Rosemary L. Pendleton Abstract Leidolf, Andreas; Wolfe, Michael L.; Pendleton, Rosemary L. 2000. Bird communities of gambel oak: a descriptive analysis. Gen. Tech. Rep. RMRS-GTR-48. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 30 p. Gambel oak (Quercus gambelii Nutt.) covers 3.75 million hectares (9.3 million acres) of the western United States. This report synthesizes current knowledge on the composition, structure, and habitat relationships of gambel oak avian communities. It lists life history attributes of 183 bird species documented from gambel oak habitats of the western United States. Structural habitat attributes important to bird-habitat relationships are identified, based on 12 independent studies. This report also highlights species of special concern, provides recommendations for monitoring, and gives suggestions for management and future research. Keywords: Avian ecology, bird-habitat relationships, neotropical migrant, oakbrush, oak woodlands, scrub oak, Quercus gambelii, Western United States The Authors ______________________________________ Andreas Leidolf is a Graduate Research Assistant in the Department of Fisheries and Wildlife at Utah State University (USU). He received a B.S. degree in Forestry/Wildlife Management from Mississippi State University in 1995. He is currently completing his M.S. degree in Fisheries and Wildlife ecology at USU. Michael L. Wolfe is a Professor in the Department of Fisheries and Wildlife at USU. He received a B.S. degree in Wildlife Management at Cornell University in 1963 and his doctorate in Forestry/Wildlife Management at the University of Göttingen, Germany, in 1967.