stone age institute publication series Series Editors Kathy Schick and Nicholas Toth

Stone Age Institute Gosport, Indiana and Indiana University, Bloomington, Indiana

Number 1. THE : Case Studies into the Earliest Stone Age Nicholas Toth and Kathy Schick, editors

Number 2. BREATHING LIFE INTO FOSSILS: Taphonomic Studies in Honor of C.K. (Bob) Brain Travis Rayne Pickering, Kathy Schick, and Nicholas Toth, editors

Number 3. THE CUTTING EDGE: New Approaches to the of Human Origins Kathy Schick, and Nicholas Toth, editors

Number 4. THE HUMAN BRAIN EVOLVING: Paleoneurological Studies in Honor of Ralph L. Holloway Douglas Broadfield, Michael Yuan, Kathy Schick and Nicholas Toth, editors STONE AGE INSTITUTE PUBLICATION SERIES NUMBER 2 Series Editors Kathy Schick and Nicholas Toth

breathing life into fossils: Taphonomic Studies in Honor of C.K. (Bob) Brain

Editors

Travis Rayne Pickering University of Wisconsin, Madison

Kathy Schick Indiana University

Nicholas Toth Indiana University

Stone Age Institute Press · www.stoneageinstitute.org 1392 W. Dittemore Road · Gosport, IN 47433 COVER CAPTIONS AND CREDITS.

Front cover, clockwise from top left.

Top left: Artist’s reconstruction of the depositional context of Swartkrans Cave, , with a leopard consuming a hominid carcass in a tree outside the cave: bones would subsequently wash into the cave and be incorporated in the breccia deposits. © 1985 Jay H. Matternes.

Top right: The Swartkrans cave deposits in South Africa, where excavations have yielded many hominids and other animal fossils. ©1985 David L. Brill.

Bottom right: Reconstruction of a hominid being carried by a leopard. © 1985 Jay H. Matternes.

Bottom left: Photograph of a leopard mandible and the skull cap of a hominid from Swartkrans, with the leopard’s canines juxtaposed with puncture marks likely produced by a leopard carrying its hominid prey. © 1985 David L. Brill.

Center: Photo of Bob Brain holding a cast of a spotted hyena skull signed by all of the taphonomy conference participants. © 2004 Kathy Schick, Stone Age Institute.

Back cover credits.

Top: © 2004 Stone Age Institute.

Bottom left: © 2004 Kathy Schick, Stone Age Institute.

Bottom right: © 2005 Greg Murphy.

Published by the Stone Age Institute. ISBN-10: 0-9792-2761-5 ISBN-13: 978-0-9792-2761-5 Copyright © 2007, Stone Age Institute Press.

All rights reserved under International and Pan-American Copyright Conventions. No part of this book may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopying, without permission in writing from the publisher. Contents ix

BREATHING LIFE INTO FOSSILS: TAPHONOMIC STUDIES IN HONOR OF C.K. (BOB) BRAIN EDITED BY TRAVIS RAYNE PICKERING, NICHOLAS TOTH AND KATHY SCHICK STONE AGE INSTITUTE PRESS, 2007

CONTENTS

Comment by Series Editors ...... ii Contributors ...... v Acknowledgements ...... vii Table of Contents ...... ix Preface ...... xi Foreword ...... xi Introduction ...... xv

Chapter 1: Fifty Years of Fun with Fossils: Some Cave Taphonomy-Related Ideas and Concepts that Emerged Between 1953 and 2003 C.K. Brain ...... 1 Chapter 2: Rather Odd Detective Stories: A View of Some Actualistic and Taphonomic Trends in Paleoindian Studies Gary Haynes ...... 25 Chapter 3: The Destruction of Skeletal Elements by Carnivores: The Growth of a General Model for Skeletal Element Destruction and Survival in Zooarchaeological Assemblages Naomi Cleghorn and Curtis W. Marean ...... 37 Chapter 4: Hominids and Carnivores at Kromdraai and Other Quaternary Sites in Southern Africa J. Francis Thackeray ...... 67 Chapter 5: Taphonomic Analysis of an Excavated Striped Hyena Den from the Eastern Desert of Jordan Kathy Schick, Nicholas Toth, Thomas Gehling and Travis Rayne Pickering ...... 75 Chapter 6: Taphonomy and Site Formation of Two Early Miocene Sites on Rusinga Island, Alan Walker ...... 107 Chapter 7: Are You In or Out (of Africa)? Site Formation at Dmanisi and Actualistic Studies in Africa Martha Tappen, David Lordkipanidze, Maia Bukshianidze, Reid Ferring and Abesalom Vekua ...... 119 x Breathing Life into Fossils: Taphonomic Studies in Honor of C.K. (Bob) Brain

Chapter 8: Changes Through Time in Carcass Survival in the Amboseli Ecosystem, Southern Kenya A. K. Behrensmeyer ...... 137 Chapter 9: Vertebrate Taphonomic Perspectives on Oldowan Hominin Land Use in the Plio-Pleistocene Olduvai Basin, Robert J. Blumenschine, Charles R. Peters, Salvatore D. Capaldo, Peter Andrews, Jackson K. Njau and Briana L. Pobiner ...... 161 Chapter 10: The Earlier Stone Age in South Africa: Site Context and the Infl uence of Cave Studies Kathleen Kuman ...... 181 Chapter 11: Taphonomy of Sterkfontein Australopithecus Skeletons Ron J. Clarke ...... 199 Chapter 12: Taphonomy of Immature Hominid Skulls and the Taung, Mojokerto, and Herto Specimens Gail E. Krovitz and Pat Shipman...... 207 Chapter 13: Carcass Foraging by Early Hominids at Swartkrans Cave (South Africa): A New Investigation of the Zooarchaeology and Taphonomy of Member 3 Travis Rayne Pickering, Manuel Domínguez-Rodrigo, Charles P. Egeland and C.K. Brain ...... 233 Chapter 14: Equifi nality in Carnivore Tooth Marks and the Extended Concept of Archaeological Palimpsests: Implications for Models of Passive Scavenging by Early Hominids Manuel Domínguez-Rodrigo, Charles P. Egeland and Travis Rayne Pickering ...... 255 Chapter 15: Butchering Backstraps and Bearing Backbones: Insights from Hadza Foragers and Implications for Archaeology Henry T. Bunn ...... 269 Chapter 16: Carnivora and Carnivory: Assessing Hominid Toothmarks in Zooarchaeology Tim D. White and Nicholas Toth ...... 281