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A Monographic Revision of the Genus Platycoelia Dejean (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini) Andrew B

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A Monographic Revision of the Genus Platycoelia Dejean (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini) Andrew B University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Bulletin of the University of Nebraska State Museum, University of Nebraska State Museum 2003 A Monographic Revision of the Genus Platycoelia Dejean (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini) Andrew B. T. Smith University of Nebraska - Lincoln, [email protected] Follow this and additional works at: http://digitalcommons.unl.edu/museumbulletin Part of the Entomology Commons, and the Other Ecology and Evolutionary Biology Commons Smith, Andrew B. T., "A Monographic Revision of the Genus Platycoelia Dejean (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini)" (2003). Bulletin of the University of Nebraska State Museum. 3. http://digitalcommons.unl.edu/museumbulletin/3 This Article is brought to you for free and open access by the Museum, University of Nebraska State at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Bulletin of the University of Nebraska State Museum by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. A Monographic Revision of the Genus Platycoelia Dejean (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini) Andrew B. T. Smith Bulletin of the University of Nebraska State Museum Volume 15 A Monographic Revision of the Genus Platycoelia Dejean (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini) by Andrew B. T. Smith UNIVERSITY. OF, ( NEBRASKA "-" STATE MUSEUM Published by the University of Nebraska State Museum Lincoln, Nebraska 2003 Bulletin of the University of Nebraska State Museum Volume 15 Issue Date: 7 July 2003 Editor: Brett C. Ratcliffe Cover Design: Angie Fox Text design and layout: Linda J. Ratcliffe Text fonts: New Century Schoolbook and Arial Bulletins may be purchased from the Museum. Address orders to: Publications Secretary W436 Nebraska Hall University of Nebraska State Museum Lincoln, NE 68588-0514 U.S.A. Price: $20.00 Copyright © by the University of Nebraska State Museum, 2003. All rights reserved. Apart from citations for the purposes of research or review, no part of this Bulletin may be reproduced in any form, mechanical or electronic, including photocopying and recording, without permission in writing from the publisher. ISSN 0093-6812 Library of Congress Catalog Card Number Printed in the United States of America The Bulletin is a peer-reviewed journal. 11 CONTENTS Abstract .............................................................................................................. 1 Introduction ........................................................................................................ 3 Tribe Anoplognathini ....................................................................................... .4 Subtribe Platycoeliina...................................................................................... 6 Taxonomic History of the genus Platycoelia ...................................................... 6 Notes on Authors, Types, and Collections .......................................................... 8 Phylogenetic Methods and Materials ............................................................... 12 Phylogenetic Analysis ................................................................................ 12 Character Definition .................................................................................. 12 Adult Morphological Character Analysis .................................................. 13 Phylogenetic ProtocoL ................................................................................ 22 Taxonomic Methods and Materials .................................................................. 22 Specimens ................................................................................................. 22 Catalog Format ............................................................................................ 24 Type Specimens ........................................................................................... 24 Designation of Lectotypes and Neotypes .................................................... 25 Species Concept.......................................................................................... 26 Phylogenetic Patterns and Biogeographic Implications ................................... 27 Key to the Neotropical Genera of Anoplognathini ........................................... 30 Subtribe Platycoeliina Burmeister, 1844 ........................................................... 31 Genus Platycoelia Dejean, 1833 ......................................................................... 31 Key to the Species of Platycoelia ... ..................................................................... 34 1. Platycoelia alternans Erichson, 1847 ................................................. .42 2. Platycoelia inflata Ohaus, 1904 ......................................................... .44 3. Platycoelia nervosa Kirsch, 1871.. ..................................................... .48 4. Platycoelia marginata Burmeister, 1844 ............................................ 50 5. Platycoelia meridensis new species ..................................................... 55 6. Platycoelia selanderi Martinez and Martinez, 1994 ........................... 57 7. Platycoelia prasina Erichson, 184 7 ...................................................... 61 8. Platycoelia simplicior Ohaus, 1909 ..................................................... 62 9. Platycoelia abdominalis Ohaus, 1904 ................................................. 64 10. Platycoelia pomacea Erichson, 1847 ................................................... 66 11. Platycoelia occidentalis Ohaus, 1904.................................................. 69 12. Platycoelia valida Burmeister, 1844 .................................................... 71 13. Platycoelia mesosternalis Ohaus, 1904 ............................................... 74 14. Platycoelia peruviana new species ...................................................... 76 15. Platycoelia forcipalis Ohaus, 1904 ...................................................... 80 16. Platycoelia sandia new species ........................................................... 84 17. Platycoelia penai Frey, 1967 ............................................................... 86 18. Platycoelia galerana new species ........................................................ 87 19. Platycoelia flavostriata (Latreille, 1813) ............................................. 91 20. Platycoelia nigrosternalis Ohaus, 1904 .............................................. 95 21. Platycoelia humeralis Bates, 1888 ..................................................... 97 22. Platycoelia intermedia Ohaus, 1925 .................................................... 101 111 23. Platycoelia convexa new species ....................................................... 102 24. Platycoelia interstincta new species ................................................. 104 25. Platycoelia variolosa Ohaus, 1904...................................................... 105 26. Platycoelia grandicula new species ................................................... 106 27. Platycoelia hiporum new species ....................................................... 111 28. Platycoelia paucarae new species ...................................................... 112 29. Platycoelia flavoscutellata Ohaus, 1904............................................ 115 30. Platycoelia puncticollis Ohaus, 1904................................................. 116 31. Platycoelia unguicularis Ohaus, 1904............................................... 120 32. Platycoelia burmeisteriana Ohaus, 1917 ........................................... 122 33. Platycoelia burmeisteri Arrow, 1899 ................................................. 124 34. Platycoelia traceyae new species ....................................................... 126 35. Platycoelia wallisi Ohaus, 1904 ........................................................ 129 36. Platycoelia flavohumeralis new species ............................................. 131 37. Platycoelia steinheili Ohaus, 1904.................................................... 132 38. Platycoelia chrysotina Ohaus, 1904.................................................. 133 39. Platycoelia butleri new species .......................................................... 136 40. Platycoelia hirta Ohaus, 1904........................................................... 138 41. Platycoelia altiplana new species ...................................................... 140 42. Platycoelia confluens Ohaus, 1904 .................................................... 141 43. Platycoelia lutescens Blanchard, 1851.. ............................................. 144 44. Platycoelia gaujoni Ohaus, 1904....................................................... 149 45. Platycoelia pusilla new species ......................................................... 152 46. Platycoelia bordoni Martinez, 1976 ................................................... 153 47. Platycoelia parva Kirsch, 1885 .......................................................... 154 48. Platycoelia furva new species ............................................................ 157 49. Platycoelia quadrilineata Burmeister, 1844 ..................................... 159 50. Platycoelia rufosignata Ohaus, 1904................................................ 160 51. Platycoelia insolita new species ......................................................... 163 52. Platycoelia signaticollis
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