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Die Echinodermen Der Rügener Schreibkreide (Maastrichtium, Deutschland) – Ein Kurzes Update Der Nachgewiesenen Arten

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Die Echinodermen Der Rügener Schreibkreide (Maastrichtium, Deutschland) – Ein Kurzes Update Der Nachgewiesenen Arten 33 Zitteliana 92 Short Communication Die Echinodermen der Rügener Schreibkreide Paläontologie Bayerische (Maastrichtium,GeoBio- Deutschland) – ein kurzes Update & Geobiologie Staatssammlung Center LMU München für Paläontologie und Geologie LMUder München nachgewiesenen Arten München, 31.03.2018 Mike Reich1,2,3*& Manfred Kutscher4 Manuscript received 1SNSB - Bayerische Staatssammlung für Paläontologie und Geologie, Richard-Wagner-Str. 10, 01.03.2018; revision 80333 München accepted 21.03.2018 2Department für Geo- und Umweltwissenschaften, Ludwig-Maximilians-Universität München, ISSN 0373-9627 Richard-Wagner-Str. 10, 80333 München 3GeoBio-CenterLMU, Richard-Wagner-Str. 10, 80333 München ISBN 978-3-946705-04-8 4Dorfstr. 10, 18546 Sassnitz *Corresponding author; E-mail: [email protected] Zitteliana 92, 33–36. Schlüsselwörter: Echinodermata; Crinoidea; Asteroidea; Ophiuroidea; Echinoidea; Holothuroidea; Schreibkreide; Ober-Kreide; Maastrichtium; Rügen; Jasmund; Vorpommern; Deutschland Key words: Echinodermata; Crinoidea; Asteroidea; Ophiuroidea; Echinoidea; Holothuroidea; chalk; Upper Cretaceous; Maastrichtian; Rügen; Jasmund; Western Pomerania; Germany 1. Einführung Reich & Stegemann (2012), Schlüter et al. (2012, 2016), Thuy (2013), Gale (2016), Gale et al. (2018), Die Rügener Schreibkreide (Maastrichtium) ist Miller et al. (2017), Neumann & Girod (2018) und eine klassische und wichtige Typuslokalität für O’Hara et al. (2018) sowie Kutscher (unveröff.) und Fossilien der europäischen Ober-Kreide (Reich & Reich (unveröff.) herangezogen. Frenzel 2002; Reich et al. 2004a). Mit mehr als 1.470 Arten fossiler Pflanzen und Tiere (Reich et al. 2018) gehört diese heute zu den am besten bekannten 2. crinoidea (Seelilien und haarsterne) Schreibkreide-Vorkommen der Welt. Innerhalb der bisher untersuchten Fossilassoziationen befinden Isocrinida sich auch Vertreter aller heute noch vorkommenden Stachelhäuter-Gruppen – den Crinoidea, Asteroidea, Austinocrinus bicoronatus (von Hagenow, 1840) Ophiuroidea, Echinoidea und Holothuroidea – mit ‘Isocrinus’ lanceolatus (Roemer, 1840) insgesamt 157 nachgewiesenen Taxa. Damit ist die Isselicrinus buchii (Roemer, 1840) Rügener Echinodermenvergesellschaftung diver- Isselicrinus stelliferus (von Hagenow, 1840) ser als jede andere bisher untersuchte Echinoder- Nielsenicrinus agassizi (von Hagenow, 1840) menfauna vergleichbaren Alters. Dabei konnten im Biotop ‘Schreibkreide’ Vertreter unterschiedlichster comatulida ökologischer Gilden, wie beispielsweise aktive Räu- ber, Sediment- und Suspensionsfresser, verschie- Amphorometra gr. conoidea (Goldfuss, 1839) dene infaunale (aber auch pelagische) Arten und [jüngeres Synonym = Amphorometra rugiana (Gislén, 1924)] wahrscheinliche nektobenthische Schwimmer nach- Bourgueticrinus bruennichinielseni Ødum in Jes- gewiesen werden (Reich et al. 2004b). sen & Ødum, 1923 Vorliegende Kurzübersicht aktualisiert eine im Bourgueticrinus constrictus (von Hagenow in Jahre 2004 veröffentlichte englischsprachige Über- Quenstedt, 1876) sicht (Reich et al. 2004b). Dazu wurden unter ande- Bourgueticrinus hagenowi (Goldfuss, 1840) rem die seitdem veröffentlichten Arbeiten von Kut- Democrinus gisleni Rasmussen, 1961 scher (2003, 2011), Villier et al. (2004), Kutscher & Hertha gr. mystica von Hagenow, 1840 Säuberlich (2009), Kroh & Smith (2010), Reich (2012), Hertha gr. plana (Brünnich Nielsen, 1913) Zitteliana 92 34 Hertha gr. pygmaea Gislén, 1924 Forcipulatida Jaekelometra gr. belgica (Jaekel, 1902) Monachocrinus? sp. gen. et sp. indet. (Asteriidae) Placometra gr. laticirra (Carpenter, 1880) [jüngeres Synonym = Placometra scutata (Gislén, 1925)] inc. ord. Semiometra pommerania Gislén, 1924 Tricalclamnella cretacea Reich, 1997 [?juvenile Sta- Millericrinida cheln] Apiocrinites? sp. 4. ophiuroidea (Schlangensterne) Roveacrinida Euryalida Applinocrinus cretaceus (Bather, 1924) Applinocrinus ramaraoi Jain & Mallikarjuna, 1996 Asteronyx? simplex Müller, 1950 Applinocrinus texanus Peck, 1973 Asteronyx? spinulosa Kutscher & Jagt in Jagt, Birgelenocrinus degraafi Jagt, 1999 2000 gen. et sp. indet. (Roveacrinidae) Trichaster? ornatus (Rasmussen, 1950) Trichaster? sp. 3. Asteroidea (Seesterne) ophiacanthida Paxillosida Manfredura curvata (Kutscher & Jagt in Jagt, 2000) Coulonia sp. Ophiacantha? sp. Lophidiaster pygmaeus Spencer, 1913 Ophiochondrus punctatus (Kutscher & Jagt in Jagt, 2000) notomyotida Ophiochondrus semirotundus (Kutscher & Jagt in Jagt, 2000) gen. et sp. indet. (Benthopectinidae) Ophiocoma? ishidai Kutscher & Jagt in Jagt, 2000 Ophiocoma? rasmusseni Hess, 1960 Valvatida Ophiocoma? senonensis (Valette, 1915) Ophioderma? radiatum Kutscher & Jagt in Jagt, Chomataster acules Spencer, 1913 2000 Crateraster sp. Ophioderma? substriatum (Rasmussen, 1950) Metopaster poulsenii Brünnich Nielsen, 1943 Ophiogaleus danicus (Rasmussen, 1952) Metopaster tumidus Spencer, 1913 Ophiojagtus alternatus (Kutscher & Jagt in Jagt, Nymphaster wrighti (Rasmussen, 1950) 2000) Ophryaster magnus Spencer, 1913 Ophiomyxa? jekerica (Berry, 1938) Parametopaster sp. Ophiomyxa? rhipidata Kutscher & Jagt in Jagt, Pycinaster rasmusseni Müller, 1953 2000 Recurvaster gibber Schulz & Weitschat, 1971 Ophiomyxa? densa Kutscher, 2011 Stauranderaster miliaris Brünnich Nielsen, 1943 Ophiomyxa? sp. nov. Stauranderaster mixtus Brünnich Nielsen, 1943 Ophiotreta striata (Kutscher & Jagt in Jagt, 2000) Teichaster reticulatus Schulz & Weitschat, 1981 Valettaster sp. Amphilepidida Weitschataster undulatus (Spencer, 1913) gen. et sp. nov. (Stauranderasteridae) Amphiura? plana Kutscher & Jagt in Jagt, 2000 Hemieuryale? parva Kutscher & Jagt in Jagt, 2000 Velatida Ophiactis? sulcata Kutscher & Jagt in Jagt, 2000 Ophiolepis? granulata Kutscher & Jagt in Jagt, Pteraster sp. 2000 Ophiolepis? linea Kutscher & Jagt in Jagt, 2000 Spinulosida Ophiothrix? bongaertsi Kutscher & Jagt in Jagt, 2000 Echinaster? sp. Ophiothrix? cristata Kutscher & Jagt in Jagt, 2000 gen. et sp. indet. (Echinasteridae) Sigsbeia sp. 35 Zitteliana 92 ophioscolecida Phymosomatida Ophiologimus rugosus (Kutscher & Jagt in Jagt, Gauthieria alterna (Kutscher, 1985) 2000) Gauthieria princeps (von Hagenow, 1840) [jüngere Ophioscolex? clivulus Kutscher & Jagt in Jagt, Synonyme = Gauthieria radiata (Sorignet, 1850), Gauthieria pseu- 2000 doradiata (Schlüter, 1883)] Ophioscolex? cretaceus Kutscher & Jagt in Jagt, Phymosoma granulosum (Goldfuss, 1826) 2000 Phymosoma koenigi (Mantell, 1822) Phymosoma ravni Schlüter, Kutscher, Smith, Jagt ophiurida & Lees, 2012 gen. inc. sp. 1 Kutscher, 1985 [„phymosomatoide Ophiomusium biconcavum Kutscher & Jagt in Kleinformen“] Jagt, 2000 gen. inc. sp. 2 Kutscher, 1985 [„phymosomatoide Ophiomusium granulosum (Roemer, 1840) Kleinformen“] Ophiomusium sentum Kutscher & Jagt in Jagt, gen. inc. sp. 3 Kutscher, 1985 [„phymosomatoide 2000 Kleinformen“] Ophiomusium sinuatum Kutscher & Jagt in Jagt, gen. inc. sp. 3a Kutscher, 1985 [„phymosomatoide 2000 Kleinformen“] Ophiotitanos serrata (Roemer, 1840) gen. inc. sp. 4 Kutscher, 1985 [„phymosomatoide Stegophiura? hagenowi (Rasmussen, 1950) Kleinformen“] gen. inc. sp. 5 Kutscher, 1985 [„phymosomatoide ophioleucida? Kleinformen“] Sinosura jasmundensis Kutscher & Jagt in Jagt, Stomopneustoida 2000 Sinosura aff. schneideri Kutscher, 1987 Trochalosoma taeniatum (von Hagenow, 1840) Sinosura sp. Echinoneoida 5. Echinoidea (Seeigel) Conulus magnificus (d’Orbigny, 1853) Galerites (Galerites) stadensis (Lambert, 1911) cidaroida Galerites (Galerites) vulgaris (Leske, 1778) Stereocidaris hagenowi (Desor, 1858) Holectypoida Stereocidaris pistillum (Quenstedt, 1852) Temnocidaris baylei Cotteau, 1863 Echinogalerus tenuiporus (Schlüter, 1902) Tylocidaris? sp. Echinogalerus hemisphaericus (Desor, 1842) Echinothurioida holasteroida Hygrosoma bruennichi (Ravn, 1928) Cardiaster granulosus (Goldfuss, 1826) Echinocorys scutatus Leske, 1778 [Synonyme = camarodonta Echinocorys ovatus Leske, 1778, Echinocorys perconicus (von Hagenow, 1840), Echinocorys jaekeli Nietsch, 1921] Hemidiadema wehrlii (Nestler, 1978) Hagenowia elongata (Brünnich Nielsen, 1942) Offaster rugius Reich, Villier & Kutscher, 2004 Diadematoida Plesiocorys (Sternotaxis) heberti (Cotteau in Cot- teau & Triger 1860) Palaeodiadema multiforme Ravn, 1928 Spatangoida Salenioida Cyclaster (Brissopneustes) ruegensis (Kutscher, Salenia (Pleurosalenia) bonissenti (Cotteau, 1866) 1978) sensu Lambert, 1898 Cyclaster platornatus Kutscher, 1978 Salenia (Salenia) anthophora (Müller, 1847) Diplodetus sp. Salenia (Salenia) belgica Lambert, 1898 Peroniaster cotteaui Gauthier, 1887 Salenidia pygmaea (von Hagenow, 1840) Zitteliana 92 36 6. holothuroidea (Seegurken) Gulf Coast (Texas, Mississippi) and southern Sweden. Papers in Palaeontology 2(4), 489–532. Apodida Gale AS, Sadorf E, Jagt JWM. 2018. Roveacrinida (Crinoidea, Articulata) from the upper Maastrichtian Peedee Formation (upper Cretaceous) of North Carolina, USA – The last pelagic Calcancora pomerania Reich, 2003 microcrinoids. Cretaceous Research 85, 176–192. Hemisphaeranthos simplex Müller, 1964 Kroh A, Smith AB. 2010. The phylogeny and classification of post- Jumaraina rara (Müller, 1964) Palaeozoic echinoids. Journal of Systematic Palaeontology Jumaraina venusta (Müller, 1964) 8(2), 147–212. Kutscher M. 2003. Bestimmungsschlüssel der Seeigel (Echinoidea) Rigaudites nudus Reich, 2003 der Weißen Schreibkreide (Kreide, Unter-Maastrichtium) von Trematrochus smirnovi Reich, 2002 Rügen (Deutschland) und Møn (Dänemark). Erratica. Mono- gen. et sp. nov. A graphien zur Geschiebekunde 5, 1–41. gen. et sp.
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