Entomofauna carpathica, 2020, 32(2): 47-87

CONTRIBUTION TO THE KNOWLEDGE OF LITHOSIINI AND (, ) OF CENTRAL AND NORTHERN LAOS, PART 4.

Karol BUCSEK

Slovak Academy of Sciences, Institute of Zoology, Dúbravská cesta 9, 845 06 Bratislava, Slovak republic; e-mail: [email protected]

BUCSEK, K. 2020. Contribution to the knowledge of Lithosiini and Arctiini (Erebidae, Arctiinae) of central and northern Laos, part 4. Entomofauna carpathica, 32(2): 47-87.

Abstract: This study is based on material of Lithosiini and Arctiini (Erebidae, Arctiinae) recorded in the Nakai (about 500 m), Bolikhamsai (200–300 m) and Huaphanh (1200–1900 m) areas of central and northern Laos. This fourth part lists 54 , including descriptions of 4 new species: Teulisna varia n. sp., Microlithosia unicolora n. sp., Danielithosia dubatolovi n. sp. and Poliosia bilineata n. sp. They are listed below 3 new combinations: Teulisna fimbriata n. comb., Cernyia pseudobrevivalva n. comb. and Churinga nigripes n. comb.

Key words: Erebidae, Lithosiini, Arctiini, new species, Laos,

MATERIAL AND METHODS

The photos of adults were made with NIKON D700 and SONY DSC-WX220. Genitalia were mounted in microscopic slides using Canada balsam, morphological observations were made using and photographed by Digital USB 2,0 microscope camera (Micro Capture Pro). Subsequently the genitalia were demounted,preserved in glycerine in the test tubes and attached to the corresponding specimen. Post-shooting image processing was done using the software Adobe® Photoshop CS5. Holotypes are stored in the Slovak National Museum, Bratislava, paratypes in the collection of Karol Bucsek.

Abbreviations HT – holotype SNM – Slovak National Museum, Bratislava PT – paratype CKB – in coll. Karol Bucsek, Bratislava

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SYSTEMATIC SECTION

Genus Teulisna Walker, 1862 Type species: Teulisna plagiata Walker, 1862 ()

Teulisna karena Černý, 2009 (Fig. 1)

Teulisna karena Černý: Cerny, Pinratana 2009, of , vol. 6: 132, pl. 26: 264, gen. 3: 264f, 264v. Teulisna karena Černý: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 110, Pl. 19: 272, 272a, Mal145. Teulisna karena Černý: Kirti, Joshi & Singh, 2014, Acta zool. cracov. 57 (1-2): 14, Figs 6-11, 18 (list). Teulisna karena Černý: Kirti, Singh, 2015, Arctiid Moths of , vol. 1: 113.

Wingspan. Male 19 - 21 mm. Type locality. N Thailand. Distribution in Laos. Huaphanh. Geographical range. NE India, Laos, Thailand, Peninsular Malaysia.

Teulisna uniplaga Hampson, 1894 (Fig. 2)

Teulisna uniplaga Hampson, 1894, Moths India 2: 88. Ilema uniplaga Hampson, 1900, Cat. Lep. Phalaenae Br. Mus. 2: 133, 135, pl. 21: 16. Lithosia uniplaga Hampson: Draut, 1914: in Seitz, Macrolep. World 10:205, pl. 14h. uniplaga Hampson: Strand, 1922, Lep. Cat. 26: 587. Teulisna uniplaga Hampson: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 230, pl. 11: 17. Teulisna uniplaga Hampson: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 133 - 134, pl. 27: 267. Teulisna uniplaga Hampson: Bucsek 2014, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, Suppl. 29, Pl. 5: 62, 62a. Teulisna uniplaga Hampson: Kirti, Joshi & Singh, 2014, Acta zool. cracov. 57 (1-2): 19 (list). Teulisna uniplaga Hampson: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 197, Pl. 18: 149.

Wingspan. Male 18–19 mm. Type locality. (= Burma). Distribution in Laos. Nakai. Geographical range. , Myanmar, Laos, Thailand, Cambodia, Peninsular Malaysia, Sumatra, Borneo.

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Teulisna fimbriata (Leech, 1890) n. comb. (Figs. 3, 4, 90, 91)

Tegulata fimbriata Leech, 1890, Entomologist 23: 81. Ilema fimbriata Leech: Hampson, 1900, Cat. Lep. Phalaenae Br. Mus. 2: 136, pl. 21, f. 17. Lithosia fimbriata Leech: Seitz, 1910, Macrolepid. World 2: 69, pl. 13: d. Eilema fimbriata Leech: Strand, 1922, in Junk, Lep. Cat. 26: 555. Eilema protuberans Moore: Fang, 1982, Icon. Het. Sin. 2: 198, fig. 1434 (nec Moore). Eilema tumida Walker: Fang, 1992, in Icon. For. Ins. p. 936, fig. 3157 (nec Walker). Thysanoptyx fimbriata Leech: Fang, 2000, Fauna Sin., Ins. 19, Lep. Arctiidae: 236, fig. 158, pl. 11: 24. Thysanoptyx fimbriata Leech: Volynkin & Dubatolov, 2017, Zootaxa, 4277 (1): 112 (list).

Wingspan. Male 25–27 mm, female 28 mm. Type locality. C China. Distribution in Laos. Huaphanh. Geographical range. China (Shaanxi, Hubei, Hunan, , , ), Laos.

Teulisna protuberans (Moore, 1878) (Figs. 5, 6, 92, 93)

Tegulata protuberans Moore, Proc. Zool. Soc.1878: 23, pl. II: 6. Ilema protuberans Moore: Hampson, 1900, Cat. Lep.Phal. Br. Mus. 2: 133-134. Lithosia protuberans Moore: Draudt, 1914: in Seitz, Macrolep. World 10: 205, pl. 14 h. Teulisna protuberans Moore: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 131, pl. 26: 262a, 262b.

Wingspan. Male 23–25 mm, female 25 mm. Type locality. India – Darjiling. Distribution in Laos. Huaphanh. Geographical range. India (Sikkim), , Thailand, Laos.

Teulisna tumida (Walker, 1862) (Figs. 7, 8)

Tegulata tumida Walker, 1862, Journ. Lin. Soc. Lond. (Zool.) 6: 110. Teulisna tumida Walker: Hampson, 1893, Ill. Het. Br. Mus. 9: 13. Ilema tumida Walker: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 133, fig. 82. Lithosia tumida Walker: Draudt, 1914: in Seitz, Macrolep. World 10: 204, pl. 14 g. Eilema tumida Walker: Strand, 1922, in Junk, Lep. Cat. 26: 586. Eilema tumida Walker: Kishida, 1993, Tinea 13 (Suppl. 3): 37, pl. 40: 17. Teulisna tumida Walker: Fang, 2000, Fauna Sin., Ins. 19, Lep. Arctiidae: 230, fig. 150, pl. 9: 18. Teulisna tumida Walker: Holloway, 2001, Moths of Borneo 7: 309, pl. 2, fig. 58. Teulisna tumida Walker: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 130, pl. 26: 261a, 261b. 49 Entomofauna carpathica, 2020, 32(2): 47-87

Teulisna tumida Walker: Bucsek 2014, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 111, Pl. 5: 62, 62a. Teulisna tumida Walker: Kirti, Joshi & Singh, 2014, Acta zool. cracov. 57 (1-2): 19 (list). Teulisna protuberans Moore: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 115.

Wingspan. Male 21–24 mm, female 25–26 mm. Type locality. Borneo – Sarawak. Distribution in Laos. Huaphanh. Geographical range. NE India, China, Thailand, Laos, Cambodia, Peninsular Malaysia, Borneo, Sumatra.

Teulisna maculata Fang, 2000 (Figs. 9, 10)

Teulisna maculata Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 231, fig. 151, pl. 11: 19. Teulisna maculata Fang: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 134-135, pl. 27: 269. Teulisna maculata Fang: Kirti, Joshi & Singh, 2014, Acta zool. cracov. 57 (1-2): 18 (list).

Wingspan. Male 20–24 mm, female 23–25 mm. Type locality. China – Yunnan. Distribution in Laos. Huaphanh. Geographical range. China (Yunnan), Laos, Thailand.

Teulisna tongdzuythanhi Dubatolov & Bucsek, 2016 (Figs. 11, 12, 94)

Teulisna tongdzuythanhi Dubatolov & Bucsek, 2016, Euroasian Ent. J. 15 (3): 229, Figs 2, 3, 21.

Wingspan. Male 18 mm, female 19 mm. Type locality. C Vietnam. Distribution in Laos. Huaphanh. Geographical range. Vietnam, Laos.

Teulisna mithunoides Černý, 2009 (Figs. 13–15, 95, 96)

Teulisna mithunoides Černý, Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 135, pl. 27: 271. Teulisna mithunoides Černý: Kirti, Joshi & Singh, 2014, Acta zool. cracov. 57 (1-2): 18 (list). Teulisna mithunoides Černý: Bucsek, 2016, Entomofauna, 37 (37): 574, pl. 1: 2, pl. 2: 1. Teulisna mithunoides Černý: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 193, Pl. 18: 146. 50 Entomofauna carpathica, 2020, 32(2): 47-87

Wingspan. Male 15 mm, female 17 mm. Type locality. SW Thailand. Distribution in Laos. Nakai. Geographical range. Thailand, Laos, Cambodia, Peninsular Malaysia.

Teulisna varia Bucsek n. sp. (Figs. 16–18, 97, 98)

Holotype. ♂ LAOS, Huaphanne prov., Mt. Phu Pane, 1200-1900m, Ban Saluei v. env., 3.-13.V.2019, 20°12´N, 103°59´E, leg. K. Bucsek, SNM with a museum serial number SZ 10 957. Paratypes. 2 ♂♂, 2 ♀♀ like holotype, CKB.

Wingspan. Male 18 - 19 mm, female 20 - 22 mm. Holotype. Male 19 mm, antennae finely pectinated, head, thorax and abdomen ochreous, anal margin straw yellow. Forewing ochreous with a distinctive brown dust, patterning dark brown. Medial line intermittent, more pronounced on costa, slightly oblique towards the base. Postmedial line intermittent, more pronounced on costa, follows the outer margin. On termen a few dark dots, cilia ochreous. Hindwing light ochreous, subterminal area brown dusted, dust on apex more conspicuous. Male genitalia. Uncus long, nearly straight, with curved tip. Valva narrow, widened to the apex, apical lobe of valva markedly sclerotized, distal costal process very pronounced, on the apex with two small spikes. Juxta sclerotized, with distinct thin spike on both sides. Aedeagus with a large cornut, which is serrated on one side. A small field of cornuti, with a sclerotized, small, serrated plate inside. Female. Habitually the same as a male, but a little darker and more cinnamon in color. Female genitalia. Posterior apophysis and dorsal plate strongly sclerotized, ductus bursae very broad and sclerotized. Corpus bursae very markedly sclerotized, cylindrical in shape, completely filled with cornuti. Silmilar species: Habitually very similar species Teulisna tongdzuythanhi Dubatolov & Bucsek, but it has the forewing color more homogenous, the subterminal area of the hindwing is without brown dust. Teulisna mithunoides Černý is smaller. The main differences are in the genitals.

Genus Mithuna Moore, 1878 Type species: Mithuna quadriplaga Moore, 1878 (Bhutan)

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Mithuna strigifera Hampson, 1900 (Fig. 19)

Mithuna strigifera Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 123, fig. 66. Mithuna strigifera Hampson: Draudt, 1914: in Seitz, Macrolep. World 10: 213, pl. 14h. Mithuna strigifera Hampson: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 151, pl. 31: 304. Mithuna strigifera Hampson: Bucsek 2014, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 117, Pl. 20: 291, 291a, Mal157a. Mithuna strigifera Hampson: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 47.

Wingspan. Male 16–18 mm. Type locality. India – Sikkim. Distribution in Laos. Huaphanh. Geographical range. India (Sikkim), Thailand, Laos, Peninsular Malaysia.

Genus Macotasa Moore, 1878 Type species: Teulisna biplagella Butler, 1877 (Borneo)

Macotasa nedoshivinae Dubatolov, 2012 (Figs. 20, 21, 99)

Macotasa nedoshivinae Dubatolov, 2012, Euroasian Ent. J. 11 (6): 511, pl. 3: 7-8, f. 7.

Wingspan. Male 22–25 mm. Type locality. S Vietnam. Distribution in Laos. Nakai, Bolikhamsai. Geographical range. Vietnam, Laos.

Genus Moore, 1878 Type species: Lithosia reticulata Moore, [1866] (Darjeeling)

Dolgoma rectoides Dubatolov, 2012 (Figs. 22, 100)

Dolgoma rectoides Dubatolov, 2012, Euroasian Ent. J. 11 (6): 507-508, pl. 3: 1, f. 1. Dolgoma rectoides Dubatolov: Singh, Kirti, Datta, Joshi & Volynkin, 2019, Zootaxa 4683 (1): 39-40.

Wingspan. Male 22 mm. Type locality. C Vietnam. Distribution in Laos. Huaphanh. Geographical range. Vietnam, Laos.

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Genus Teuloma Volynkin & Singh, 2019 Type species: Lithosia nebulosa Walker, 1862 (Borneo)

Teuloma nebulosa (Walker, 1862) (Figs. 23, 24, 101)

Lithosia nebulosa Walker, 1862, Journ. Soc. Zool. VI: 106. Lithosia nebulosa Walker: Swinhoe, 1892, Cat. Het. Mus. Oxon: 127, pl. IV, fig. 18. Lithosia nebulosa Walker: Kirby, 1892, Cat. Het.: 320. Ilema perdentata Druce, 1899, Ann. Mag. Nat. Hist. (7), 4: 201. Ilema nebulosa Walker: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 143. Ilema perdentata Druce: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 151, pl. 21: 26 (female). Lithosia perdentata Druce: Draudt, 1914: in Seitz, Macrolep. World 10: 208, pl. 14 k. Teulisna nebulosa Walker: Holloway, 2001, Moths of Borneo 7: 313, pl. 3, figs 66, 82. Eilema perdentata Druce: Fang, 1982, Icon. Het. Sin. 2: 199, fig. 1443. Systropha perdentata Druce: Fang, 2000, Fauna Sin., Ins. 19, Lep. Arctiidae: 249-250, fig. 172, pl. 12: 10. Teulisna nebulosa Walker: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 134, pl. 27: 268a, 268b. Teulisna nebulosa Walker: Bucsek 2014, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 115, Pl. 20: 285, 285a, 285b, 285c, 285d, 285e, Mal153. Teulisna nebulosa Walker: Kirti, Joshi & Singh, 2014, Acta zool. cracov. 57 (1-2): 18 (list). Teulisna nebulosa Walker: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 55. Teulisna nebulosa Walker: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 194, Pl. 18: 147. Teuloma nebulosa Walker: Singh, Kirti, Datta, Joshi & Volynkin, 2019, Zootaxa 4683 (1): 53, Figs 30–32, 47, 48, 56.

Wingspan. Male 21–25 mm, female 24–26 mm. Type locality. Borneo - Sarawak. Distribution in Laos. Huaphanh. Geographical range. NE India (Arunachal Pradesh), China (Yunnan), Thailand, Laos, Cambodia and Malaysia (Malay Peninsula and Borneo).

Genus Gampola Moore, 1878 Type species: Gampola fasciata Moore, 1878 (Ceylon=)

Gampola sinica Dubatolov, Kishida & Wang, 2012 (Figs. 25, 102)

Gampola sinica Dubatolov, Kishida & Wang, 2012, Tinea 22 (1): 35. Gampola fasciata Moore: Cerny & Pinratana, 2009, Moths Thailand 6: 153-154, pl. 31, fig. 310a, 310b. Gampola sinica Dubatolov: Joshi, Kirti & Singh, 2016, Tinea 23 (4): 220 (note). 53 Entomofauna carpathica, 2020, 32(2): 47-87

Wingspan. Female 26 mm. Type locality. S China. Distribution in Laos. Huaphanh. Geographical range. China (), , Thailand, Laos.

Genus Asiapistosia Dubatolov & Kishida, 2012 Type species: Eilema stigma Fang, 2000 (China - )

Asiapistosia subnigra (Leech, 1899) (Figs. 26, 27, 103, 104)

Oeonistis subnigra Leech, 1899, Trans. Ent. Soc. Lond. 1899 (1): 179. Apistosia subnigra Leech: Hampson, Cat. Lep. Phalaenae Br. Mus. 2: 227, pl. 24, f. 22. Apistosia subnigra Leech: Seitz, 1910, Groß-Schmett. Erde 2: 64, fig. 12e. Apistosia subnigra Leech: Strand, 1922, Lepid. Cat. 26: 627. Apistosia subnigra Leech: Daniel, 1953, Bonn. Zool. Beitr. 3 (3-4): 320. Apistosia subnigra Leech: Fang, 2000, Fauna Sinica, Insecta vol.19: 196, pl. 9, fig. 31. Asiapistosia subnigra Leech: Dubatolov, Kishida & Wang, 2012, Tinea 22 (1): 28, f. 3.

Wingspan. Male 29 mm, female 29 - 30 mm. Type locality. W China. Distribution in Laos. Huaphanh. Geographical range. China (Shaanxi, , , Hubei, Hunan, Sichuan, Yunnan, Guangdong), Laos.

Genus Thysanoptyx Hampson, 1894 Type species: Lithosia tetragona Walker, 1854 ()

Thysanoptyx indosinica Volynkin & Dubatolov, 2017 (Figs. 28, 29, 105)

Thysanoptyx indosinica Volynkin & Dubatolov, 2017, Zootaxa, 4277 (1): 113, 118, Figs. 1, 2, 17, 25. Thysanoptyx indosinica Volynkin & Dubatolov: Singh, Joshi, Kirti & Volynkin, 2017, Zootaxa 4319 (2): 375 (note).

Wingspan. Male 25–27 mm. Type locality. N Thailand. Distribution in Laos. Nakai. Geographical range. Laos, Thailand, Vietnam, Cambodia.

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Genus Brunia Moore, 1878 Type species: Lithosia antica Walker, 1854 (Ceylon=Sri Lanka)

Brunia sarawaca (Butler, 1877) (Figs. 30, 106)

Lithosia sarawaca Butler, 1877, Trans. Ent. Soc. 1877: 350. Eilema griseadisca Holloway, 1982: in Barlow, Introduction to Moths South-east Asia: 210. Brunia sarawaca Butler: Holloway, 2001, Moths of Borneo 7: 325, pl. 3, figs 110, 115. Brunia sarawaka [sic, recte sarawaca] Butler: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 149, pl. 30: 300a, 300b. Brunia sarawaca Butler: Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula-Malaysia, 121- 122, pl. 21:302, 302a. Brunia sarawaca Butler: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 27. Brunia sarawaka [sic, recte sarawaca] Butler: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 210, Pl. 19: 163. Brunia sarawaca Butler: Bucsek, 2017, Entomofauna 38 (8): 144, Pl. 3, 8, 10: fig. 2.-2l.

Wingspan. Female 24 mm. Type locality. Borneo – Sarawak. Distribution in Laos. Nakai. Geographical range. NE Himalaya, China, Thailand, Cambodia, Laos, Vietnam, Peninsular Malaysia, Java, Borneo, -Palawan.

Genus Cernyia Bucsek, 2012 Type species: Eilema pseudocretacea Holloway, 2001 (Borneo)

Cernyia pseudocretacea (Holloway, 2001) (Figs. 31, 32, 107, 108)

"Eilema" pseudocretacea Holloway, 2001, Moths of Borneo 7: 323, pl. 3, fig. 105. Cernyia pseudocretacea Holloway: Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula-Malaysia, 125, pl. 22: 310, 310a, Mal173. Cernyia pseudocretacea Holloway: Dubatolov & Bucsek, 2013, Tinea 22 (4): 279 (note). Cernyia pseudocretacea Holloway: Kirti, Singh, 2016, Arctiid Moths of India, vol. 2: 141.

Wingspan. Male 24–25 mm, female 28 mm. Type locality. Borneo – Sarawak. Distribution in Laos. Huaphanh. Geographical range. Borneo, Brunei, Peninsular Malaysia, Laos, Vietnam, India (Nagaland).

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Cernyia pseudobrevivalva (Dubatolov & Bucsek, 2016) n. comb. (Figs. 33, 34, 109, 110)

Katha pseudobrevivalva Dubatolov & Bucsek, 2016, Euroasian. Ent. J. 15 (3): 230, Figs 5, 23. pseudobrevivalva Dubatolov & Bucsek: Joshi, Singh & Singh, 2018, Zootaxa 4407 (3): 440 (list).

Wingspan. Male 25–26 mm, female 30–31 mm. Type locality. N Vietnam. Distribution in Laos. Huaphanh. Geographical range. Laos, Vietnam.

Genus Katha Moore, 1878 Type species: Bombyx helvola Hübner, [1803] (Europe)

Katha montana Bucsek, 2012 (Figs. 35, 36, 111)

Katha montana Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula-Malaysia, 123, pl. 22: 306, 306a, 306b, Mal166.

Wingspan. Male 25 mm. Type locality. Malaysia – Pahang. Distribution in Laos. Huaphanh. Geographical range. Laos, Peninsular Malaysia.

Genus Moore, 1878 Type species: Lithosia serva Walker, 1854 ()

Gandhara vietnamica Dubatolov, 2012 (Figs. 37, 38)

Gandhara vietnamica Dubatolov, 2012, Euroasian Ent. J., 11 (6): 508, Pl. 3: 4, Fig. 3. Gandhara serva Walker: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 147, pl. 30: 297.

Wingspan. Male 20–28 mm, female 26–30 mm. Type locality. Vietnam. Distribution in Laos. Huaphanh. Geographical range. Vietnam, Laos, Thailand.

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Genus Bucsekia Dubatolov, 2012 Type species: yazakii Dubatolov, 2012 (China)

Bucsekia mediumpilosa Bucsek, 2012 (Figs. 39, 40, 112, 113)

Bucsekia mediumpilosa Bucsek, 2012, Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula-Malaysia, 135, pl. 24: 334, 334a, Mal175. Bucsekia mediumpilosa Bucsek: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 228, Pl. 21: 183.

Wingspan. Male 20–22 mm, female 21 mm. Type locality. Malaysia - Pahang. Distribution in Laos. Huaphanh. Geographical range. Laos, Cambodia, Peninsular Malaysia.

Genus Microlithosia Daniel, 1954 Type species: Microlithosia shaowuica Daniel, 1954 (China – Fukien)

Microlithosia unicolora Bucsek n. sp. (Figs. 41, 42, 114)

Holotype. ♂ LAOS, Huaphanne prov., Mt. Phu Pane, 1200-1900m, Ban Saluei v. env., 3.-13.V.2019, 20°12´N, 103°59´E, leg. K. Bucsek, SNM with a museum serial number SZ 10 958. Paratype. 1 ♂ like holotype, CKB.

Wingspan. Male 17–18 mm. Holotype. Male 18 mm, antennae finely pectinated, head, thorax and abdomen are dark yellow, anal margin lighter. Forewing dark yellow without patterning, hindwing a little lighter. Male genitalia. Uncus long and narrow, slightly bent, pointed at the end. Valva wide, on the apical lobe of valva a series of small spikes. Discal saccular process long and narrow, strongly arched inward with a blunt termination, with two pointed processes in the upper third on the outside. Juxta roof-shaped. Aedeagus ovoid with a long medial process, which has three different sized spines on the base. Female. Unknown. Similar species: Microlithosia jagbiri Singh & Joshi has a discal saccular process terminated by a sharp point.

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Microlithosia nanlingica Dubatolov, Kishida & Wang, 2012 (Figs. 43, 115)

Microlithosia shaowuica (part.): Fang, 2000, Fauna Sinica (Insecta) 19: 287-288, fig. 202 (male genitalia), pl. 13, fig. 23. Microlithosia nanlingica Dubatolov, Kishida & Wang, 2012, Tinea 22 (1): 39, f. 47, 99. Microlithosia nanlingica Dubatolov, Kishida & Wang: Dubatolov & Kishida, 2012, Amurian Zool. J. 4 (2): 179, pl. 4: 2. Microlithosia nanlingica Dubatolov, Kishida & Wang: Bayarsaikhan & Bae, 2018, J. Asia- Pacif. Biodiv. 11 (3): 387, 388, Figs 1D, 4D. Microlithosia nanlingica Dubatolov, Kishida & Wang: Bayarsaikhan, Lee, Bae, 2018, Zootaxa 4420 (4): 505, Figs. 3, 6.

Wingspan. Male 16–18 mm. Type locality. S China. Distribution in Laos. Huaphanh. Geographical range. China (Guangdong), Laos, Vietnam.

Genus Danielithosia Dubatolov & Kishida, 2012 Type species: Tigrioides aureolata Daniel, 1954 (China – Chekiang)

Danielithosia zolotuhini Dubatolov, 2012 (Figs. 44–46, 116–118)

Danielithosia zolotuhini Dubatolov, 2012, Euroasian Ent. J. 11 (6): 510, pl. 3: 5, f. 4. Danielithosia zolotuhini Dubatolov: Dubatolov, 2013, Acta Ent. Mus. Nat. Pragae 53 (1): 386 (list). Danielithosia zolotuhini Dubatolov: Bayarsaikhan, Dubatolov, Heppner & Bae, 2017, Zootaxa 4291 (3): 574, Figs. 2, 3.

Wingspan. Male 19–20 mm, female 19–22 mm. Type locality. N Vietnam. Distribution in Laos. Huaphanh. Geographical range. Laos, Vietnam.

Danielithosia dubatolovi Bucsek n. sp. (Figs. 47–50, 119, 120)

Holotype. ♂ LAOS, Bolikhamsai, Khoun Ngeun env., 200–300 m, 18°08′48.9″N, 04°27′44.7″E, 29. IV.–1. V. 2019, K. Bucsek leg., SNM with a meseum serial number SZ 10 959. Paratypes. 4 ♂♂, 1 ♀ like holotype, 1 ♀ LAOS, Khammouane prov., Nakai env., 500 m, 17°34´N, 105°10´E, 25.- 29. IV. 2019, leg. K. Bucsek, CKB.

58 Entomofauna carpathica, 2020, 32(2): 47-87

Wingspan. Male 17–19 mm, female 20 mm. Holotype. Male 19 mm, antennae finely pectinated, head light brown, thorax dark brown abdomen creamy brown, anal margin creamy. Forewing homogenously brown, costa from base to 2/3 light gray-brown. Hindwing light brown. Male genitalia. Uncus straight, pointed. Valva at the base very wide, slightly narrowed to the top, termination rounded. Sacculus indistinct, distal saccular process spatulate. Juxta massive, cylindrical, with two asymmetrical processes. The left process long and bent inward (extends to about 1/2 uncus), with small node at the base on the inside; the right process short (about 1/3 of the length of the left) and straight. Aedeagus cylindrical, curved, with one distinct cornut, a row of four smaller cornuti and a field of very small cornuti. Female. Habitually the same as a male, antennae filiform. Female genitalia. Antevaginal plate sclerotized, corpus bursae massive with a distinct teardrop-shaped signum. Appendix bursae massive, ovoid. Etymology. I named the species after my friend and excellent expert in the Arctiinae Palearct and Oriental region Vladimir Viktorovich Dubatolov, ScDr (Novosibirsk, Russia).

Genus Poliosia Hampson, 1900 Type species: Lithosia muricolor Walker, 1862 (Borneo)

Poliosia cubitifera (Hampson, 1894) (Figs. 51, 52, 121, 122)

Lithosia cubitifera Hampson, 1894, Fauna Br. India (Moths) 2: 82. Poliosia cubitifera Hampson: Hampson, 1900, Cat. Lep. Phalaenae Br. Mus. 2: 108, pl. 20, f. 23. Poliosia cubitifera Hampson: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 51. Poliosia cubitifera Hampson: Bayarsaikhan, Na, & Bae, 2018, Zootaxa 4482 (1): 147 (list). Poliosia cubitifera Hampson: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 158, pl. 32: 320.

Wingspan. Male 17–18 mm. Type locality. India - Assam. Distribution in Laos. Huaphanh. Geographical range. India (Sikkim, Assam), China, Laos, Thailand.

Poliosia rectilinea de Joannis, 1928 (Figs. 53, 123)

Poliosia rectilinea de Joannis, 1928, Ann. Soc. Ent. Fr. 97: 253, Pl. 1, fig. 2. Poliosia rectilinea de Joannis: Bayarsaikhan, Na & Bae, 2018, Zootaxa 4482 (1): 148 (list).

59 Entomofauna carpathica, 2020, 32(2): 47-87

Wingspan. Male 16 mm. Type locality. N Vietnam. Distribution in Laos. Nakai. Geographical range. Vietnam, Laos.

Poliosia bilineata Bucsek n. sp. (Figs. 54, 124)

Holotype. ♂ LAOS, Khammouane prov., Nakai env., 500 m, 17°34´N, 105°10´E, 25.- 29. IV. 2019, leg. K. Bucsek, SNM with a meseum serial number SZ 10 960.

Wingspan. Male 16 mm. Holotype. Antennae finely pectinated, head ochreous, thorax and abdomen dark brown. Forwing ochreous, with dark brown stripes. The medial line is indistinct, slightly bent inwards in the middle, postmedial line indistinctive zigzag, more or less parallel termen. Hindwing ochreous with strong brown dust in subterminal area. Male genitalia. Uncus thick, more or less straight, pointed at the end. Valva narrow, at the top bent inside. Discal saccular process thick, curved inside, pointed. Juxta roof-shaped. Vinculum massive in the shape of Y. Aedeagus cylindrical without diagnostic characters. Female. Unknown.

Poliosia cardamomensis Bayarsaikhan & Bae, 2018 (Figs. 55–57, 125, 126)

Poliosia cardamomensis Bayarsaikhan & Bae, 2018, Zootaxa 4482 (1): 142, 145, Figs. 1a, 1b, 6a˗d, 7.

Wingspan. Male 13–14 mm, female 14 mm. Type locality. Cambodia. Distribution in Laos. Nakai. Geographical range. Laos, Cambodia.

Genus Veslema Bucsek, 2012 Type species: binotata Hampson, 1893 (Ceylon=Sri Lanka)

Veslema bipunctulata Bucsek, 2012 (Fig. 58)

Veslema bipunctulata Bucsek, 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 134-135, pl. 24: 333, 333a, Mal179.

60 Entomofauna carpathica, 2020, 32(2): 47-87

Wingspan. Female 18 mm. Type locality. Malaysia – Pahang. Distribution in Laos. Huaphanh. Geographical range. Laos, Peninsular Malaysia.

Genus Nishada Moore, 1878 Type species: Nishada flabrifera Moore, 1878 (India - Calcutta District)

Nishada flabrifera Moore, 1878 (Figs. 59, 60, 127)

Nishada flabrifera Moore, 1878, Proc. Zool. Soc. 1878: 23. Nishada flabrifera Moore: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 113, fig. 53. Nishada flabrifera Moore: Kishida, 1995, Moths of Nepal 4: 42, pl. 108: 14. Nishada flabrifera Moore: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 155, pl. 32: 312. Nishada flabrifera Moore: Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula- Malaysia, 131-132, pl. 23: 325, 325a. Nishada flabrifera Moore: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 48. Nishada flabrifera Moore: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 123. Nishada flabrifera Moore: Joshi, Kirti & Singh, 2016, Zootaxa 4179 (1): 128 (note).

Wingspan. Male 26–28 mm, female 29–30 mm. Type locality. India – Calcutta District. Distribution in Laos. Huaphanh, Nakai. Geographical range. Sri Lanka, India, Myanmar, Thailand, Laos, Java, Peninsular Malaysia.

Genus Agrisius Walker, 1855 Type species: Agrisius guttivitta Walker, 1855 (India – Sikkim)

Agrisius leloii Volynkin, Dubatolov & Kishida, 2018 (Fig. 61)

Agrisius leloii Volynkin, Dubatolov & Kishida, 2018, Zootaxa 4459 (2): 131, 137, Figs 9, 10, 26, 27, 36.

Wingspan. Female 51 mm. Type locality. C Vietnam. Distribution in Laos. Huaphanh. Geographical range. Vietnam, Laos.

61 Entomofauna carpathica, 2020, 32(2): 47-87

Genus Vamuna Moore, 1878 Type species: Lithosia remelana Moore, 1865 (India – Darjeeling)

Vamuna remelana (Moore, 1865) (Fig. 62)

Lithosia remelana Moore, 1865, Proc. Zool. Soc. Lond. 1865: 798 Crambomorpha ramelana Moore: Butler, 1877, Trans. Ent. Soc. Lond. 1877: 357. Vamuna ramelana Moore: Moore, 1878, Proc. Zool. Soc. Lond. 1878: 10. Sidyma ramelana Moore: Hampson, 1894, Moths. Ind. 2: 68. Agylla ramelana Moore: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 211, pl. 24: 18 Agylla ramelana Moore: Draudt, 1914: in Seitz, Macroler. World 10: 200, pl. 15 i. Agylla ramelana Moore: Daniel, 1952, Bonn. Zool. Beitr. 3: 321. Agylla ramelana Moore: Fang, 1982, Icon. Het. Sin. 2: 201, fig. 1472 Vamuna ramelana Moore: Kishida, 1993, Tinea 13 (Suppl. 3): 36. Vamuna ramelana Moore: Fang, 2000, Fauna Sin., Ins. 19, Lep. Arctiidae: 208, fig. 130, pl. 10: 14. Vamuna remelana Moore: Holloway, 2001, Moths of Borneo 7: 289, pl. 1, figs 13, 14. Vamuna remelana Moore: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 124, pl. 25: 248a, 248b. Vamuna remelana Moore: Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula-Malaysia, 106-107, pl. 18: 263, 263a, 263b, 263c, 263d. Vamuna remelana Moore: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 58. Vamuna remelana Moore: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 165.

Wingspan. Male 43 mm. Type locality. India – Darjeeling. Distribution in Laos. Huaphanh. Geographical range. India (Himalaya), China, Vietnam, Thailand, Peninsular Malaysia, Borneo, Sumatra, Java.

Genus Herrich-Schäffer, [1855] Type species: Macrobrochis interstitialis Herrich-Schäffer, 1856 (=gigas Walker, 1854) (India)

Macrobrochis fukiensis (Daniel, 1952) (Fig. 63)

Paraona fukiensis Daniel, 1953, Bonn. zool. Beitr. 3 (3–4): 319, (1-2): pl. f. 56. Macrobrochis fukiensis Daniel: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 191, pl. 9: 23. Macrobrochis fukiensis Daniel: Cerny & Pinratana, 2009, Moths of Thailand, vol. 6: 121, pl. 24: 243. Macrobrochis fukiensis Daniel: Bucsek, 2016, Entomofauna 37 (37): 574, pl. 1: 1. Macrobrochis fukiensis Daniel: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 233, Pl. 22: 188.

62 Entomofauna carpathica, 2020, 32(2): 47-87

Wingspan. Male 56 mm. Type locality. China – Fujian. Distribution in Laos. Bolikhamsai. Geographical range. China, Vietnam, Laos, Thailand, Cambodia, Peninsular Malaysia.

Genus Churinga Moore, 1878 Type species: Churinga rufifrons Moore, 1878 (India – Darjiling)

Churinga nigripes (Hampson, 1900) n. comb. (Figs. 64, 128)

Ilema nigripes Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 150, 175-176, pl. XXIII, fig. 6. Eilema nigripes Hampson: Cerny & Pinratana, 2009, Moths of Thailand, vol. 6: 146-147, pl. 30: 295. Eilema nigripes Hampson: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 35.

Wingspan. Male 26 mm. Type locality. India – Sikkim. Distribution in Laos. Bolikhamsai. Geographical range. India (Sikkim, Assam), Laos, Thailand.

Genus Chrysaeglia Butler, 1877 Type species: Lithosia magnifica Walker, 1862 (Borneo)

Chrysaeglia magnifica Walker, 1866 (Figs. 65)

Chrysaeglia magnifica Walker, 1866, J. Linn. Soc. (Zool.) 6: 103. Chrysaeglia ferrifasciata Moore, 1878, Proc. Zool. Soc. Lond. 1878: 13. Chrysaeglia magnifica Walker: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 185, f. 102. Chrysaeglia magnifica Walker: Hampson, 1914, Cat. Lep. Phal. Br. Mus. (Suppl.) 1: 511. Chrysaeglia magnifica Walker: Fang, 2000, Fauna Sin. Ins.19, Lep. Arctiidae: 222, f. 142, 143, pl.11: 7. Chrysaeglia magnifica Walker: Holloway, 2001, Moths of Borneo 7: 292, pl 1, figs 1d, 26. Chrysaeglia magnifica Walker: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 28. Chrysaeglia magnifica Walker: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 180.

Wingspan. Male 42–52 mm, female 50–60 mm. Type locality. Borneo – Sarawak. Distribution in Laos. Huaphanh. Geographical range. NE Himalaya, China, Laos, Thailand, , Borneo, Sulawesi. 63 Entomofauna carpathica, 2020, 32(2): 47-87

Genus Hübner, [1819] Type species: Phalaena ornatrix Linne, 1758 (North America)

Utetheisa pulchelloides vaga Jordan, 1939 (Fig. 66)

Utetheisa pulchelloides ssp. vaga, Jordan, 1939, Nov. Zool. XLI: 284-285. Linne: Hampson, 1901, Cat. Lep. Phal. Br. Mus. 3: 483, fig. 217 (partim). Utetheisa lotrix Cramer: Rothschild, 1914: in Seitz, Macrolep. World 10: 260 (partim). Utetheisa pulchelloides Hampson: Holloway, 1988: Moths of Borneo, Vol. 6: 56, pl. 4, figs 92, 94, 95. Utetheisa pulchelloides ssp. vaga Jordan: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 310, fig. 217, pl. 5: 28, 29. Utetheisa pulchelloides ssp. vaga Jordan: Cerny & Pinratana, 2009, Moths of Thailand, vol. 6: 192, pl. 44: 384a, 384b. Utetheisa pulchelloides ssp. vaga Jordan: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 261, Pl. 24: 214.

Wingspan. Female 34–35 mm. Type locality. . Distribution in Laos. Bolikhamsai, Nakai. Geographical range. Seychelles, Sri Lanka, India, Myanmar, China, Laos, Cambodia, Thailand, , Indo-Australian and Pacific tropics, Australia, New Zealand.

Genus Hübner, [1820] Type species: Phalaena (Geomerta) lacticinia Cramer, [1777] (India)

Nyctemera adversata (Schaller, 1788) (Fig. 67)

Phalaena (Geometra) adversata Schaller, 1788, Beytr.Ges.Exot. Pap. Naturf. 23: 54, pl. 1: 13. Nyctemera plagifera Walker, 1854, List Spec. Lepid. Ins. Coll. Br. Mus. 2: 400. Nyctemera plagifera Walker: Hampson, 1894, Moths Ind. 2: 474. Nyctemera plagifera Walker: Seitz, 1910, Macrolep. World 2: 103, pl. 18h. Nyctemera plagifera Walker: Daniel 1943, Mitt. Münch. Ent. Ges., Munich 33: 269. Nyctemera plagifera Walker: Roepke, 1957, Tijd. Ent. 100: 155. Schaller: Holloway, 1988: Moths of Borneo, Vol. 6: 68, pl. 5, figs 118, 137. Nyctemera adversata Schaller: Fang, 1992, Icon of For. Ins. in Hunan China: 931, fig. 3134. Nyctemera adversata Schaller: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 474 - 475, fig. 330, pl. 20: 2. Nyctemera adversata Schaller: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 199, pl. 46: 398. Nyctemera (Orphanos) adversata Schaller: Dubatolov V.V., 2010, Neue Ent. Nach. 65: 13.

64 Entomofauna carpathica, 2020, 32(2): 47-87

Nyctemera adversata Schaller: Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula- Malaysia, 140, pl. 25: 344. Nyctemera adversata Schaller: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 28. Nyctemera adversata Schaller: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 180. Nyctemera adversata Schaller: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 261, Pl. 24: 214.

Wingspan. Male 43 mm, female 43–50 mm. Type locality. Hong Kong. Distribution in Laos. Huaphanh. Geographical range. Himalaya, Nepal, W. and S. China, Laos, Cambodia, Japan, Thailand, Peninsular Malaysia, Sumatra, Borneo, Philippines.

Nyctemera tripunctaria Linne, 1758 (Fig. 68)

Nyctemera tripunctaria Linne, 1758, Syst. Nat. (Ed. 10): 523. Nyctemera tripunctaria Linne: Walker, 1854, List Spec. Lepid. Ins. Coll. Br. Mus. 2: 397. Nyctemera tripunctaria Linne: Hampson, 1894, Moths Ind. 2: 47. Deilemera tripunctaria Linne: Swinhoe, 1903, Trans. Ent. Soc. Lond. 1903: 53, 80. Nyctemera tripunctaria Linne: Seitz, 1915, Macrolep. World 10: 267, pl. 29a. Nyctemera tripunctaria Linne: van Eecke, 1928, Zool. Meded. Mus. Leiden 11: 58-59. Nyctemera tripunctaria Linne: Fang, 1982, Icon. Het. Sin. 2: 226, fig. 1660. Nyctemera tripunctaria Linne: Fang, 1985, Econ. Ins. Fauna of China 33: 88, pl. 10: 142. Nyctemera tripunctaria Linne: Holloway, 1988: Moths of Borneo, Vol. 6: 69, pl. 5, figs 124, 136. Nyctemera tripunctaria Linne: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 472, pl. 19: 13. Nyctemera tripunctaria Linne: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 199, pl. 46: 397. Nyctemera tripunctaria Linne: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 13.

Wingspan. Male 45 mm. Type locality. Not stated. Distribution in Laos. Huaphanh. Geographical range. S. China, Laos, Thailand, Vietnam, Sundaland, Philippines, Sulawesi.

Genus Hübner, [1819] Type species: Phalaena interrupta Linne, 1767 (=gangis Linne)

Creatonotos transiens (WALKER, 1855) (Figs. 69, 70)

Spilosoma transiens Walker, 1855, List. Spec. Lep. Ins. Coll. Br. Mus. 3: 675. Amphissa vacillans Walker, 1855, Ibid. 3: 685. 65 Entomofauna carpathica, 2020, 32(2): 47-87

Aloa isabellina Walker, 1855, Ibid. 3: 705. Phisama transiens Walker: Hampson, 1894, Moths Ind. 2: 29, fig. 10. Creatonotus transiens Walker: Hampson, 1901, Cat. Lep. Phal. Br. Mus. 3: 334 - 335, fig. 150. Phisama transiens Walker: Seitz, 1910, Macrolep. World 2: 90, pl. 17a. Creatonotus transiens Walker: Rothschild, 1910, Nov. Zool. 17: 160 - 161. Walker: Strand, 1919, Lep. Cat. 22: 246-247. Creatonotos transiens orientalis Walker: Nakamura, 1976,Tyô to Ga 27 (3): 116,fig.1 (3), 3 (5). Creatonotus transiens Walker: Fang, 1982, Icon. Het. Sin. 2: 222, fig. 1632. Creatonotos transiens Walker: Koda, 1988, Tyo to Ga 39 (1): 39, fig. 82, 93, 84, 113 G, H. Creatonotos transiens Walker: Holloway, 1988: Moths of Borneo 6: 52, pl.3, figs 88, 107. Creatonotos transiens Walker: Kuroko & Lewvanich, 1993: Lepid. Pests Tropic. Fruit Trees Thailand. Creatonotos transiens Walker: Fang, 2000, Fauna Sin., Ins. 19, Lep. Arctiidae: 367 - 368, fig. 256, pl. 15: 9. Creatonotos transiens vacillans Walker: Dubatolov V., Holloway J. 2007, Bonner zoologische Beitr[ae]ge. - Bd 55, Heft 2, 118, f. 4, 16 - 18. Creatonotos transiens Walker: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 167, pl. 35: 337a, 337b. Creatonotos (Phissama) transiens vacillans Walker: Dubatolov V.V., 2010, Neue Ent. Nach. 65: 44. Creatonotos (Phissama) transiens vacillans Walker: Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula-Malaysia, 144, pl. 25: 352, 352a. Creatonotos transiens Walker: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 242, Pl. 22: 195. Creatonotos (Phissama) transiens Walker: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 38.

Wingspan. Male 38–45 mm, female 45–50 mm. Type locality. Hong Kong. Distribution in Laos. Huaphanh, Bolikhamsai, Nakai. Geographical range. Himalaya, Nepal, W. and S. China, Laos, Cambodia, Japan, Thailand, Peninsular Malaysia, Sumatra, Borneo, Philippines, Borneo, Sulawesi.

Creatonotos gangis (Linne, 1763) (Figs. 71, 72)

Phalaena gangis Linne, 1763, Amoenitates Acad., 6: 410. Noctua interrupta Linne, 1767, Syst. Nat. (Edn 12) 1 (2): 840. Bombyx francisca Fabricius, 1787, Mantissa Insectorum 2: 131, 200. Creatonotum continuatus Moore, 1877; Ann. Mag. nat. Hist. (4) 20 (118): 344. Creatonotus interruptus Moore, 1882, Lepid. Ceylon 2 (2): 73, pl. 108, f. 5, 5a. Creatonotus continuatus Moore, 1882, Lepid. Ceylon 2 (2): 73, pl. 108, f. 4. Creatonotus gangis Linne: Hampson, 1901, Cat. Lepid. Phalaenae Br. Mus. 3: 333. Creatonotus gangis Linne: Rothschild, 1910, Novit. Zool. 17 (2): 160. Creatonotus gangis Linne: Seitz, 1910, Macrolep. World 2: 90, pl. 17a. Linne: Strand, 1919, Lep. Cat. 22: 245-246. 66 Entomofauna carpathica, 2020, 32(2): 47-87

Creatonotos gangis Linne: Fang, 1985, Econ. Ins. Fauna of China 33:68, pl. 6: 101. Creatonotos gangis Linne: Koda, 1988, Tyo to Ga 39 (1):39, fig. 113 E, F. Creatonotos gangis Linne: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 368-369, fig. 257, pl. 15: 10. Creatonotos gangis Linne: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 166, pl. 35: 336a, 336b. Creatonotos gangis Linne: Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula-Malaysia, (Suppl.), 33, pl. 6: 73. Creatonotos gangis Linne: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 241, Pl. 22: 194. Creatonotos gangis Linne: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 37, 38.

Wingspan. Male 36–42 mm, female 45 mm. Type locality. Not stated. Distribution in Laos. Huaphanh. Geographical range. India and China to New Guinea and N. Australia.

Genus Curtis, 1825 Type species: Bombyx menthastri Denis & Schiffermüller, 1775 (Europe)

Spilosoma nydia tienmushanica (Daniel, 1943) (Fig. 73)

Spilarctia tienmushanica Daniel, 1943, Mitt. Münch. Ent. Ges., Munich 33: 694, pl 17: 6 - 9. Spilarctia tienmushanica Daniel: Fang, 1985, Econ. Ins. Fauna of China 33: 58, pl. 5: 81. Spilarctia tienmushanica Daniel: Fang, 2000, Fauna Sin., Ins. 19, Lep. Arctiidae: 442 - 443, fig. 317, pl. 18: 6. Spilosoma tienmushanica Daniel: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 182, pl. 41: 367a, 367b, gen. 4: 367f, 367v. Spilarctia nydia tienmushanica Daniel: Dubatolov V.V., 2010, Neue Ent. Nach. 65: 56. Spilosoma tienmushanica Daniel: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 259, Pl. 24: 212.

Wingspan. Male 42 mm. Type locality. China - Zhejiang. Distribution in Laos. Huaphanh. Geographical range. China, Vietnam, Laos, Thailand, Cambodia.

Spilosoma bisecta Leech, [1889] (Figs. 74–76)

Spilosoma bisecta Leech, [1889], Proc. Zool. Soc. London 1888: 618-619, pl. 31, fig. 3. Spilosoma bisecta Leech: Leech, 1899, Trans. Ent. Soc. Lond. 1899 (1): 148. Spilosoma mandarina Moore, 1877, Ann. Mag. nat. Hist. (4) 20 (116): 88. 67 Entomofauna carpathica, 2020, 32(2): 47-87

Spilosoma howqua Moore, 1877, Ann. Mag. nat. Hist. (4) 20 (116): 88. Spilarctia bisecta shanghaiensis Daniel, 1943, Mitt. Münch. Ent. Ges. 33 (3): 692, pl. 18, f. 9-12.

Wingspan. Male 40–43 mm, female 48 mm. Type locality. Hong Kong. Distribution in Laos. Huaphanh. Geographical range. China, Myanmar, Vietnam, Laos, Thailand.

Spilosoma sagittifera (Moore, 1888) (Figs. 77–79)

Spilarctia sagittifera Moore, 1888, Proc. zool. Soc. Lond. 1888: 394. Spilosoma sagittifera Moore: Thomas, 1994, Heteroc. Sumatr. 7 (2): 184-186. Spilarctia sagittifera Moore: Kishida, 1995, Tinea 14 (suppl. 2): 43, pl. 107, f. 13-14. Spilarctia sagittifera Moore: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 18.

Wingspan. Male 40–44 mm. Type locality. India – Dharmsala. Distribution in Laos. Huaphanh. Geographical range. , India, Nepal, Thailand, Vietnam, Laos, China, Taiwan.

Genus Lemyra Walker, 1856 Type species: Lemyra extensa Walker, 1856 (Sulawesi)

Lemyra quadrisaccus (Holloway, 1982) (Figs. 80)

Paralacydes quadrisaccus Holloway, 1982, in Barlow, An introduction to the moths of South-east Asia: 217. Lemyra quadrisaccus Holloway: Thomas, 1990, Nachr. Ent. Ver. Apollo, Supp. 9, 52: fig. 29. Lemyra quadrisaccus Holloway: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 177, pl. 39: 357a, 357b. Lemyra quadrisaccus Holloway: Dubatolov V.V., 2010, Neue Ent. Nach. 65: 49. Lemyra quadrisaccus Holloway: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 246, Pl. 23: 199.

Wingspan. Male 28–33 mm. Type locality. Malysia - W. Pahang. Distribution in Laos. Huaphanh. Geographical range. Malaysia (Pahang), Thailand, Vietnam, Laos.

68 Entomofauna carpathica, 2020, 32(2): 47-87

Lemyra gloria Fang, 1993 (Figs. 81, 129)

Spilarctia multivittata assama Rothschild: Fang, 1982, Icon. Het. Sin. 2: 215 fig. 1590. Spilarctia multivittata assama Rothschild: Fang, 1985, Econ. Ins. Fauna of China 33: 47-48, pl. 3: 49. Lemyra gloria Fang, 1993, Scient.Treatise on Systematic and Evol. Zoology 2: 181, 191, fig. 10, pl. 8: 11. Lemyra gloria Fang: Fang, 2000, Fauna Sin., Ins. 19, Lep. Arctiidae: 389, fig. 274, pl. 16: 6.

Wingspan. Male 27–28 mm. Type locality. China – Yunnan. Distribution in Laos. Huaphanh. Geographical range. China (Yunnan), Laos.

Lemyra ?multivittata (Moore, 1865) (Figs. 82, 130)

Spilosoma multivittata Moore, 1865, Proc. Zool. Soc. Lond. 1865: 808. Thyrgorina multivittata Moore: Swinhoe, 1892, Cat. Het. Oxford Mus. 1: 180. Diacrisia multivittata Moore: Hampson, 1901, Cat. Lep. Phal. Br. Mus. 3: 257, 265, pl. 44, 44. Diacrisia multivittata Moore: Rothschild, 1914: in Seitz, Macrolep. World 10: 239, pl. 20 e. Lemyra multivittata Moore: Fang, 1993, Scient. Treatise and Evolut. Zool. 2: 180, fig. 9, pl. 8: 10. Spilarctia multivittata assama Rothschild: Fang, 1982, Icon. Ins. Fauna of China 33: 47, pl. 3: 49. Thanatarctia multivittata Moore: Koda, 1988, Tyo to Ga 39 (1): 19, fig. 110C, D. Lemyra multivittata Moore: Thomas, 1990, Nachr.ent.Ver.Apollo, Frankfurt, Suppl. 9: 26, Abb. 11. Lemyra multivittata Moore: Kishida, 1993, Tinea 14 (Suppl. 1): 68, pl. 18: 21. Lemyra multivittata Moore: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 381-382, fig. 267, pl. 15: 21. Lemyra multivittata Moore: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 10.

Wingspan. Male 27 mm. Type locality. India. Distribution in Laos. Huaphanh. Geographical range. India (Darjeeling, Sikkim), Nepal, China, Laos, Thailand, Vietnam.

Lemyra ?crama Černý, 2009 (Figs. 83, 131)

Lemyra crama Černý, 2009, Cerny, Pinratana, Moths of Thailand, vol. 6: 178, pl. 39: 358, gen. 3: 358f, 358v.

Wingspan. Male 31 mm. Type locality. N Thailand. Distribution in Laos. Huaphanh. Geographical range. Thailand, ?Laos. 69 Entomofauna carpathica, 2020, 32(2): 47-87

Lemyra ?anormala (Daniel, 1943) (Figs. 84, 132)

Spilarctia rhodophila anormala Daniel, 1943, Mitt. Münch. Ent. Ges., Munich 33: 710, pl. 19: 16-17. Lemyra anormala Daniel: Thomas, 1990, Nachr.ent.Ver.Apollo, Frankfurt, Suppl. 9: 16, Abb. 42, 43. Lemyra anormala Daniel: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 376-377, figs 260, pl. 15: 15. Lemyra anormala Daniel: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 168-169, pl. 36: 339a, 339b.

Wingspan. Male 30 mm. Type locality. China – Yunnan. Distribution in Laos. Huaphanh. Geographical range. China (Yunnan), Laos, Thailand, Myanmar.

Genus Paraeuchaetes Grote, [1866] Type species: Paraeuchaetes cadaverosa Grote, [1866] (Cuba)

Paraeuchaetes pseudoinsulata Rego Barros, 1956 (Fig. 85)

Paraeuchaetes pseudoinsulata Rego Barros, 1956, Revta Bras. Ent. 6: 79. Paraeuchaetes pseudoinsulata Rego Barros: Holloway, 1988: Moths of Borneo, Vol. 6: 63, pl.4. Paraeuchaetes pseudoinsulata Rego Barros: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 190, pl. 43: 380a, 380b. Paraeuchaetes pseudoinsulata Rego Barros: Dubatolov V.V., 2010, Neue Ent. Nach. 65: 2. Paraeuchaetes pseudoinsulata Rego Barros: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 59. Paraeuchaetes pseudoinsulata Rego Barros: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 253, Pl. 24: 206.

Wingspan. Male 28–33 mm, female 35–43 mm. Type locality. Venezuela – Caracas. Distribution in Laos. Huaphanh. Geographical range. Venezuela, Trinidad; introduced and established in Sri Lanka, India, Laos, Thailand, Cambodia, Peninsular Malaysia, Singapore, Borneo, Palawan, Guam.

Genus Calpenia Moore, 1872 Type species: Calpenia saundersi Moore, 1872 (India)

70 Entomofauna carpathica, 2020, 32(2): 47-87

Calpenia saundersi Moore, 1872 (Fig. 86)

Calpenia saundersi Moore, 1872, Proc. zool. Soc. Lond. 1872 (2): 571, pl. 33, f. 1. Calpenia saundersi Moore: Seitz, 1915, Macrolep. World 10: 265, pl. 26f. Calpenia saundersi Moore: Fang, 1985, Econ. Ins. Fauna of China 33: 83-84, pl. 9: 131. Calpenia saundersi Moore: Fang, 2000, Fauna Sin., Ins. 19, Lep. Arctiidae: 457, pl. 19: 2. Calpenia saundersi Moore: Dubatolov, 2004, Atalanta 35(1/2): 75, figs 4, 5 (genitalia). Calpenia saundersi Moore: Witt & Speidel, 2006, Entomofauna 27 (2): 38 (list). Calpenia saundersi Moore: Singh, Singh & Joshi, 2014, Rec. Zool. Survey India Occ. Pap 367: 6. Calpenia saundersi Moore: Kishida, 2015, Sel. arctiid moths of the World, 55, pl. 2: 2, 3, 4.

Wingspan. Female 85 mm. Type locality. India – Sikkim. Distribution in Laos. Huaphanh. Geographical range. India (Sikkim), China (Yunnan, Tibet), Laos.

Genus Areas Walker, 1855 Type species: Areas orientalis Walker, 1855 (N India)

Areas galactina inouei Dubatolov, Haynes & Kishida, 2009 (Fig. 87)

Areas galactina inouei Dubatolov, Haynes & Kishida, 2009, Tinea 20 (5): 319-322, fig. 6, gen. 37. Areas galactina inouei Dubatolov, Haynes & Kishida: Nakao, Černý, 2013, Nachr. Ent. Ver. Apollo, NF 34 (1/2): 89, Figs. 1a, 1b. Areas galactina inouei Dubatolov, Haynes & Kishida: Kishida, 2015, Sel. arctiid moths of the World, 81, pl. 34: 4, 5.

Wingspan. Male 73 mm. Type locality. N Vietnam. Distribution in Laos. Huaphanh. Geographical range. China (Yunnan), Vietnam, Laos, Thailand.

Areas aquila Nakao & Černý, 2013 (Figs. 88, 89)

Areas aquila Nakao & Černý, 2013, Nachr. Ent. Ver. Apollo, NF 34 (1/2): 89, Figs. 2a, 2b.

Wingspan. Male 70 mm, female 74 mm. Type locality. N Laos. Distribution in Laos. Huaphanh, Xienkhouang. Geographical range. Laos, Thailand. 71 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 1-12. Adults. 1. Teulisna karena, ♂. 2. Teulisna uniplaga, ♂. 3-4. Teulisna fimbriata, 3. ♂, 4. ♀. 5-6. Teulisna protuberans, 5. ♂, 6. ♀. 7-8. Teulisna tumida, 7. ♂, 8. ♀. 9-10. Teulisna maculata, 9. ♂, 10. ♀. 11-12. Teulisna tongdzuythangi, 11. ♂, 12. ♀. 72 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 13-26. Adults. 13-15. Teulisna mithunoides, 13-14. ♂♂, 15. ♀,. 16-18. Teulisna varia n. sp., 16. ♂, HT, 17. ♂, PT, 18. ♀, PT. 19. Mithuna strigifera, ♂. 20-21. Macotasa nedoshivinae, ♂♂. 22. Dolgoma rectoides, ♂. 23-24. Teuloma nebulosa, 23. ♂, 24. ♀. 25. Gampola sinica, ♀. 26. Asiapistosia subnigra, ♂. 73 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 27-36. Adults. 27. Asiapistosia subnigra, ♀. 28-29. Thysanoptyx indosinica, ♂♂. 30. Brunia sarawaca, ♀. 31-32. Cernyia pseudocretacea, 31. ♂, 32. ♀. 33-34. Cernyia pseudobrevivalva, 33. ♂, 34. ♀. 35-36. Katha montana, ♂♂.

74 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 37-58. Adults. 37-38. Gandhara vietnamica, 37. ♂, 38. ♀. 39-40. Bucsekia mediumpilosa, 39. ♂, 40. ♀. 41-42. Microlithosia unicolora n. sp., 41. ♂, HT, 42. ♂, PT. 43. Microlithosia nanlingica, ♂. 44-46. Danielithosia zolotuhini, 44-45. ♂♂, 46. ♀. 47-50. Danielithosia dubatolovi n. sp., 47. ♂, HT, 48. ♂, PT, 49-50. ♀♀, PT. 51-52. Poliosia cubitifera, ♂♂. 53. Poliosia rectilinea, ♂. 54. Poliosia bilineata n. sp., ♂, HT. 55-57. Poliosia cardamomensis, 55-56. ♂♂, 57. ♀. 58. Veslema bipunctulata, ♀. 75 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 59-68. Adults. 59-60. Nishada flabrifera, 59. ♂, 60. ♀. 61. Agrisius leloii, ♀. 62. Vamuna remelana, ♂. 63. Macrobrochis fukiensis, ♂. 64. Churinga nigripes, ♂. 65. Chrysaeglia magnifica, ♂. 66. Utetheisa pulchelloides vaga, ♀. 67. Nyctemera adversata, ♂. 68. Nyctemera tripunctaria, ♂. 76 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 69-78. Adults. 69-70. Creatonotos transiens, 69. ♂, 70. ♀. 71-72. Creatonotos gangis, 71. ♂, 72. ♀. 73. Spilosoma nydia tienmushanica, ♂. 74-76. Spilosoma bisecta, 74-75. ♂♂, 76. ♀. 77-78. Spilosoma sagittifera, ♂♂. 77 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 79-89. Adults. 79. Spilosoma sagittifera, ♂. 80. Lemyra quadrisaccus, ♂. 81. Lemyra gloria, ♂. 82. Lemyra ?multivittata, ♂. 83. Lemyra ?crama, ♂. 84. Lemyra ?anormala, ♂. 85. Paraeuchaetes pseudoinsulata, ♂. 86. Calpenia saundersi, ♀. 87. Areas galactina inouei, ♂. 88-89. Areas aquila, 88. ♂, 89. ♀. Moths above the line has been enlarget more as moths below the line. 78 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 90-98. Genitalia. 90-91. Teulisna fimbriata, 90. ♂, 91. ♀. 92-93. Teulisna protuberans, 92. ♂, 93. ♀. 94. Teulisna tongdzuythanhi, ♂. 95-96. Teulisna mithunoides, 95. ♂, 96. ♀. 97-98. Teulisna varia n. sp., 97. ♂, HT, 98. ♀, PT. 79 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 99-107. Genitalia. 99. Macotasa nedoshivinae, ♂. 100. Dolgoma rectoides, ♂. 101. Teuloma nebulosa, ♂. 102. Gampola sinica, ♀. 103-104. Asiapistosia subnigra, 103. ♂, 104. ♀. 105. Thysanoptyx indosinica, ♂. 106. Brunia sarawaca, ♀. 107. Cernyia pseudocretacea, ♂.

80 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 108-115. Genitalia. 108. Cernyia pseudocretacea, ♀. 109-110. Cernyia pseudobrevivalva, 109. ♂, 110. ♀. 111. Katha montana, ♂. 112-113. Bucsekia mediumpilosa, 112. ♂, 113. ♀. 114. Microlithosia unicolora n. sp., ♂, HT. 115. Microlithosia nanlingica, ♂.

81 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 116-124. Genitalia. 116-118. Danielithosia zolotuhini, 116-117. ♂♂, 118. ♀. 119-120. Danielithosia dubatolovi n. sp., 119. ♂, HT, 120. ♀, PT. 121-122. Poliosia cubitifera, ♂♂. 123. Poliosia rectilinea, ♂. 124. Poliosia bilineata n. sp., ♂, HT. 82 Entomofauna carpathica, 2020, 32(2): 47-87

Figures 125-132. Genitalia. 125-126. Poliosia cardamomensis, 125. ♂, 126. ♀. 127. Nishada flabrifera, ♂. 128. Churinga nigripes, ♂. 129. Lemyra gloria, ♂. 130. Lemyra ?multivittata, ♂. 131. Lemyra ?crama, ♂. 132. Lemyra ?anormala, ♂. 83 Entomofauna carpathica, 2020, 32(2): 47-87

ACKNOWLEDGEMENTS

Thanks to Mr. Henrich Novák, Bratislava for translating the manuscript into English and Dr. Ladislav Roller, Slovak Academy of Sciences, Bratislava for English correction. I also express my thanks to Dr. Ľubomír Vidlička, Slovak Academy of Sciences, Bratislava for technical help during editing of the manuscript.

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