~ SNAKE PARK in NAMIBIA Emanuele Cimatti
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Freshwater Fishes
WESTERN CAPE PROVINCE state oF BIODIVERSITY 2007 TABLE OF CONTENTS Chapter 1 Introduction 2 Chapter 2 Methods 17 Chapter 3 Freshwater fishes 18 Chapter 4 Amphibians 36 Chapter 5 Reptiles 55 Chapter 6 Mammals 75 Chapter 7 Avifauna 89 Chapter 8 Flora & Vegetation 112 Chapter 9 Land and Protected Areas 139 Chapter 10 Status of River Health 159 Cover page photographs by Andrew Turner (CapeNature), Roger Bills (SAIAB) & Wicus Leeuwner. ISBN 978-0-620-39289-1 SCIENTIFIC SERVICES 2 Western Cape Province State of Biodiversity 2007 CHAPTER 1 INTRODUCTION Andrew Turner [email protected] 1 “We live at a historic moment, a time in which the world’s biological diversity is being rapidly destroyed. The present geological period has more species than any other, yet the current rate of extinction of species is greater now than at any time in the past. Ecosystems and communities are being degraded and destroyed, and species are being driven to extinction. The species that persist are losing genetic variation as the number of individuals in populations shrinks, unique populations and subspecies are destroyed, and remaining populations become increasingly isolated from one another. The cause of this loss of biological diversity at all levels is the range of human activity that alters and destroys natural habitats to suit human needs.” (Primack, 2002). CapeNature launched its State of Biodiversity Programme (SoBP) to assess and monitor the state of biodiversity in the Western Cape in 1999. This programme delivered its first report in 2002 and these reports are updated every five years. The current report (2007) reports on the changes to the state of vertebrate biodiversity and land under conservation usage. -
A Molecular Phylogeny of the Lamprophiidae Fitzinger (Serpentes, Caenophidia)
Zootaxa 1945: 51–66 (2008) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA Copyright © 2008 · Magnolia Press ISSN 1175-5334 (online edition) Dissecting the major African snake radiation: a molecular phylogeny of the Lamprophiidae Fitzinger (Serpentes, Caenophidia) NICOLAS VIDAL1,10, WILLIAM R. BRANCH2, OLIVIER S.G. PAUWELS3,4, S. BLAIR HEDGES5, DONALD G. BROADLEY6, MICHAEL WINK7, CORINNE CRUAUD8, ULRICH JOGER9 & ZOLTÁN TAMÁS NAGY3 1UMR 7138, Systématique, Evolution, Adaptation, Département Systématique et Evolution, C. P. 26, Muséum National d’Histoire Naturelle, 43 Rue Cuvier, Paris 75005, France. E-mail: [email protected] 2Bayworld, P.O. Box 13147, Humewood 6013, South Africa. E-mail: [email protected] 3 Royal Belgian Institute of Natural Sciences, Rue Vautier 29, B-1000 Brussels, Belgium. E-mail: [email protected], [email protected] 4Smithsonian Institution, Center for Conservation Education and Sustainability, B.P. 48, Gamba, Gabon. 5Department of Biology, 208 Mueller Laboratory, Pennsylvania State University, University Park, PA 16802-5301 USA. E-mail: [email protected] 6Biodiversity Foundation for Africa, P.O. Box FM 730, Bulawayo, Zimbabwe. E-mail: [email protected] 7 Institute of Pharmacy and Molecular Biotechnology, University of Heidelberg, INF 364, D-69120 Heidelberg, Germany. E-mail: [email protected] 8Centre national de séquençage, Genoscope, 2 rue Gaston-Crémieux, CP5706, 91057 Evry cedex, France. E-mail: www.genoscope.fr 9Staatliches Naturhistorisches Museum, Pockelsstr. 10, 38106 Braunschweig, Germany. E-mail: [email protected] 10Corresponding author Abstract The Elapoidea includes the Elapidae and a large (~60 genera, 280 sp.) and mostly African (including Madagascar) radia- tion termed Lamprophiidae by Vidal et al. -
Nyika and Vwaza Reptiles & Amphibians Checklist
LIST OF REPTILES AND AMPHIBIANS OF NYIKA NATIONAL PARK AND VWAZA MARSH WILDLIFE RESERVE This checklist of all reptile and amphibian species recorded from the Nyika National Park and immediate surrounds (both in Malawi and Zambia) and from the Vwaza Marsh Wildlife Reserve was compiled by Dr Donald Broadley of the Natural History Museum of Zimbabwe in Bulawayo, Zimbabwe, in November 2013. It is arranged in zoological order by scientific name; common names are given in brackets. The notes indicate where are the records are from. Endemic species (that is species only known from this area) are indicated by an E before the scientific name. Further details of names and the sources of the records are available on request from the Nyika Vwaza Trust Secretariat. REPTILES TORTOISES & TERRAPINS Family Pelomedusidae Pelusios rhodesianus (Variable Hinged Terrapin) Vwaza LIZARDS Family Agamidae Acanthocercus branchi (Branch's Tree Agama) Nyika Agama kirkii kirkii (Kirk's Rock Agama) Vwaza Agama armata (Eastern Spiny Agama) Nyika Family Chamaeleonidae Rhampholeon nchisiensis (Nchisi Pygmy Chameleon) Nyika Chamaeleo dilepis (Common Flap-necked Chameleon) Nyika(Nchenachena), Vwaza Trioceros goetzei nyikae (Nyika Whistling Chameleon) Nyika(Nchenachena) Trioceros incornutus (Ukinga Hornless Chameleon) Nyika Family Gekkonidae Lygodactylus angularis (Angle-throated Dwarf Gecko) Nyika Lygodactylus capensis (Cape Dwarf Gecko) Nyika(Nchenachena), Vwaza Hemidactylus mabouia (Tropical House Gecko) Nyika Family Scincidae Trachylepis varia (Variable Skink) Nyika, -
"Herpetofauna of Oak Ridge Area."
.p o RNC.<,7 .Jiiiiiire . , .. - ' [3.., *i . '& " J .) 7/C .f q }g Q - i '" - . - y' o - - ___ ORNL.3653. 't c |;,, UC.48 - Bielegy and Me ' ' TID.4500 (34th ed. REFEREflCE 2-35 6 I . t ! - . .t * , ,e . !. THE HERPETOFAUNA OF THE CAK RIDGE AREA ~}^i.'- i ti R. M. Jchnson s ' , , p ?.... , . - e . 'a l .a: . *)f* J*.*tf t if 9 . ' .85 - . p* e ' W C&fi . %' g. ey cdev ; : , , %$ 9pto ur .s s <- o .y-=- s%,<=. o ;u n ___ II G.k ' e .a wt . i, ;j f'$~MS):if%--~L.$, ;A s o1.. f =;'Q::.+Eli(. t - ]s- < a ~ . n= ~ h . 9:!;;s-rf C L. %- s. ,, % ~- ~, ~g. ;"3 - a - l " )< ,\.:-m5W?.;; ..- ! ,- n _ .' | H o r - %P, O AK RIDGE N AT1014 AL L ABOR ATORY .Vj '.g t(e,C ' .hl t ENERGT cperated by ( ' (Er. i, | [.*jt" .ti UNION C ARBIDE CORPOR ATION i f or the | -$N,. ) | I U.S. ATGMIC E N E RGY C O.V. MISSION ' - . ~:.* 4 .h' . tih- > . - '8201060379 811231 | PDR ADGCK 05000 G . ' ' " ' ' ' e __ _ _ . .s- * e * . * .. , . 0 .- . e . * O . e - e . e e . i I I e } ' Petened in USA. Peise 12.00. A.e.lebte fee es e Close.agne se fee Fedwet c.on, he..eael Seceee of 1seadeeds, l [ Sciene fie ead Teebe.s et tale I * U.5 2eere.... el Co...eee, Se..a s t.eid. v.eg.eie . LEGAL MOTICE 4 No.ehee ene Va.eed $ ewes, ' - TMe eeene .es e.eewed es e. ess eeae e8 Co...--iew eseaneced .we. -
Ancestral Reconstruction of Diet and Fang Condition in the Lamprophiidae: Implications for the Evolution of Venom Systems in Snakes
Journal of Herpetology, Vol. 55, No. 1, 1–10, 2021 Copyright 2021 Society for the Study of Amphibians and Reptiles Ancestral Reconstruction of Diet and Fang Condition in the Lamprophiidae: Implications for the Evolution of Venom Systems in Snakes 1,2 1 1 HIRAL NAIK, MIMMIE M. KGADITSE, AND GRAHAM J. ALEXANDER 1School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Johannesburg. PO Wits, 2050, Gauteng, South Africa ABSTRACT.—The Colubroidea includes all venomous and some nonvenomous snakes, many of which have extraordinary dental morphology and functional capabilities. It has been proposed that the ancestral condition of the Colubroidea is venomous with tubular fangs. The venom system includes the production of venomous secretions by labial glands in the mouth and usually includes fangs for effective delivery of venom. Despite significant research on the evolution of the venom system in snakes, limited research exists on the driving forces for different fang and dental morphology at a broader phylogenetic scale. We assessed the patterns of fang and dental condition in the Lamprophiidae, a speciose family of advanced snakes within the Colubroidea, and we related fang and dental condition to diet. The Lamprophiidae is the only snake family that includes front-fanged, rear-fanged, and fangless species. We produced an ancestral reconstruction for the family and investigated the pattern of diet and fangs within the clade. We concluded that the ancestral lamprophiid was most likely rear-fanged and that the shift in dental morphology was associated with changes in diet. This pattern indicates that fang loss, and probably venom loss, has occurred multiple times within the Lamprophiidae. -
Annotated Checklist and Provisional Conservation Status of Namibian Reptiles
Annotated Checklist - Reptiles Page 1 ANNOTATED CHECKLIST AND PROVISIONAL CONSERVATION STATUS OF NAMIBIAN REPTILES MICHAEL GRIFFIN BIODIVERSITY INVENTORY MINISTRY OF ENVIRONMENT AND TOURISM PRIVATE BAG 13306 WINDHOEK NAMIBIA Annotated Checklist - Reptiles Page 2 Annotated Checklist - Reptiles Page 3 CONTENTS PAGE ABSTRACT 5 INTRODUCTION 5 METHODS AND DEFINITIONS 6 SPECIES ACCOUNTS Genus Crocodylus Nile Crocodile 11 Pelomedusa Helmeted Terrapin 11 Pelusios Hinged Terrapins 12 Geochelone Leopard Tortoise 13 Chersina Bowsprit Tortoise 14 Homopus Nama Padloper 14 Psammobates Tent Tortoises 15 Kinixys Hinged Tortoises 16 Chelonia GreenTurtle 16 Lepidochelys Olive Ridley Turtle 17 Dermochelys Leatherback Turtle 17 Trionyx African Soft-shelled Turtle 18 Afroedura Flat Geckos 19 Goggia Dwarf Leaf-toed Geckos 20 Afrogecko Marbled Leaf-toed Gecko 21 Phelsuma Namaqua Day Gecko 22 Lygodactylus Dwarf Geckos 23 Rhoptropus Namib Day Geckos 25 Chondrodactylus Giant Ground Gecko 27 Colopus Kalahari Ground Gecko 28 Palmatogecko Web-footed Geckos 28 Pachydactylus Thick-toed Geckos 29 Ptenopus Barking Geckos 39 Narudasia Festive Gecko 41 Hemidactylus Tropical House Geckos 41 Agama Ground Agamas 42 Acanthocercus Tree Agama 45 Bradypodion Dwarf Chameleons 46 Chamaeleo Chameleons 47 Acontias Legless Skinks 48 Typhlosaurus Blind Legless Skinks 48 Sepsina Burrowing Skinks 50 Scelotes Namibian Dwarf Burrowing Skink 51 Typhlacontias Western Burrowing Skinks 51 Lygosoma Sundevall’s Writhing Skink 53 Mabuya Typical Skinks 53 Panaspis Snake-eyed Skinks 60 Annotated -
SQUAMATA: Vlperldae Crotalus Ruber
840.1 REPTILIA: SQUAMATA: VlPERlDAE Crotalus ruber Catalogue of American Amphibians and Reptiles. Beaman, K.R. and E.A. Dugan. 2006. Crotalus ruber. Crotalus ruber Cope Red Diamond Rattlesnake Crotalus adamanteus atrox: Cope 1875:33 (part). Crotalus exsul: Garman 1884 [dated 1883]:114. Crotalus adamanteus ruber Cope 1892:690. Type- locality not given, restricted to "Dulzura, San Diego County, California," by Smith and Taylor (1950). Holotype, National Museum of Natural History (USNM) 9209, adult male, collector and date of collection not given (not examined by authors). Crotalus atrox ruber: Stejneger 1895:439. Crotalus rube^ Van Denburgh 1896: 1007. Figure 1. Crotalus ruberfrom Chino Hills State Park, Crotalus lucasensis Van Denburgh 1920:29. Type- San Bernardino County, California, USA (above); locality, "Agua Caliente, Cape Region of Lower Crotalus ruberfrom the Jacumba Mountains, Imperial [Baja] California, Mexico." Holotype, California County, California, USA (below). Photographs by Academy of Sciences (CAS) 45888, adult male, E.A. Dugan. collected by J.R. Slevin on 26 July 1919. Crotalus atrox lucasensis Schmidt 1922:698. Crotalus atrox elegans Schmidt 1922:699. Type- locality, "Angel de la Guardia Island, Gulf of Cali- fornia" [Mexico]. Holotype, National Museum of Natural History (USNM) 64452, age and sex not given, collected by C.H. Townsend on 10 April 1911. Crotalus exsul ruber: Kallert 1927:372. Crotalus ruber ruber: Klauber 194959. Crotalus ruber lucasensis Klauber 194959. Crotalus ruber lorenzoensis Radcliffe and Maslin 1975:490. Type-locality, "San Lorenzo Sur Island in the Gulf of California, Baja California Norte, Mexico." Holotype, San Diego Society of Natural History (SDSNH) 46009, adult male, collected by C.E. -
Amphibian and Reptile Checklist
KEY TO ABBREVIATIONS ___ Eastern Rat Snake (Pantherophis alleghaniensis) – BLUE RIDGE (NC‐C, VA‐C) Habitat: Varies from rocky timbered hillsides to flat farmland. The following codes refer to an animal’s abundance in ___ Eastern Hog‐nosed Snake (Heterodon platirhinos) – PARKWAY suitable habitat along the parkway, not the likelihood of (NC‐R, VA‐U) Habitat: Sandy or friable loam soil seeing it. Information on the abundance of each species habitats at lower elevation. AMPHIBIAN & comes from wildlife sightings reported by park staff and ___ Eastern Kingsnake (Lampropeltis getula) – (NC‐U, visitors, from other agencies, and from park research VA‐U) Habitat: Generalist at low elevations. reports. ___ Northern Mole Snake (Lampropeltis calligaster REPTILE C – COMMON rhombomaculata) – (VA‐R) Habitat: Mixed pine U – UNCOMMON forests and open fields under logs or boards. CHECKLIST R – RARE ___ Eastern Milk Snake (Lampropeltis triangulum) – (NC‐ U, VA‐U) Habitat: Woodlands, grassy balds, and * – LISTED – Any species federally or state listed as meadows. Endangered, Threatened, or of Special Concern. ___ Northern Water Snake (Nerodia sipedon sipedon) – (NC‐C, VA‐C) Habitat: Wetlands, streams, and lakes. Non‐native – species not historically present on the ___ Rough Green Snake (Opheodrys aestivus) – (NC‐R, parkway that have been introduced (usually by humans.) VA‐R) Habitat: Low elevation forests. ___ Smooth Green Snake (Opheodrys vernalis) – (VA‐R) NC – NORTH CAROLINA Habitat: Moist, open woodlands or herbaceous Blue Ridge Red Cope's Gray wetlands under fallen debris. Salamander Treefrog VA – VIRGINIA ___ Northern Pine Snake (Pituophis melanoleucus melanoleucus) – (VA‐R) Habitat: Abandoned fields If you see anything unusual while on the parkway, please and dry mountain ridges with sandier soils. -
Part A: Details of Applicant
ANNEXURE 1 FORMS Form 1 REPUBLIC OF NAMIBIA ENVIRONMENTAL MANAGEMENT ACT, 2007 (Section 32) APPLICATION FOR ENVIRONMENTAL CLEARANCE CERTIFICATE Revenue stamp Revenue stamp Revenue stamp or revenue or revenue or revenue franking franking franking machine machine machine impression impression impression PART A: DETAILS OF APPLICANT Name :(person or business) : Pure Tantalum (Pty) L td Business Registration/Identity No . (if applicable) Correspondence Address : P.O. Box 68 Uis Namibia Name of Contact Person: Pieter. H Position of Contact Person: Telephone No: +2 64 61 402246 Fax No: E - mail Address: [email protected] Tick the appropriate box PART B. SCOPE OF THE ENVIRONMENTAL CLEARANCE CERTIFICATE 1. The environmental clearance certificate is for: T he exploration activities on exclusive prospecting license ( EPL ) 7004 in Dorob National Park, Erongo region. 2. Details of the activity(s) covered by the environmental clearance certificate : [Note: Please attach plans to show the location and scope of the designated activity(s ), and use additional sheets if ne cessary: Title of Activity: E xploration activities on exclusive pr ospecting license (EPL) 7004 in Dorob National Park, Erongo region Nature of Activity : T he exploration activities on exclusive prospecting license (EPL) 7004 in Dorob National Park, Erongo region. The project activities will involve mapping , drilling and sampling of Base and Rare metals, 1. Name of current holder of Environmental Clearance Certificate: Industrial Minerals , Non Nuclear Fuel Minerals and Precious metals. Location of Activity: EPL 7004 in Dorob National Park , Erongo Region Scale and Scope of Activity: Exploration activities of Base and Rare Metals, Industrial Minerals, Non - Nuclear Fuel Minerals and Precious Metals on EPL 7004, area size of 1796.3841 Hectares . -
Australasian Journal of Herpetology ISSN 1836-5698 (Print)1 Issue 12, 30 April 2012 ISSN 1836-5779 (Online) Australasian Journal of Herpetology
Australasian Journal of Herpetology ISSN 1836-5698 (Print)1 Issue 12, 30 April 2012 ISSN 1836-5779 (Online) Australasian Journal of Herpetology Hoser 2012 - Australasian Journal of Herpetology 9:1-64. Available online at www.herp.net Contents on pageCopyright- 2. Kotabi Publishing - All rights reserved 2 Australasian Journal of Herpetology Issue 12, 30 April 2012 Australasian Journal of Herpetology CONTENTS ISSN 1836-5698 (Print) ISSN 1836-5779 (Online) A New Genus of Coral Snake from Japan (Serpentes:Elapidae). Raymond T. Hoser, 3-5. A revision of the Asian Pitvipers, referred to the genus Cryptelytrops Cope, 1860, with the creation of a new genus Adelynhoserea to accommodate six divergent species (Serpentes:Viperidae:Crotalinae). Raymond T. Hoser, 6-8. A division of the South-east Asian Ratsnake genus Coelognathus (Serpentes: Colubridae). Raymond T. Hoser, 9-11. A new genus of Asian Snail-eating Snake (Serpentes:Pareatidae). Raymond T. Hoser, 10-12-15. The dissolution of the genus Rhadinophis Vogt, 1922 (Sepentes:Colubrinae). Raymond T. Hoser, 16-17. Three new species of Stegonotus from New Guinea (Serpentes: Colubridae). Raymond T. Hoser, 18-22. A new genus and new subgenus of snakes from the South African region (Serpentes: Colubridae). Raymond T. Hoser, 23-25. A division of the African Genus Psammophis Boie, 1825 into 4 genera and four further subgenera (Serpentes: Psammophiinae). Raymond T. Hoser, 26-31. A division of the African Tree Viper genus Atheris Cope, 1860 into four subgenera (Serpentes:Viperidae). Raymond T. Hoser, 32-35. A new Subgenus of Giant Snakes (Anaconda) from South America (Serpentes: Boidae). Raymond T. Hoser, 36-39. -
A Phylogeny and Revised Classification of Squamata, Including 4161 Species of Lizards and Snakes
BMC Evolutionary Biology This Provisional PDF corresponds to the article as it appeared upon acceptance. Fully formatted PDF and full text (HTML) versions will be made available soon. A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes BMC Evolutionary Biology 2013, 13:93 doi:10.1186/1471-2148-13-93 Robert Alexander Pyron ([email protected]) Frank T Burbrink ([email protected]) John J Wiens ([email protected]) ISSN 1471-2148 Article type Research article Submission date 30 January 2013 Acceptance date 19 March 2013 Publication date 29 April 2013 Article URL http://www.biomedcentral.com/1471-2148/13/93 Like all articles in BMC journals, this peer-reviewed article can be downloaded, printed and distributed freely for any purposes (see copyright notice below). Articles in BMC journals are listed in PubMed and archived at PubMed Central. For information about publishing your research in BMC journals or any BioMed Central journal, go to http://www.biomedcentral.com/info/authors/ © 2013 Pyron et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/2.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited. A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes Robert Alexander Pyron 1* * Corresponding author Email: [email protected] Frank T Burbrink 2,3 Email: [email protected] John J Wiens 4 Email: [email protected] 1 Department of Biological Sciences, The George Washington University, 2023 G St. -
Patterns of Species Richness, Endemism and Environmental Gradients of African Reptiles
Journal of Biogeography (J. Biogeogr.) (2016) ORIGINAL Patterns of species richness, endemism ARTICLE and environmental gradients of African reptiles Amir Lewin1*, Anat Feldman1, Aaron M. Bauer2, Jonathan Belmaker1, Donald G. Broadley3†, Laurent Chirio4, Yuval Itescu1, Matthew LeBreton5, Erez Maza1, Danny Meirte6, Zoltan T. Nagy7, Maria Novosolov1, Uri Roll8, 1 9 1 1 Oliver Tallowin , Jean-Francßois Trape , Enav Vidan and Shai Meiri 1Department of Zoology, Tel Aviv University, ABSTRACT 6997801 Tel Aviv, Israel, 2Department of Aim To map and assess the richness patterns of reptiles (and included groups: Biology, Villanova University, Villanova PA 3 amphisbaenians, crocodiles, lizards, snakes and turtles) in Africa, quantify the 19085, USA, Natural History Museum of Zimbabwe, PO Box 240, Bulawayo, overlap in species richness of reptiles (and included groups) with the other ter- Zimbabwe, 4Museum National d’Histoire restrial vertebrate classes, investigate the environmental correlates underlying Naturelle, Department Systematique et these patterns, and evaluate the role of range size on richness patterns. Evolution (Reptiles), ISYEB (Institut Location Africa. Systematique, Evolution, Biodiversite, UMR 7205 CNRS/EPHE/MNHN), Paris, France, Methods We assembled a data set of distributions of all African reptile spe- 5Mosaic, (Environment, Health, Data, cies. We tested the spatial congruence of reptile richness with that of amphib- Technology), BP 35322 Yaounde, Cameroon, ians, birds and mammals. We further tested the relative importance of 6Department of African Biology, Royal temperature, precipitation, elevation range and net primary productivity for Museum for Central Africa, 3080 Tervuren, species richness over two spatial scales (ecoregions and 1° grids). We arranged Belgium, 7Royal Belgian Institute of Natural reptile and vertebrate groups into range-size quartiles in order to evaluate the Sciences, OD Taxonomy and Phylogeny, role of range size in producing richness patterns.