Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009

Report Citation: Parkinson, L., S.A. Blanchette, J. Heron. 2009. Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009. B.C. Ministry of Environment, Ecosystems Branch, Wildlife Science Section, Vancouver, B.C. 51 pp.

Cover illustration: Euphyes vestris, taken 2007, lower Fraser Valley, photo by Denis Knopp.

Photographs may be used without permission for non-monetary and educational purposes, with credit to this report and photographer as the source. The cover photograph is credited to Denis Knopp.

Contact Information for report: Jennifer Heron, Invertebrate Specialist, B.C. Ministry of Environment, Ecosystems Branch, Wildlife Science Section, 316 – 2202 Main Mall, Vancouver, B.C., Canada, V6T 1Z1. Phone: 604-222-6759. Email: [email protected]

Acknowledgements

Fieldwork was conducted by Laura Parkinson and Sophie-Anne Blanchette, B.C. Conservation Corps Invertebrate Species at Risk Crew. Jennifer Heron (B.C. Ministry of Environment) provided maps, planning and guidance for this project.

The B.C. Invertebrate Species at Risk Inventory project was administered by the British Columbia Conservation Foundation (Joanne Neilson). Funding was provided by the B.C. Ministry of Environment through the B.C. Conservation Corp program (Ben Finkelstein, Manager and Bianka Sawicz, Program Coordinator), the B.C. Ministry of Environment Wildlife Science Section (Alec Dale, Manager) and Conservation Framework Funding (James Quayle, Manager). Joanne Neilson (B.C. Conservation Foundation) was a tremendous support to this project. This project links with concurrent invertebrate stewardship projects funded by the federal Habitat Stewardship Program for species at risk.

B.C. Conservation Data Centre provided previous records and location information for Dun Skipper. Special thanks to Denis Knopp who provided background information on Dun Skipper, recommendations for survey sites and habitats in the Harrison Lake area. Special thanks to Crispin Guppy who provided help with identification of specimens, photographs and further insights into habitat for various butterfly species recorded during this survey.

Thanks to everyone who identified specimens collected opportunistically. Rob Cannings (Royal British Columbia Museum) identified terrestrial insects collected by hand and Rex Kenner (Beaty Biodiversity Museum Spencer Entomological Collection) identified aquatic insects. Kristiina Ovaska and Lennart Sopuck identified aquatic molluscs. Karen Needham (Beaty Biodiversity Museum Spencer Entomological Collection) helped with resources from the entomological museum.

Survey information from this project was sent to the B.C. Conservation Data Centre and the Wildlife Species Inventory database. Butterfly specimens collected were deposited at the Beaty Biodiversity Museum Spencer Entomological Collection (Vancouver) (Karen Needham, Curator) and at the Royal British Columbia Museum (Victoria) (Claudia Copley, Collections Manager).

Photographs were taken by Sophie-Anne Blanchette and Laura Parkinson. Appropriate credit is next to each photograph. Denis Knopp is credited with the cover photograph.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 iv

Executive Summary

Dun Skipper (Euphyes vestris) is a widespread species in North America, but the western population (E.v. vestris) is scattered, disjunct and considered vulnerable throughout its range in British Columbia and the states of Washington, Oregon and California. In British Columbia, Dun Skipper is blue listed due to limited records and anthropogenic changes to its habitat (B.C. Conservation Data Center, 2009; Shepard, 2000). Previous records in B.C include southern Vancouver Island, Hornby Island, Salt Spring Island, Denman Island, throughout the lower Fraser River area from Hope to Lillooet, and three isolated sites in Pemberton, Powell River and Mission.

This survey was initiated to survey for additional occurrences of Dun Skipper in the Harrison Lake area. Nine days of surveys targeting Dun Skipper habitats were completed during July 2009. The survey areas focused on crown land accessible via forest service roads on the east and west sides of Harrison Lake. Habitats targeted for surveys included regenerating wet clear cuts and natural open areas where the host plant, grasses and sedges of Carex and Cyperus species could potentially occur. All butterfly observations were recorded.

Twelve butterfly species were observed during this survey. Neither Dun Skipper butterflies nor species of conservation concern were recorded. Two introduced butterfly species were observed, the Cabbage White (Pieris rapae) and the European Skipper (Thymelicus lineola).

Future inventory should focus on areas south of the Fraser in the lower mainland, Mission, Burns Bog (Delta), and southeastern Vancouver Island and the Gulf Islands.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 v

Table of Contents

Acknowledgements ...... iv

Executive Summary ...... v

Introduction ...... 10

Background Information ...... 12

Survey Area ...... 12

Summary of Existing Information ...... 16

Methods ...... 17 Habitat Characteristics for Dun Skipper ...... 17 Field surveys ...... 17 Butterflies ...... 17 Aquatic Samples ...... 18 Hand searching for other terrestrial insects ...... 18

Results ...... 19

Other Butterfly Observations ...... 20 Mylitta Crescent (Phyciodes mylitta) ...... 21 Clodius Parnassian (Parnassius clodius) ...... 21 Cabbage White (Pieris rapae) ...... 22 Western Tiger Swallowtail (Papilio rutulus) ...... 22 Lorquin’s Admiral (Limenitus lorquini) ...... 23 Pale Swallowtail (Papilio eurymedon) ...... 23 Hydaspe Fritillary (Speyeria hydaspe) ...... 24 California Tortoiseshell (Nymphalis californica) ...... 25 Pine White (Neophasia menapia) ...... 26 Northern Blue (Plebejus idas) ...... 26 European Skipper (Thymelicus lineola) ...... 26 Purplish Copper (Lycaena helloides) ...... 27 Western Spring Azure (Celestrina echo) ...... 27

Other Invertebrate Observations ...... 28 Aquatic Samples ...... 28 Hand Collected Terrestrial Insects ...... 28

Discussion ...... 30

Recommendations for future surveys ...... 30

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 vi

References ...... 31

Appendix ...... 32

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 vii

List of Tables

Table 1: Aquatic sampling sites in the Harrison Lake area ...... 15 Table 2: Summary table of butterfly species observed in Harrison Lake Surveys (*introduced species; **species of conservation concern) ...... 19 Table 3: Summary of observations of the five most commonly recorded butterfly species seen in the Harrison Lake area, July 2009 ...... 20 Table 4: Aquatic molluscs collected opportunistically from roadside lakes and marshes near Harrison Lake, as identified by Kristiina Ovaska and Lennart Sopuck ...... 28 Table 5: Terrestrial insects collected opportunistically in the Harrison Lake area, as identified by Rob Cannings ...... 29

List of Figures and Photographs

Figure 1: Generalized locations for Dun Skipper in B.C.(B. C. Conservation Data Centre, 2009)...... 11 Figure 2: Transects surveyed for Dun Skipper in Harrison Lake area July 2009. Each colour represents a different survey day...... 13 Figure 3: Photo of a recent clear cut on the West side of Harrison Lake, July 17 2009. Photo by Laura Parkinson...... 14 Figure 4: Photo of Tolson’s FSR, July 19 2009. Photo by Laura Parkinson...... 14 Figure 5: Photo of overgrown unmarked road branching off of South Talc FSR into regenerating clear cut/meadow habitat, July 22 2009. Photo by Laura Parkinson .. 14 Figure 6: Photo of Slollicum FSR taken on July 17 2009. Photo by Sophie-Anne Blanchette...... 14 Figure 7: Photos of unnamed marshy area adjacent to Harrison Lake FSR taken on July 5 2009. Photos by Laura Parkinson...... 15 Figure 8: Photo of Wood Lake adjacent to West Harrison FSR where aquatic samples were conducted taken on July 6 2009. Photo by Sophie-Anne Blanchette...... 15 Figure 9: Photo of unnamed swamp adjacent to West Harrison FSR where aquatic samples were conducted taken on July 6 2009. Photo by Sophie-Anne Blanchette...... 15 Figure 10: Photo of Grace Lake adjacent to West Harrison FSR where aquatic samples were conducted taken on July 6 2009. Photo by Laura Parkinson...... 16 Figure 11: Photo of Wolf Lake adjacent to West Harrison FSR where aquatic samples were conducted taken on July 6 2009. Photo by Laura Parkinson...... 16 Figure 12: Photos of the dorsal and ventral views of a Clodius Parnassian taken on an unmarked road branching off of Chehalis FSR on July 5th 2009. Photos by Laura Parkinson...... 22 Figure 13: Photo of Lorquin’s admiral taken on North Statlu FSR, July 4 2009. Photo by Sophie-Anne Blanchette...... 23 Figure 14: Photo of a small stream running across Bear Creek FSR near where Hydaspe Fritillary was observed taken on July 18 2009. Photo by Laura Parkinson...... 24 Figure 15: Dorsal and ventral view of a Hydaspe Fritillary observed on Bear Creek FSR taken on July 18 2009. Photos by Sophie-Anne Blanchette...... 25

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 viii

Figure 16: California tortoiseshell taken on North Statlu Creek FSR on July 4th 2009. Photo by Laura Parkinson...... 25 Figure 17: Photos of an Northern Blue observed on Tolson’s FSR on July 19th 2009. Photo by Laura Parkinson...... 26 Figure 18: Purplish Copper observed on Bear Creek FSR on July 18 2009. Photo by Sophie-Anne Blanchette...... 27

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 ix

Introduction

Introduction

Two subspecies of Dun Skipper are found in Canada: the western population (E. v. vestris) and the eastern population (E. v. metacomet). The eastern population occurs in central and eastern Canada, from Saskatchewan to Nova Scotia and south to Arizona and Florida in the United States, (Guppy and Shepard, 2001), where this subspecies is common and widespread. The western population is the only subspecies of Dun Skipper that occurs in B.C. Until as recently as the late 1990s B.C. populations of Dun Skipper were not recognized as distinct from the eastern population of Dun Skipper, and all Dun Skipper in Canada were classified as E. v. metacomet (Shepard, 2000).

Dun Skipper, western population, is found from southern British Columbia through western Washington and Oregon as far south as central California. Despite being widespread, populations of this subspecies are disjunct and scattered and the butterfly is considered vulnerable throughout most of its provincial range. Dun Skipper is blue- listed (special concern) and S3 (vulnerable) in B.C., due to the low number of occurrences throughout the province and threats to its habitat (B. C. Conservation Data Center, 2009). Federally, it is listed as threatened by the Committee on the Status of Endangered Wildlife in Canada (COSEWIC) and by the species at Risk Act (SARA, Schedule 1, Appendix 1). Globally Dun Skipper has a conservation status rank of G5T4 (species is widespread and secure, but that the subspecies E. v. vestris is widespread but uncommon) (Natureserve, 2009).

In British Columbia, Dun Skipper records are from Vancouver Island (greater Victoria area northward to Port Alberni), on the Gulf Islands, as well as throughout the Lower Mainland from Metro Vancouver east towards the lower Fraser Valley to Lillooet and Pemberton (Figure 2) (B. C. Conservation Data Center, 2009). The first record for this species in B.C. dates back to 1902 in Goldstream on Vancouver Island (Shepard, 2000).

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 10

Introduction

Figure 1: Generalized locations for Dun Skipper in B.C.(B. C. Conservation Data Centre, 2009).

The B.C. Conservation Corps Invertebrate Species at Risk Crew conducted surveys for Dun Skipper in the Harrison Lake area July 3-5 and July 17-22 2009. This survey aims to fill some of the knowledge gaps around distribution of the species in B.C. in the Lower Mainland.

The objectives of this survey are to: 1) Fill some of the knowledge gaps around distribution of Dun Skipper in the lower Fraser Valley. 2) Survey potentially suitable habitats for Dun Skipper on the borders of the species known range in southwestern B.C. 3) Gain information on the habitat preferences and life history of Dun Skipper Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 11

Background Information

4) Survey opportunistically for other terrestrial and aquatic invertebrates

Background Information

Dun Skipper is a small chocolate brown butterfly with a wingspan of 25-30 mm. The male Dun Skipper has a black stigmal patch on the dorsal forewing which may be surrounded by a slight tan flush. The female has a few white spots on the dorsal forewing which can be very inconspicuous and a pale purple crescent on the ventral hindwing (Guppy and Shepard, 2001). It is one of the few uniformly dark brown skippers in B.C. and can only be confused with the superficially similar tawny-edged skipper (Polites themistocles) and the common roadside skipper (Amblyscirtes vialis) (Acorn and Sheldon, 2006).

Dun Skipper flies from mid-May through to mid-August in one single brood. The females lay their pale green eggs at the base of sedge or grass host plant. After a few months, the pale green larvae hatches and forms a larval refuge site. This small silk chamber will be used throughout the larvae’s developmental stage as a protective refuge between feeding sessions (Guppy and Shepard, 2001). The exact length of the larval development in B.C. is unknown but based on observations from other parts of its range it is suspected to between 24 to 36 days (Guppy and Shepard, 2001).

The species of host plant used by Dun Skipper in B.C. is currently unknown. In Eastern North America the eastern subspecies of Dun Skipper (E. v. metacomet) has been known to use Cyperus esculentus and several Carex spp. and Poa spp. as host plants (Knopp et al., 2008). The Carex spp. documented as host plants for Dun Skipper in eastern North America do not occur in British Columbia, although different Carex species are available in B.C. Several species of Cyperus, including C. esculentus also occur in B.C. These genera of grasses and sedges are typically associated with open or edge habitats that may be moist to dry and are often disturbed, such as roadside ditches (Pojar and McKinnon, 1994).

Presence of a host plant may be a factor limiting the distribution of the western population of Dun Skipper, as this butterfly may exhibit varying host plant specificity in different habitats and locations (SHEPARD, 2000).

Survey Area

Surveys for Dun Skipper were conducted on the west side and east sides of Harrison Lake (Figure 2), on crown land accessible by active and deactivated forest service roads (FSRs).

There is a long history of forestry and resource extraction within the Harrison Lake area. Recent and historic forest harvesting within the Harrison Lake area has resulted in a patchy landscape composed of different forest ages and tree species composition.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 12

Survey Area

Recent clear cuts (less than 1 year; Figure 4) were avoided because they represented generally poor habitat for butterflies (i.e. little to no flowering vegetation growth, high winds, and dry conditions thus causing premature plant senescence).

Harrison Lake is a very high-use recreational area, particularly for the purposes of camping and off-road vehicle use, which have contributed to the highly disturbed ecosystems on both sides of the lake. Main roads with high vehicle use were not selected for surveys due to herbicide application (signs were posted where recent herbicide application had been completed) and high level of disturbance to nearby vegetation.

Figure 2: Transects surveyed for Dun Skipper in Harrison Lake area July 2009. Each colour represents a different survey day.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 13

Survey Area

Figure 3: Photo of a recent clear cut on the West side of Figure 4: Photo of Tolson’s FSR, July 19 2009. Harrison Lake, July 17 2009. Photo by Laura Parkinson. Photo by Laura Parkinson.

Figure 5: Photo of overgrown unmarked road branching Figure 6: Photo of Slollicum FSR taken on July off of South Talc FSR into regenerating clear 17 2009. Photo by Sophie-Anne Blanchette. cut/meadow habitat, July 22 2009. Photo by Laura Parkinson

Aquatic Surveys

Aquatic surveys were conducted opportunistically in rainy or cold and overcast weather inappropriate for butterfly surveys on July 5 and July 6 2009. On July 5 surveys were conducted in a small, unnamed marsh adjacent to Harrison Lake FSR on the east side of Harrison Lake (Figure 7). On July 6, surveys were conducted in Wood (Figure 8), Grace (Figure 10) and Wolf Lakes (Figure 11), all accessible off of West Harrison FSR. All of these lakes are adjacent to high use forest recreation sites, and as a result were relatively disturbed. A small unnamed roadside swampy area was also sampled adjacent to West Harrison FSR on July 6 (Figure 9). See table below for UTM locations of wetlands sampled.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 14

Survey Area

Table 1: Aquatic sampling sites in the Harrison Lake area UTM zone; 10 NAD 83 Survey Site Name Date Easting Northing Length (min) Unnamed marsh July 5 2009 584213 5470195 60 Wood Lake July 6 2009 582929 5478967 120 Unnamed July 6 2009 584497 5465733 30 swamp Grace Lake July 6 2009 582762 5464448 30 Wolf Lake July 6 2009 582479 5464983 65

Figure 7: Photos of unnamed marshy area adjacent to Harrison Lake FSR taken on July 5 2009. Photos by Laura Parkinson.

Figure 8: Photo of Wood Lake adjacent to West Figure 9: Photo of unnamed swamp adjacent to Harrison FSR where aquatic samples were West Harrison FSR where aquatic samples were conducted taken on July 6 2009. Photo by Sophie- conducted taken on July 6 2009. Photo by Sophie- Anne Blanchette. Anne Blanchette.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 15

Survey Area

Figure 10: Photo of Grace Lake adjacent to West Figure 11: Photo of Wolf Lake adjacent to West Harrison FSR where aquatic samples were Harrison FSR where aquatic samples were conducted taken on July 6 2009. Photo by Laura conducted taken on July 6 2009. Photo by Laura Parkinson. Parkinson.

Summary of Existing Information

The most recent surveys for Dun Skipper were completed in the lower Fraser Valley in 2007 (Knopp et al., 2008) and in 2009 (Knopp et al., 2009 in prep). During their surveys, Knopp et al., 2008 found Dun Skipper west of Hope and south of Lytton. They also observed Dun Skippers in the Morris Valley and Nahatlatch Valleys, extending the known range of this species in B.C. Knopp et al. (2008) conducted some surveys on the east side of Harrison Lake, but no Dun Skipper were observed in this area. Other recent sightings of Dun Skipper were recorded near Boston Bar, Hope, Yale and Lytton in 2002, on Hornby Island, Burns Bog, Salt Spring Island, in 2004 and Denman Island in 2007. A summary of Dun Skipper records is provided by Knopp et al. (2008) and B.C. Conservation Data Centre (2009).

Little information is available on the distribution and life history of Dun Skipper in B.C. Many museum records for Dun Skipper were collected prior to the 1950s, and have no associated habitat information or specific occurrence data, making habitat threats and trends difficult to assess.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 16

Methods

Methods

Habitat Characteristics for Dun Skipper

Dun Skipper has been observed in moist, open grassland or forest meadow habitats (Knopp, et al., 2007; Guppy and Shepard, 2001). Additional habitats include disturbed areas such as power line right-of ways, roadside and adjacent ditches and abandoned road or trails (Shepard, 2000; Opler et al., 2009). Dun Skipper has also been observed in dry areas susceptible to seasonal flooding.

In recent surveys in the lower Fraser Valley (Knopp et al., 2008), Dun Skipper observations were typically associated with habitat that contained ephemeral water bodies such as puddles and vernal pools and often with cliff sides or slopes that provided a wind shelter and a warmer microclimate.

Potential habitat surveyed for Dun Skipper focused on open areas such as regenerating clear cuts or natural openings such as meadows adjacent to forest service roads, where the grass and/or sedge host plants could potentially occur. These sites also offered abundant and diverse flowers for foraging. There was little moist or wet habitat that is associated with some potential host plants available at the time of survey, with the exception of a few marshes. Transects often bisected a patchwork of intact and harvested forests, although survey effort was focused on open areas.

Field surveys

BUTTERFLIES

Using the habitat search image described above, maps and ground-reconnaissance were used to select survey sites. Surveys were conducted on July 3rd - 5th, 2009 and July 17th - 22nd, 2009. Surveys occurred from 9:00 to 17:00 during the peak of species’ flight period (late June to mid August). Although surveys prioritized Dun Skipper habitat, all butterflies were recorded as incidental observations (see Appendix). While open areas were targeted for surveys, forest service roads often dissected a patchwork of forested and clear cut areas, so incidental butterfly observations were also recorded from forested areas. Detailed habitat notes were not taken for incidental observations. Surveys were conducted by the B.C. Conservation Corps Invertebrate Species at Risk crew: two people completing wandering transects through potential habitat. Wandering transects follow no set transects, pre-determined grid, or fixed route and allow the surveyor to change course depending on habitat suitability. Wandering transects are an efficient method of determining butterfly presence when little information is available.

Wandering transects were recorded as tracks using a Garmin handheld GPS (geographic positioning system) unit. When butterflies were observed (including incidental observations), the following information was recorded: 1) butterfly species Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 17

Methods

and number of individuals observed; 2) point location using GPS handheld device (accuracy within 10m); 3) general weather comments; 4) general location description.

Butterflies were identified by sight when possible. A net was used to capture individuals and confirm identification using field guides when it was not possible to identify the species in flight, as well as to photograph specimens.

Due to limited field time sites were only visited once with a minimum of fifteen minutes search time at the site.

Butterflies were identified using Butterflies of British Columbia (Acorn and Sheldon, 2006).

AQUATIC SAMPLES

Aquatic sampling was conducted opportunistically at lakes or marshes, or during weather too cold or rainy for butterfly surveys. Samples were collected using aquatic sampling round-rimmed dip nets, which were used to scrape off the top soft layer of sediment at a depth of up to approximately 1.25 m along the shore of small lakes, roadside marshes and swampy areas. The sediment in the net was then sifted through and any aquatic molluscs or insects observed were taken as specimens. Specimens were preserved in vials in 95% ethanol. Specimens were sent to experts for identification. Sampling was conducted along accessible shoreline. A minimum of 30 and maximum of 90 minutes were spent at each site based on accessibility.

HAND SEARCHING FOR OTHER TERRESTRIAL INSECTS

During butterfly surveys other terrestrial insects were collected opportunistically by hand or using the butterfly net and stored in vials in 95% ethanol. Although several different families were collected, the bias was towards collecting beetles (Coleoptera), particularly Tiger Beetles (sub-family Cicindelinae) and Lady Beetles (Coccinellidae) and grasshoppers (Orthoptera).

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 18

Results

Results

A total of 856 individual butterflies comprising 12 species were observed during surveys in the Harrison Lake area (some individuals could be identified to genera only) (Table 1, Appendix). Dun Skipper, the target species of the surveys was not observed. Two introduced butterfly species were recorded (Cabbage White and European Skipper), comprising 138 observations (approximately 17% of the all observations). No red or blue listed butterfly species were observed (Table 1).

A total of approximately 91.7 hours (5440 minutes) were spent surveying potential habitat for Dun Skipper in the Harrison Lake area. A total distance of 217.8 km of transects were surveyed at 17 different sites over nine days. Time spent surveying was maximized for sites with suitable habitat and minimized for sites with marginal habitat (see potential habitat description above).

Table 2: Summary table of butterfly species observed in Harrison Lake Surveys (*introduced species; **species of conservation concern) Conservation Status Rank* British Number English Name Butterfly Species Columbia Provincial Global Observed Mylitta Crescent Phycoides mylitta Yellow S5 G5 324 Clodius Parnassian Parnassius clodius Yellow S5 G5 195 Cabbage White* Pieris rapae Exotic SNA G5 146 Western Tiger Swallowtail Papilio rutulus Yellow S5 G5 72 Lorquin's Admiral Limenitus lorquini Yellow S5 G5 51 Pale Swallowtail Papilio eurymedon Yellow S5 G5 20 Unidentified Angelwing/ Tortoiseshell 19 Hydaspe Frittilary Speyeria hydaspe Yellow S4S5 G4G5 8 Nymphalis California Tortoiseshell californica Yellow S5 G5 6 Neophasia Pine White menapia Yellow S5 G5 5 Swallowtail spp. Papilio spp. 4 Parnassian spp Parnassius spp 2 Northern Blue Plebejus idas Yellow S5 G5 1 European Skipper* Thymelicus lineola Exotic SNA G5 1 Purplish Copper Lycaena helloides Yellow S5 G5 1 Western Spring Azure Celestrina echo Yellow S5 G5 1 Unknown species 1

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 19

Other Butterfly Observations

Other Butterfly Observations

A total of 856 incidental butterflies comprising twelve species were recorded during surveys in the Harrison Lake area (see Appendix). All of the incidental butterfly species observed are yellow listed (apparently secure) species, with the exception of two introduced species, the Cabbage White and the European skipper. Five species accounted for approximately 90% of all butterfly observations: Lorquin’s Admiral, Western Tiger Swallowtail, Clodius Parnassian, Mylitta Crescent, and Cabbage White (Table 1 & 2).

Table 3: Summary of observations of the five most commonly recorded butterfly species seen in the Harrison Lake area, July 2009 Western Lorquin's Tiger Clodius Mylitta Cabbage Admiral Swallowtail Parnassian Crescent White (Limenitus (Papilio (Parnassius (Phycoides (Pieris Site Date lorquini) rutulus) clodius) mylitta) rapae) Chehalis FSR Branch 1 3-Jul-2009 3 6 Chehalis FSR branch 2 3-Jul-2009 4 Chehalis FSR 1 4-Jul-2009 1 2 1 North Statlu Creek Rd. 4-Jul-2009 7 3 7 1 Chehalis FSR 2 4-Jul-2009 2 1 Harrison Lake FSR Branch 1 5-Jul-2009 2 3 5 3 Harrison Lake FSR Branch 2 5-Jul-2009 2 6 Harrison Lake FSR 5-Jul-2009 3 3 Slollicum FSR 17-Jul-2009 2 4 16 28 47 Bear Creek FSR 18-Jul-2009 5 16 47 66 16 Tolson's FSR 19-Jul-2009 9 28 45 63 28 North Talc Mainline 20-Jul-2009 9 7 34 33 10 Clear Creek FSR 21-Jul-2009 1 2 2 1 unmarked branch road 1 21-Jul-2009 1 Cogburn Creek FSR 21-Jul-2009 4 15 3 unmarked branch road 22-Jul-2009 6 4 12 109 35 South Talc FSR 22-Jul-2009 1 2 10

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Other Butterfly Observations

Table 3: Summary of observations of the five most commonly recorded butterfly species seen in the Harrison Lake area, July 2009 Western Lorquin's Tiger Clodius Mylitta Cabbage Admiral Swallowtail Parnassian Crescent White (Limenitus (Papilio (Parnassius (Phycoides (Pieris Site Date lorquini) rutulus) clodius) mylitta) rapae) Total 51 72 193 324 146 % of Total Observations 6.0 8.4 22.5 37.9 15.2

Mylitta Crescent (Phyciodes mylitta)

Mylitta Crescent is a common butterfly found in southern British Columbia, Washington, Oregon and California. The species is commonly observed in meadows, parks and along roads. Caterpillars utilize thistles (Cirsium and Caardus spp.) as food plant (Opler et al., 2009).

This was the most ubiquitous of any butterfly observed in the Harrison Lake area, although individuals were only observed on the east side of Harrison Lake. Mylitta Crescents were typically observed in regenerating clear cuts or along relatively open stretches of road. They were observed both in flight and perching on roadside vegetation. A total of 324 Mylitta Crescent observations were recorded at almost all sites surveyed on the east side of the lake, accounting for approximately 38% of total butterfly observations (Table 2).

Clodius Parnassian (Parnassius clodius)

Clodius Parnassian is a relatively large white butterfly. This species is endemic to North America where it is found in Southern Alaska, B.C., northern California, Idaho, Montana, Wyoming and Utah (B. C. Conservation Data Centre, 2009). Adults inhabit open alpine woods and openings where its larval food plant, bleeding heart (Dicentra spp.) can be found in adjacent canyon and ridges (B.C. Conservation Data Centre, 2009).

Clodius Parnassian was the second most common butterfly observed. A total of 193 individuals of this species were observed on almost all transects surveyed, accounting for approximately 23% of the total butterflies observed (Table 2).Clodius Parnassian was typically observed adjacent to regenerating clearcuts or natural meadows, although there were some observations in forests adjacent to open areas. Two or more individuals were often observed flying together. This species was observed in flight and perched on roadside flowers and vegetation. It was one of the few butterflies observed in a short survey of Harrison Lake FSR, one of the more major forest service roads that proved to be too disturbed and heavily used to warrant further survey.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 21

Other Butterfly Observations

Figure 12: Photos of the dorsal and ventral views of a Clodius Parnassian taken on an unmarked road branching off of Chehalis FSR on July 5th 2009. Photos by Laura Parkinson.

Cabbage White (Pieris rapae)

Cabbage White butterflies are an introduced species that has now invaded North America south of the Arctic. They are present in highly disturbed areas as well as in more natural settings (B.C. Conservation Data Centre, 2009). Cabbage White utilizes host plants from the mustard family.

Cabbage White was the third most common species observed, with 146 individuals accounting for approximately 15% of the total butterfly observations. This butterfly occurred in almost all the sites surveyed (Table 2).Cabbage White butterflies were ubiquitous both in regenerating clear cuts and in open forest. They were often observed flying in groups of two or more.

Western Tiger Swallowtail (Papilio rutulus)

Western Tiger Swallowtail is found in the southern part of the province from Vancouver Island to the lower Fraser River Valley, east to the Okanagan and Kootenay valleys. The species is found in or adjacent to forested, woody areas nearby running water where moisture is available for mud-puddling (Species Bank, 2003). This includes along creeks, rivers, canyons and urban parks. The larvae are known to feed on a diversity of trees and shrub including Willow (Larix spp.), Poplar (Populus spp.) and Birch (Betula spp.) (Natureserve, 2009).

Western Tiger Swallowtail was the fourth most common butterfly observed with 72 individuals accounting for approximately 8.5% of the total butterfly observations. This butterfly was recorded at a majority of transects surveyed (Table 2). Western Tiger Swallowtail was observed along stretches of forest service road adjacent to mature forest with natural openings, in patches of mature forest adjacent to regenerating

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 22

Other Butterfly Observations clearcuts, or in regenerating clearcuts. It was often observed in flight but also perching on roadside shrubs or flowers.

Lorquin’s Admiral (Limenitus lorquini)

Lorquin’s Admiral is wide spread across the southern part of B.C. including Vancouver Island up to Emerald Lake. It is found in or adjacent to forested areas, parks and orchards. Its larval food plant includes many deciduous trees like Willow, Poplar and Cottonwood, and Cherry (Prunus spp.) (Guppy and Shepard, 2001).

Lorquin’s Admiral was the fifth most common species observed with a total of 51 individuals accounting for approximately 6% of the total butterfly observations. It was observed at on a majority of the transects surveyed (Table 2). Lorquin’s Admiral was observed adjacent to areas with dense shrubs, both young and mature forests, or in open areas adjacent to forests. This distinctive butterfly is easy to recognize in flight and was sometimes observed flying high in the foliage. It was also observed resting on the road or roadside shrubs.

Figure 13: Photo of Lorquin’s admiral taken on North Statlu FSR, July 4 2009. Photo by Sophie-Anne Blanchette.

Pale Swallowtail (Papilio eurymedon)

Pale Swallowtail occurs only in the southern portion of B.C., as far north as Wells Gray and South Tweedsmuir Provincial parks (Species Bank, 2003). The adults prefer open woodlands, foothills and streamsides. This butterfly is often found hilltopping and mud- puddling (Species Bank, 2003) Larval food plant includes trees and shrubs in the Rose (Rosaceae), Buckthorn (Rhamnaceae) and Birch (Betulaceae) families including Cherry (Prunus emarginata) and Ash (Fraxinus spp.)(Opler et al., 2009).

Twenty pale swallowtails were observed at four different sites: seven on North Talc Mainline, nine on Tolson’s Fsr, three on unmarked branch road, and one on unmarked branch road 2 (see Table 2 for dates transects were surveyed)Pale Swallowtail was

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 23

Other Butterfly Observations

typically observed in flight along stretches of forest service road adjacent to mature forest with natural openings, or in patches of mature forest adjacent to regenerating clearcuts. It was sometimes observed flying in the foliage of trees and shrubs lining the road. This species was only observed on the east side of Harrison Lake.

Hydaspe Fritillary (Speyeria hydaspe)

Hydaspe Fritillary is found throughout B.C. from the Rocky Mountains west to Vancouver Island. The butterfly most abundant in July (flies from June to August) and prefers moist clearings in forested areas where lots of flowers are available for nectaring. The rhodope subspecies is the only one present in Canada (Species Bank, 2003).

Nine individuals were observed at three different sites on the east side of Harrison Lake: four on Bear Creek forest service road one on Tolson’s Fsr and three on North Talc Mainline. All observations were made in the afternoon. Hydaspe Fritillary was observed in regenerating clearcuts with tall shrubs adjacent to forests, often near a small creek or stream (Fig 9).

Figure 14: Photo of a small stream running across Bear Creek FSR near where Hydaspe Fritillary was observed taken on July 18 2009. Photo by Laura Parkinson.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 24

Other Butterfly Observations

Figure 15: Dorsal and ventral view of a Hydaspe Fritillary observed on Bear Creek FSR taken on July 18 2009. Photos by Sophie-Anne Blanchette.

California Tortoiseshell (Nymphalis californica)

California Tortoiseshell is found in woodland and forest openings, brushy areas and forest edges (Opler et al., 2009). This migratory species uses Redstem Caenothus (Caenothus sanguineus) or other Caenothus spp. as its larval host plant. California Tortoiseshell hibernates in B.C. under woodpiles, dead woods, old buildings and the adults emerge in the spring (Guppy and Shepard, 2001). The adults feed on a diversity of wild flowers.

Six individuals were observed at six different sites: Chehalis FSR 2, North Statlu Creek FSR, Harrison Lake FSR branch 2, North Talc Mainline, Bear Creek FSR and unmarked branch road (see Table 2 for dates transects were surveyed). This species was typically observed along forest services roads adjacent to patches of mature forest bordered by regenerating clearcuts or in mature forest with natural openings.

Figure 16: California tortoiseshell taken on North Statlu Creek FSR on July 4th 2009. Photo by Laura Parkinson.

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 25

Other Butterfly Observations

Pine White (Neophasia menapia)

Pine White butterflies are found in southern B.C. and Vancouver Island and as far north as the Chilcotin in the Fraser Valley and the Bella Coola Valley on the coast (Species Bank, 2003). The adults prefer coniferous forest where they lay their eggs on Pine (Pinus) and Fir (Abies) species (Guppy and Shepard 2001).

A total of five Pine Whites were observed at only two sites: four butterflies on Cogburn Creek FSR and one on Tolson’s FSR (Table 2). All observations were made along sections of road adjacent to patches of mature forest.

Northern Blue (Plebejus idas)

The Northern Blue is found throughout mainland British Columbia with the exception of the north coast and the Cascades. It is found flying in open meadows up to and above the timberline from July to early August. It has been observed ovipositing on Dwarf Bilberry (Vaccinium caespitosum) in Wisconsin and Ontario and this is thought to be the host plant for the Northern Blue in B.C. (Guppy and Shepard, 2001).

One tattered Northern Blue butterfly was observed on Tolson’s FSR on July 19 2009, perched on roadside grasses.

Figure 17: Photos of an Northern Blue observed on Tolson’s FSR on July 19th 2009. Photo by Laura Parkinson.

European Skipper (Thymelicus lineola)

European Skipper is a small burnt orange butterfly with a black edge along the dorsal wing surface. This exotic species was first recorded in Terrace in the 1960’s and has spread across the southern interior and lower mainland (Guppy and Shepard, 2001). The butterfly nectars on low-growing flowers such as Orange Hawkweed (Hieracium aurantiacum), Oxeye Daisy (Leucanthemum vulgare), thistles and clover (Trifolium spp.) can be found in disturbed areas (Opler et al., 2009). European Skipper uses grasses,

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 26

Other Butterfly Observations

such as Timothy Grass (Phleum pratense), as a host plant and can be found in meadows, fields and along roadsides (Guppy and Shepard, 2001).

One individual of this exotic species was observed on July 4th on Chehalis FSR, perched on some roadside flowers.

Purplish Copper (Lycaena helloides)

Purplish Copper is found throughout southern B.C. The butterfly is usually the most common copper species observed but its population size can vary from year to year. The adults are known to fly in two generations, the first one in June and July and the second in August and September. Purplish Copper prefers open fields and roadsides but can also be found near wet meadows. The adults tend to wander far form their larval food plant, usually a member of the Buckwheat family (Polygonaceae), including Knotweed (Polygonum spp.) and Docks (Rumex spp.) (Species Bank, 2003).

One Purplish Copper individual was observed on Bear Creek road on July 18 2009, adjacent to a regenerating clearcut.

Figure 18: Purplish Copper observed on Bear Creek FSR on July 18 2009. Photo by Sophie-Anne Blanchette.

Western Spring Azure (Celestrina echo)

Western Spring Azure is found in southern B.C. and on southeastern Vancouver Island to the tip of Baja California and Northern Mexico (Guppy and Shepard, 2001).The adults prefer openings and edges of deciduous forests and wooded freshwater marshes and swamps. Larval plants consist of a variety of woody shrubs and herbs such as dogwood (Cornus florida), New Jersey tea (Ceanothus americana) and meadowsweet (Spiraea salicifolia). Butterflies nectar on many species including Blackberry (Rubus spp.),

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 27

Other Butterfly Observations

Spreading Dogbane (Apocynum androsaemifolium) and Showy Milkweed (Asclepius speciosa).

One worn individual was observed on July 5th on an unmarked road branching off of Harrison Lake Forest Service Road. This deactivated road had some open areas with a few flowers where the western spring azure individual was observed which ended in mature forest.

Other Invertebrate Observations

Aquatic Samples

Five hours and five minutes were spent conducting aquatic surveys at five different sites on July 5 and July 6 2009 (see Table 1). A majority of aquatic invertebrates found in the samples were either Odonate nymphs or molluscs (see Table 4).

One aquatic beetle was found in an unnamed marsh adjacent to Harrison FSR (see Table 1; Figure 7), identified as Gyrinus consobrinus. Rex Kenner, the expert who identified the specimen, considered this an interesting record.

Table 4: Aquatic molluscs collected opportunistically from roadside lakes and marshes near Harrison Lake, as identified by Kristiina Ovaska and Lennart Sopuck Common Name Species Name Date Location Number Pea/fingernail Sphaeridae 05-Jul-09 Unnamed 1 Clam roadside marsh Two-ridge Rams- Helisoma 06-Jul-09 Wood Lake 4 horn (anceps?) Rams-horn Helisoma sp. 06-Jul-09 Wood Lake 6 Pea/fingernail Sphaeridae 06-Jul-09 Grace Lake 1 Clam Gyro Gyraulus sp. 06-Jul-09 Wolf Lake 2

Long Fingernailclam Musculium 06-Jul-09 Wolf Lake 1 transversum Pea/fingernailclam Sphaeridae 06-Jul-09 Wolf Lake 1 Rams-horn Helisoma sp. 06-Jul-09 Wolf Lake 2 (juvenile) Pea/fingernailclam Sphaeridae 06-Jul-09 Wolf Lake 2

Hand Collected Terrestrial Insects

Terrestrial insects were collected opportunistically by hand in the Harrison Lake area. Due to time constraints, not all specimens were identified, and some were not identified to greater precision than the level of family or genus. Grasshopper specimens collected are awaiting identification.

None of the species below been assessed and assigned a conservation status by the B.C. Conservation Data Center, with the exception of the Western Tiger Beetle, which is Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 28

Other Invertebrate Observations yellow listed in B.C. The Asian Spotted Ladybird Beetle is an exotic species introduced from Japan. It was introduced intentionally as a form of biological control for aphids, and has become very common in some areas, particularly urban areas (Haggard and Haggard, 2006).

Table 5: Terrestrial insects collected opportunistically in the Harrison Lake area, as identified by Rob Cannings Common Name Order Family Genus Species Date Snail-killer Carabid Coleoptera Carabidae Scaphinotus angusticollis 3-Jul-09 Multi-coloured Asian Ladybird Beetle Coleoptera Coccinellidae Harmonia axyridis 3-Jul-09 Ground Beetles Coleoptera Carabidae Ditylus quadricollis 5-Jul-09 Long-horned Beetles Coleoptera Cerambycidae 5-Jul-09 Western Blood-red Ladybird Beetle Coleoptera Coccinellidae Cycloneda polita 5-Jul-09 Multi-coloured Asian Ladybird Beetle Coleoptera Coccinellidae Harmonia axyridis 5-Jul-09 Ants Hymenoptera Formicidae Camponotus 5-Jul-09 Cicada Homoptera Cicadidae 18-Jul-09 Eyed Elater Coleoptera Elateridae Alaus melanops 19-Jul-09 Western Tiger Beetle Coleoptera Carabidae Cicindela oregona 21-Jul-09 Long-horned Beetles Coleoptera Cerambycidae 21-Jul-09 Black Pine Sawyer Coleoptera Cerambycidae Monochamus scutellatus 21-Jul-09 Western Blood-red Ladybird Beetle Coleoptera Coccinellidae Cycloneda polita 21-Jul-09 Multi-coloured Asian Ladybird Beetle Coleoptera Coccinellidae Harmonia axyridis 21-Jul-09 Multi-coloured Asian Ladybird Beetle Coleoptera Coccinellidae Harmonia axyridis 22-Jul-09 Scentless Plant-bug Hemiptera Rhopalidae Harmostes reflexulus 22-Jul-09

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 29

Discussion

Discussion

The target species of the surveys, Dun Skipper, was not observed. In recent surveys in the lower Fraser Valley, Knopp et al. (2008) often found Dun Skipper associated with ephemeral water bodies, and some potential host plants of Dun Skipper are also associated with wet or moist habitats. Most of the habitat surveyed in the Harrison Lake area was dry at the time of survey, with the exception of some small creeks and streams. This may have limited the potential habitat available for Dun Skipper, although it has also recently been recorded in dry habitats (Knopp et al., 2008).

Little information is available on the general biology and distribution of Dun Skipper in B.C. Much of Dun Skippers basic biology in B.C. including, the population size, larval food plant, and detailed habitat requirements, are unknown. More research on these topics would be helpful in the selection of suitable habitat for surveys and to appropriately select future protected areas.

It is important to note that some forest service roads had been sprayed with pesticides prior to our surveys, on both sides of the lake. Since butterflies may utilize plants targeted by those pesticides, it could have an impact on the butterfly population in the Harrison Lake area. Disturbance to habitat caused by high traffic on main forest service roads and high recreational use on both sides of the Harrison Lake may also be impacting butterflies in this area. Previous observations of Dun Skipper have been recorded from disturbed habitat types, so this does not preclude Dun Skipper from occurring around Harrison Lake.

In general, all butterflies were observed in greater numbers on the east side of Harrison Lake. Several species were observed only on the east side of the lake, including Pale Swallowtails, Pine Whites, Hydaspe Fritillaries and Mylitta Crescents, the latter by far the most ubiquitous species observed accounting for approximately 38% of the total butterfly observations. The reason for the differing results on either side of the lake is uncertain, as habitat surveyed on both sides of the lake was similar, and weather conditions were optimal for a majority of transects surveyed.

Recommendations for future surveys

Additional areas in the lower mainland region (south of the Fraser, Mission area), are recommended for further surveys. Surveys should also be conducted in Burns Bog, where Dun Skipper has been recently found (Knopp et al., 2008). Few surveys have been conducted in this expansive wetland which has some level of protection through its status as a park.

Survey efforts should also focus on Vancouver Island and the Gulf Islands, where the most recent voucher specimens of Dun Skipper were collected in the 1960s, although a handful of individuals were also observed there in the 1990s (Shepard, 2000).

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 30

References

References

Acorn, J. and I. Sheldon. 2006 Butterflies of British Columbia. Lone Pine Publishing, 360 pp.

B.C. Conservation Data Centre. 2008. BC Species and Ecosystems Explorer. B.C.Minist. Of Environ. Victoria, B.C. Available: http://a100.gov.bc.ca/pub/eswp/ (accessed August 20, 2009).

Guppy, C. and J. Shepard. 2001. Butterflies of British Columbia. UBC Press: Vancouver. 414 pp.

Haggard, P. and J. Haggard. 2006. Insects of the Pacific Northwest. Timber Press Field Guide: Oregon. 295 pp.

Heron, J. 2007. Butterfly Surveys in Burns Bog. Ministry of Environment, Ecosystem Branch, Wildlife Science Section. Vancouver, B.C., 21pp.

Knopp, D, L. Larkin, and J.Heron. 2008. Survey for Dun Skipper in the Lower Fraser Valley, B.C., B.C. Ministry of Environment, UBC Campus, 315 – 2202 Main Mall, Vancouver, B.C. 53pp.

NatureServe. 2009. NatureServe Explorer: An online encyclopedia of life [web application]. Version 7.1. NatureServe, Arlington, Virginia. Available http://www.natureserve.org/explorer.

Opler, P. A., K. Lotts, and T. Naberhaus, coordinators. 2009. Butterflies and Moths of North America. Bozeman, MT: Big Sky Institute. http://www.butterfliesandmoths.org/ (accessed August 18, 2009).

Pojar, J. and A. McKinnon. 1994. Plants of Coastal British Columbia including Washington, Oregon and Alaska. B.C. Ministry of Forests and Lone Pine Publishing, Vancouver, B.C. 526pp.

Shepard, J.H. 2000. Status of five butterflies and skippers in British Columbia. B.C. Minist. Environ., Lands and Parks, Wildl. Branch and Resour. Inv. Branch, Victoria, BC. 27pp.

Species Bank, Canadian Biodiversity Information Facility. 2003. Government of Canada. http://www.cbif.gc.ca/spp_pages/butterflies/speciesindex_e.php. (Accessed August 31, 2009).

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 31

Appendix

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting 05/07/09 Anglewing or Tortoiseshell 1 5477250 582588 05/07/09 Anglewing or Tortoiseshell 10 5477186 582399 20/07/09 Anglewing or Tortoiseshell 1 5485588 594514 Polygonia 04/07/09 Anglewing spp. spp. 1 5468901 564582 Polygonia 20/07/09 Anglewing spp. spp. 2 5485003 595445 Polygonia 22/07/09 Anglewing spp. spp. 1 5476924 590072 04/07/09 Anglewing spp.? 1 5466443 568496 Plebejus 19/07/09 Northern Blue idas 1 5483725 590164 04/07/09 Cabbage White Pieris rapae 1 5467766 566708 04/07/09 Cabbage White Pieris rapae 1 5465687 572114 04/07/09 Cabbage White Pieris rapae 1 5463996 574135 05/07/09 Cabbage White Pieris rapae 1 5471705 583078 05/07/09 Cabbage White Pieris rapae 1 5471611 583592 05/07/09 Cabbage White Pieris rapae 1 5471567 583658 17/07/09 Cabbage White Pieris rapae 2 5470791 591211 17/07/09 Cabbage White Pieris rapae 2 5470709 591504 17/07/09 Cabbage White Pieris rapae 1 5470615 591584 17/07/09 Cabbage White Pieris rapae 3 5470307 591437 17/07/09 Cabbage White Pieris rapae 1 5469579 592235 17/07/09 Cabbage White Pieris rapae 2 5470788 591250 17/07/09 Cabbage White Pieris rapae 1 5470697 591262 17/07/09 Cabbage White Pieris rapae 1 5470725 591454 17/07/09 Cabbage White Pieris rapae 1 5470723 591515 17/07/09 Cabbage White Pieris rapae 1 5470632 591542 17/07/09 Cabbage White Pieris rapae 3 5470807 591519 17/07/09 Cabbage White Pieris rapae 3 5470859 591524 17/07/09 Cabbage White Pieris rapae 4 5470935 591511 17/07/09 Cabbage White Pieris rapae 1 5470679 591202 17/07/09 Cabbage White Pieris rapae 1 5470481 591200 17/07/09 Cabbage White Pieris rapae 1 5470217 591504 17/07/09 Cabbage White Pieris rapae 4 5470161 591523

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 32

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting 17/07/09 Cabbage White Pieris rapae 2 5469885 591609 17/07/09 Cabbage White Pieris rapae 2 5469749 591663 17/07/09 Cabbage White Pieris rapae 3 5469520 591715 17/07/09 Cabbage White Pieris rapae 2 5469316 592237 17/07/09 Cabbage White Pieris rapae 6 5469608 592217 18/07/09 Cabbage White Pieris rapae 2 5482050 589700 18/07/09 Cabbage White Pieris rapae 2 5481038 589866 18/07/09 Cabbage White Pieris rapae 3 5482289 589599 18/07/09 Cabbage White Pieris rapae 1 5481892 589786 18/07/09 Cabbage White Pieris rapae 2 5481818 589804 18/07/09 Cabbage White Pieris rapae 1 5481539 589852 18/07/09 Cabbage White Pieris rapae 1 5480568 589941 18/07/09 Cabbage White Pieris rapae 2 5480056 592144 18/07/09 Cabbage White Pieris rapae 1 5479996 592212 18/07/09 Cabbage White Pieris rapae 1 5479431 593555 19/07/09 Cabbage White Pieris rapae 1 5483507 590080 19/07/09 Cabbage White Pieris rapae 3 5483429 590057 19/07/09 Cabbage White Pieris rapae 3 5483184 590082 19/07/09 Cabbage White Pieris rapae 2 5482847 589918 19/07/09 Cabbage White Pieris rapae 2 5482991 590076 19/07/09 Cabbage White Pieris rapae 3 5483021 590165 19/07/09 Cabbage White Pieris rapae 2 5482912 590145 19/07/09 Cabbage White Pieris rapae 2 5482794 590073 19/07/09 Cabbage White Pieris rapae 2 5482690 590176 19/07/09 Cabbage White Pieris rapae 1 5482519 590199 20/07/09 Cabbage White Pieris rapae 3 5485781 594263 20/07/09 Cabbage White Pieris rapae 1 5488365 591166 20/07/09 Cabbage White Pieris rapae 2 5487608 592526 20/07/09 Cabbage White Pieris rapae 1 5486801 593313 20/07/09 Cabbage White Pieris rapae 1 5485665 594375 20/07/09 Cabbage White Pieris rapae 1 5485588 594514 21/07/09 Cabbage White Pieris rapae 1 5488041 589710 21/07/09 Cabbage White Pieris rapae 1 21/07/09 Cabbage White Pieris rapae 1 5488569 589823 21/07/09 Cabbage White Pieris rapae 1 5498722 584978 22/07/09 Cabbage white Pieris rapae 2 5477259 589843 22/07/09 Cabbage White Pieris rapae 2 5476946 589401

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 33

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting 22/07/09 Cabbage White Pieris rapae 4 5477267 589840 22/07/09 Cabbage White Pieris rapae 1 5477502 589791 22/07/09 Cabbage White Pieris rapae 1 5477067 590031 22/07/09 Cabbage White Pieris rapae 3 5476349 590096 22/07/09 Cabbage White Pieris rapae 3 5476098 590123 22/07/09 Cabbage White Pieris rapae 2 5475817 590098 22/07/09 Cabbage White Pieris rapae 4 5475799 590199 22/07/09 Cabbage White Pieris rapae 10 5475603 590239 19/07/09 Cabbage White Pieris rapae 2 5482965 590195 California or Compton Nymphalis 18/07/09 Tortoiseshell californica 1 5482009 589719 Nymphalis 04/07/09 California Tortoiseshell? californica 1 5467427 566959 Nymphalis 22/07/09 California Tortoiseshell? californica 1 5476637 590152 Nymphalis 04/07/09 California Tortoiseshell californica 1 5465348 572515 Nymphalis 05/07/09 California Tortoiseshell californica 1 5477206 583045 Nymphalis 20/07/09 California Tortoiseshell californica 1 5487573 592561 Parnassius 03/07/09 Clodius Parnassian clodius 2 5457991 575091 Parnassius 03/07/09 Clodius Parnassian clodius 1 5458080 574771 Parnassius 03/07/09 Clodius Parnassian clodius 1 5457673 574486 Parnassius 03/07/09 Clodius Parnassian clodius 1 5457988 575098 Parnassius 03/07/09 Clodius Parnassian clodius 1 5458084 574772 Parnassius 04/07/09 Clodius Parnassian clodius 1 5466389 569062 Parnassius 04/07/09 Clodius Parnassian clodius 1 5468278 565670 Parnassius 04/07/09 Clodius Parnassian clodius 1 5468904 564576 Parnassius 04/07/09 Clodius Parnassian clodius 1 5469139 564766 Parnassius 04/07/09 Clodius Parnassian clodius 1 5468834 564780 Parnassius 04/07/09 Clodius Parnassian clodius 1 5468098 565891 Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 34

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Parnassius 04/07/09 Clodius Parnassian clodius 1 5464965 573341 Parnassius 04/07/09 Clodius Parnassian clodius 1 5466416 568443 Parnassius 04/07/09 Clodius Parnassian clodius 1 5465688 572116 Parnassius 04/07/09 Clodius Parnassian clodius 1 5465686 572113 Parnassius 04/07/09 Clodius Parnassian clodius 1 5463997 574124 Parnassius 05/07/09 Clodius Parnassian clodius 1 5471483 583384 Parnassius 05/07/09 Clodius Parnassian clodius 1 5471676 583107 Parnassius 05/07/09 Clodius Parnassian clodius 1 5472291 582842 Parnassius 05/07/09 Clodius Parnassian clodius 1 5468251 584430 Parnassius 05/07/09 Clodius Parnassian clodius 1 5468347 584465 Parnassius 05/07/09 Clodius Parnassian clodius 1 5468769 584365 Parnassius 05/07/09 Clodius Parnassian clodius 1 5471817 582941 Parnassius 05/07/09 Clodius Parnassian clodius 1 5471611 583593 Parnassius 05/07/09 Clodius Parnassian clodius 2 5477148 583332 Parnassius 05/07/09 Clodius Parnassian clodius 3 5477083 583268 Parnassius 05/07/09 Clodius Parnassian clodius 1 5477155 583187 Parnassius 05/07/09 Clodius Parnassian clodius 1 5477203 583033 Parnassius 05/07/09 Clodius Parnassian clodius 1 5477186 582399 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470871 591320 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470706 591502 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470640 591543 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470761 591508

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 35

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Parnassius 17/07/09 Clodius Parnassian clodius 1 5470749 591419 Parnassius 17/07/09 Clodius Parnassian clodius 1 5469166 591830 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470777 591444 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470706 591475 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470632 591542 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470859 591524 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470556 591276 Parnassius 17/07/09 Clodius Parnassian clodius 1 5470161 591523 Parnassius 17/07/09 Clodius Parnassian clodius 1 5469090 591875 Parnassius 17/07/09 Clodius Parnassian clodius 1 5468997 592253 Parnassius 17/07/09 Clodius Parnassian clodius 2 5469833 591641 Parnassius 18/07/09 Clodius Parnassian clodius 1 5480245 591073 Parnassius 18/07/09 Clodius Parnassian clodius 1 5480194 591602 Parnassius 18/07/09 Clodius Parnassian clodius 2 5480069 591962 Parnassius 18/07/09 Clodius Parnassian clodius 2 5480054 592133 Parnassius 18/07/09 Clodius Parnassian clodius 5 5479946 592525 Parnassius 18/07/09 Clodius Parnassian clodius 10 5479894 592732 Parnassius 18/07/09 Clodius Parnassian clodius 1 5480686 589933 Parnassius 18/07/09 Clodius Parnassian clodius 1 5480475 589955 Parnassius 18/07/09 Clodius Parnassian clodius 1 5480235 590780 Parnassius 18/07/09 Clodius Parnassian clodius 1 5480242 591351 Parnassius 18/07/09 Clodius Parnassian clodius 1 5480057 592030

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 36

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Parnassius 18/07/09 Clodius Parnassian clodius 1 5480056 592144 Parnassius 18/07/09 Clodius Parnassian clodius 2 5479996 592212 Parnassius 18/07/09 Clodius Parnassian clodius 2 5479951 592267 Parnassius 18/07/09 Clodius Parnassian clodius 3 5479926 592333 Parnassius 18/07/09 Clodius Parnassian clodius 1 5479893 592442 Parnassius 18/07/09 Clodius Parnassian clodius 2 5479945 592581 Parnassius 18/07/09 Clodius Parnassian clodius 2 5479901 592666 Parnassius 18/07/09 Clodius Parnassian clodius 6 5479679 593198 Parnassius 18/07/09 Clodius Parnassian clodius 2 5479431 593555 Parnassius 19/07/09 Clodius Parnassian clodius 3 5482850 589922 Parnassius 19/07/09 Clodius Parnassian clodius 1 5483034 590179 Parnassius 19/07/09 Clodius Parnassian clodius 2 5482965 590195 Parnassius 19/07/09 Clodius Parnassian clodius 2 5482721 590160 Parnassius 19/07/09 Clodius Parnassian clodius 1 5482180 590288 Parnassius 19/07/09 Clodius Parnassian clodius 1 5484021 589790 Parnassius 19/07/09 Clodius Parnassian clodius 1 5483573 589830 Parnassius 19/07/09 Clodius Parnassian clodius 2 5483553 590122 Parnassius 19/07/09 Clodius Parnassian clodius 2 5483507 590080 Parnassius 19/07/09 Clodius Parnassian clodius 2 5483481 590062 Parnassius 19/07/09 Clodius Parnassian clodius 1 5483184 590082 Parnassius 19/07/09 Clodius Parnassian clodius 2 5482968 589976 Parnassius 19/07/09 Clodius Parnassian clodius 2 5482847 589918

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 37

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Parnassius 19/07/09 Clodius Parnassian clodius 2 5482799 589899 Parnassius 19/07/09 Clodius Parnassian clodius 2 5482991 590076 Parnassius 19/07/09 Clodius Parnassian clodius 2 5483021 590165 Parnassius 19/07/09 Clodius Parnassian clodius 1 5482970 590205 Parnassius 19/07/09 Clodius Parnassian clodius 2 5482912 590145 Parnassius 19/07/09 Clodius Parnassian clodius 1 5482794 590073 Parnassius 19/07/09 Clodius Parnassian clodius 2 5482536 590035 Parnassius 19/07/09 Clodius Parnassian clodius 2 5482602 590072 Parnassius 19/07/09 Clodius Parnassian clodius 2 5482690 590176 Parnassius 19/07/09 Clodius Parnassian clodius 1 5482519 590199 Parnassius 20/07/09 Clodius Parnassian clodius 1 5488351 590197 Parnassius 20/07/09 Clodius Parnassian clodius 1 5488995 590977 Parnassius 20/07/09 Clodius Parnassian clodius 1 5488184 591383 Parnassius 20/07/09 Clodius Parnassian clodius 1 5487991 591813 Parnassius 20/07/09 Clodius Parnassian clodius 2 5487623 592475 Parnassius 20/07/09 Clodius Parnassian clodius 7 5486704 593329 Parnassius 20/07/09 Clodius Parnassian clodius 1 5488826 590911 Parnassius 20/07/09 Clodius Parnassian clodius 1 5488033 591463 Parnassius 20/07/09 Clodius Parnassian clodius 2 5487864 592001 Parnassius 20/07/09 Clodius Parnassian clodius 1 5487697 592297 Parnassius 20/07/09 Clodius Parnassian clodius 1 5487648 592432 Parnassius 20/07/09 Clodius Parnassian clodius 1 5487534 592667

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 38

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Parnassius 20/07/09 Clodius Parnassian clodius 2 5487079 593079 Parnassius 20/07/09 Clodius Parnassian clodius 2 5486952 593238 Parnassius 20/07/09 Clodius Parnassian clodius 3 5486868 593269 Parnassius 20/07/09 Clodius Parnassian clodius 2 5486801 593313 Parnassius 20/07/09 Clodius Parnassian clodius 1 5485964 593958 Parnassius 20/07/09 Clodius Parnassian clodius 1 5485588 594514 Parnassius 20/07/09 Clodius Parnassian clodius 1 5485299 595107 Parnassius 21/07/09 Clodius Parnassian clodius 1 5488637 589887 Parnassius 21/07/09 Clodius Parnassian clodius 2 5488837 590129 Parnassius 21/07/09 Clodius Parnassian clodius 1 5494527 584890 Parnassius 21/07/09 Clodius Parnassian clodius 1 5488662 589936 Parnassius 21/07/09 Clodius Parnassian clodius 1 5498591 584931 Parnassius 21/07/09 Clodius Parnassian clodius 1 5499386 584929 Parnassius 22/07/09 Clodius Parnassian clodius 5488095 590616 Parnassius 22/07/09 Clodius Parnassian clodius 1 5477941 589492 Parnassius 22/07/09 Clodius parnassian clodius 2 5477180 590018 Parnassius 22/07/09 Clodius parnassian clodius 1 5476316 590089 Parnassius 22/07/09 Clodius parnassian clodius 1 5475593 590255 Parnassius 22/07/09 Clodius Parnassian clodius 1 5488071 590438 Parnassius 22/07/09 Clodius Parnassian clodius 1 5478195 589488 Parnassius 22/07/09 Clodius Parnassian clodius 1 5477067 590031 Parnassius 22/07/09 Clodius Parnassian clodius 1 5475603 590239

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 39

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Parnassius 22/07/09 Clodius Parnassian clodius 1 5477152 590059 Parnassius 22/07/09 Clodius Parnassian clodius 1 5476650 590157 Parnassius 22/07/09 Clodius Parnassian clodius 2 5476637 590152 Comma/Tortoiseshell 17/07/09 unknown 1 5470784 591436 Thymelicus 04/07/09 European Skipper? lineola 1 5464164 573853 Speyeria 20/07/09 Frittilary (Hydaspe?) hydaspe 1 5488810 590900 Speyeria 18/07/09 Hydaspe Fritillary hydaspe 1 5479899 592695 Speyeria 20/07/09 Hydaspe Fritillary hydaspe 1 5488208 591375 Speyeria 18/07/09 Hydaspe Frittilary hydaspe 1 5480054 592133 Speyeria 18/07/09 Hydaspe Frittilary hydaspe 1 5479894 592732 Speyeria 18/07/09 Hydaspe Frittilary hydaspe 1 5479055 593790 Speyeria 20/07/09 Hydaspe Frittilary hydaspe 1 5484999 595449 Limenitus 22/07/09 Lorquins Admiral lorquini 1 5477785 589598 Limenitus 03/07/09 Lorquin's Admiral lorquini 1 5457983 575072 Limenitus 03/07/09 Lorquin's Admiral lorquini 1 5458144 574851 Limenitus 03/07/09 Lorquin's Admiral lorquini 1 5457978 575052 Limenitus 03/07/09 Lorquin's Admiral lorquini 1 5459921 575052 Limenitus 03/07/09 Lorquin's Admiral lorquini 1 5459521 574779 Limenitus 03/07/09 Lorquin's Admiral lorquini 1 5459556 574759 Limenitus 04/07/09 Lorquin's Admiral lorquini 1 5468024 565980 Limenitus 04/07/09 Lorquin's Admiral lorquini 1 5468517 565415 Limenitus 04/07/09 Lorquin's Admiral lorquini 1 5466366 569055

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 40

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Limenitus 04/07/09 Lorquin's Admiral lorquini 1 5468138 565870 Limenitus 04/07/09 Lorquin's Admiral lorquini 1 5468357 565612 Limenitus 04/07/09 Lorquin's Admiral lorquini 1 5468522 565433 Limenitus 04/07/09 Lorquin's Admiral lorquini 1 5468841 564774 Limenitus 04/07/09 Lorquin's Admiral lorquini 1 5466046 571559 Limenitus 05/07/09 Lorquin's Admiral lorquini 1 5472049 582763 Limenitus 05/07/09 Lorquin's Admiral lorquini 1 5471697 583072 Limenitus 05/07/09 Lorquin's Admiral lorquini 1 5477202 583034 Limenitus 05/07/09 Lorquin's Admiral lorquini 1 5477367 581969 Limenitus 17/07/09 Lorquin's Admiral lorquini 1 5470867 591239 Limenitus 17/07/09 Lorquin's Admiral lorquini 1 5470743 591237 Limenitus 18/07/09 Lorquin's Admiral lorquini 1 5481460 589922 Limenitus 18/07/09 Lorquin's Admiral lorquini 1 5480069 591962 Limenitus 18/07/09 Lorquin's Admiral lorquini 1 5482053 589711 Limenitus 18/07/09 Lorquin's Admiral lorquini 1 5481432 589970 Limenitus 18/07/09 Lorquin's Admiral lorquini 1 5480568 589941 Limenitus 19/07/09 Lorquin's Admiral lorquini 1 5484036 590149 Limenitus 19/07/09 Lorquin's Admiral lorquini 1 5483481 590071 Limenitus 19/07/09 Lorquin's Admiral lorquini 1 5482850 589922 Limenitus 19/07/09 Lorquin's Admiral lorquini 1 5484660 589842 Limenitus 19/07/09 Lorquin's Admiral lorquini 1 5483481 590062 Limenitus 19/07/09 Lorquin's Admiral lorquini 1 5483287 590108

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 41

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Limenitus 19/07/09 Lorquin's Admiral lorquini 1 5483184 590082 Limenitus 19/07/09 Lorquin's Admiral lorquini 1 5482799 589899 Limenitus 20/07/09 Lorquin's Admiral lorquini 2 5487121 593024 Limenitus 20/07/09 Lorquin's Admiral lorquini 1 5485543 594577 Limenitus 20/07/09 Lorquin's Admiral lorquini 1 5487648 592432 Limenitus 20/07/09 Lorquin's Admiral lorquini 1 5487608 592526 Limenitus 20/07/09 Lorquin's Admiral lorquini 1 5487534 592667 Limenitus 20/07/09 Lorquin's Admiral lorquini 1 5487079 593079 Limenitus 20/07/09 Lorquin's Admiral lorquini 1 5486801 593313 Limenitus 20/07/09 Lorquin's Admiral lorquini 1 5485452 594758 Limenitus 21/07/09 Lorquin's Admiral lorquini 1 5498747 584974 Limenitus 22/07/09 Lorquin's Admiral lorquini 2 5477192 589587 Limenitus 22/07/09 Lorquin's Admiral lorquini 1 5477650 589675 Limenitus 22/07/09 Lorquin's Admiral lorquini 1 5477532 589870 Phycoides 17/07/09 Mylitta Crescent mylitta 3 5470852 591240 Phycoides 17/07/09 Mylitta Crescent mylitta 1 5470668 591194 Phycoides 17/07/09 Mylitta Crescent mylitta 1 5470561 591197 Phycoides 17/07/09 Mylitta Crescent mylitta 3 5470299 591386 Phycoides 17/07/09 Mylitta Crescent mylitta 2 5470121 591513 Phycoides 17/07/09 Mylitta Crescent mylitta 3 5469973 591584 Phycoides 17/07/09 Mylitta Crescent mylitta 4 5469792 591640 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5481976 589741

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 42

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Phycoides 18/07/09 Mylitta Crescent mylitta 1 5481725 589765 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5481524 589814 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5481460 589922 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5481038 589866 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5480748 589912 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5480355 590034 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5480107 590415 Phycoides 18/07/09 Mylitta Crescent mylitta 4 5480207 590669 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5480241 591055 Phycoides 18/07/09 Mylitta Crescent mylitta 5 5480243 591205 Phycoides 18/07/09 Mylitta Crescent mylitta 4 5480054 592133 Phycoides 18/07/09 Mylitta Crescent mylitta 4 5479946 592525 Phycoides 19/07/09 Mylitta Crescent mylitta 2 5483522 590051 Phycoides 19/07/09 Mylitta Crescent mylitta 2 5483034 590179 Phycoides 20/07/09 Mylitta Crescent mylitta 1 5488444 590715 Phycoides 20/07/09 Mylitta Crescent mylitta 2 5488637 590697 Phycoides 20/07/09 Mylitta Crescent mylitta 6 5487804 592116 Phycoides 20/07/09 Mylitta Crescent mylitta 5 5487552 592611 Phycoides 21/07/09 Mylitta Crescent mylitta 2 5488906 590206 Phycoides 21/07/09 Mylitta Crescent mylitta 6 5488972 590087 Phycoides 22/07/09 Mylitta Crescent mylitta 1 5487950 590302 Phycoides 22/07/09 Mylitta Crescent mylitta 4 5475894 590158

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 43

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Phycoides 22/07/09 Mylitta Crescent mylitta 10 5475871 590245 Phycoides 22/07/09 Mylitta Crescent mylitta 6 5476575 590187 Phycoides 17/07/09 Mylitta Crescent mylitta 1 5470743 591237 Phycoides 17/07/09 Mylitta Crescent mylitta 1 5470697 591262 Phycoides 17/07/09 Mylitta Crescent mylitta 2 5470765 591281 Phycoides 17/07/09 Mylitta Crescent mylitta 1 5470893 591400 Phycoides 17/07/09 Mylitta Crescent mylitta 1 5470831 591441 Phycoides 17/07/09 Mylitta Crescent mylitta 1 5470723 591515 Phycoides 17/07/09 Mylitta Crescent mylitta 1 5470411 591227 Phycoides 17/07/09 Mylitta Crescent mylitta 1 5470127 591520 Phycoides 17/07/09 Mylitta Crescent mylitta 2 5469749 591663 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5482480 589513 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5482021 589737 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5481539 589852 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5481432 589970 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5481076 589895 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5480568 589941 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5480160 590248 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5480107 590381 Phycoides 18/07/09 Mylitta Crescent mylitta 2 5480244 591098 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5480242 591351 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5480056 592144

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 44

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Phycoides 18/07/09 Mylitta Crescent mylitta 1 5479996 592212 Phycoides 18/07/09 Mylitta Crescent mylitta 2 5479951 592267 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5479926 592333 Phycoides 18/07/09 Mylitta Crescent mylitta 3 5479893 592442 Phycoides 18/07/09 Mylitta Crescent mylitta 1 5479899 592695 Phycoides 19/07/09 Mylitta Crescent mylitta 1 5484268 589658 Phycoides 19/07/09 Mylitta Crescent mylitta 3 5484202 589711 Phycoides 19/07/09 Mylitta Crescent mylitta 1 5484021 589790 Phycoides 19/07/09 Mylitta Crescent mylitta 2 5483752 589786 Phycoides 19/07/09 Mylitta Crescent mylitta 4 5483573 589830 Phycoides 19/07/09 Mylitta Crescent mylitta 2 5483284 589759 Phycoides 19/07/09 Mylitta Crescent mylitta 2 5484660 589842 Phycoides 19/07/09 Mylitta Crescent mylitta 7 5484367 590080 Phycoides 19/07/09 Mylitta Crescent mylitta 3 5484264 590115 Phycoides 19/07/09 Mylitta Crescent mylitta 2 5483954 590114 Phycoides 19/07/09 Mylitta Crescent mylitta 2 5483725 590164 Phycoides 19/07/09 Mylitta Crescent mylitta 1 5483609 590134 Phycoides 19/07/09 Mylitta Crescent mylitta 1 5483184 590082 Phycoides 19/07/09 Mylitta Crescent mylitta 4 5482968 589976 Phycoides 19/07/09 Mylitta Crescent mylitta 3 5483062 590191 Phycoides 20/07/09 Mylitta Crescent mylitta 2 5488132 590051 Phycoides 20/07/09 Mylitta Crescent mylitta 3 5488715 590520

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 45

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Phycoides 20/07/09 Mylitta Crescent mylitta 5 5488482 590697 Phycoides 20/07/09 Mylitta Crescent mylitta 5 5488883 590938 Phycoides 20/07/09 Mylitta Crescent mylitta 1 5488033 591463 Phycoides 20/07/09 Mylitta Crescent mylitta 2 5487864 592001 Phycoides 20/07/09 Mylitta Crescent mylitta 1 5486952 593238 Phycoides 21/07/09 Mylitta Crescent mylitta 2 5488225 589675 Phycoides 21/07/09 Mylitta Crescent mylitta 2 5489005 590161 Phycoides 21/07/09 Mylitta Crescent mylitta 3 5488645 589585 Phycoides 22/07/09 Mylitta Crescent mylitta 1 5487900 590261 Phycoides 22/07/09 Mylitta Crescent mylitta 5 5488087 590342 Phycoides 22/07/09 Mylitta Crescent mylitta 3 5488083 590605 Phycoides 22/07/09 Mylitta Crescent mylitta 1 5476946 589401 Phycoides 22/07/09 Mylitta Crescent mylitta 3 5477020 589439 Phycoides 22/07/09 Mylitta Crescent mylitta 1 5477192 589587 Phycoides 22/07/09 Mylitta Crescent mylitta 10 5477502 589791 Phycoides 22/07/09 Mylitta Crescent mylitta 5 5477650 589675 Phycoides 22/07/09 Mylitta Crescent mylitta 5 5478054 589457 Phycoides 22/07/09 Mylitta Crescent mylitta 2 5476349 590096 Phycoides 22/07/09 Mylitta Crescent mylitta 3 5476098 590123 Phycoides 22/07/09 Mylitta Crescent mylitta 5 5475817 590098 Phycoides 22/07/09 Mylitta Crescent mylitta 4 5475799 590199 Phycoides 22/07/09 Mylitta Crescent mylitta 8 5475603 590239

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 46

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Phycoides 22/07/09 Mylitta Crescent mylitta 10 5475612 590289 Phycoides 22/07/09 Mylitta Crescent mylitta 1 5477152 590059 Phycoides 22/07/09 Mylitta Crescent mylitta 5 5476650 590157 Phycoides 22/07/09 Mylitta Crescent mylitta 1 5476429 590228 Phycoides 22/07/09 Mylitta Crescent mylitta 4 5477560 589865 Phycoides 19/07/09 Myllita Crescent mylitta 3 5484317 589585 Phycoides 19/07/09 Myllita Crescent mylitta 8 5484135 589725 Phycoides 19/07/09 Myllita Crescent mylitta 3 5483859 589841 Phycoides 19/07/09 Myllita Crescent mylitta 5 5483603 589830 Papilio 19/07/09 Pale Swallowtail eurymedon 1 5483654 590136 Papilio 19/07/09 Pale Swallowtail eurymedon 1 5483299 590127 Papilio 19/07/09 Pale Swallowtail eurymedon 1 5482738 590078 Papilio 19/07/09 Pale Swallowtail eurymedon 1 5483609 590134 Papilio 19/07/09 Pale Swallowtail eurymedon 1 5483553 590122 Papilio 19/07/09 Pale Swallowtail eurymedon 1 5483287 590108 Papilio 19/07/09 Pale Swallowtail eurymedon 1 5482536 590035 Papilio 19/07/09 Pale Swallowtail eurymedon 1 5482519 590199 Papilio 20/07/09 Pale Swallowtail eurymedon 1 5488194 590096 Papilio 20/07/09 Pale Swallowtail eurymedon 1 5488642 590768 Papilio 20/07/09 Pale Swallowtail eurymedon 1 5486021 593890 Papilio 20/07/09 Pale Swallowtail eurymedon 1 5488132 590051 Papilio 20/07/09 Pale Swallowtail eurymedon 1 5488208 591375

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 47

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Papilio 20/07/09 Pale Swallowtail eurymedon 1 5485806 594154 Papilio 21/07/09 Pale Swallowtail eurymedon 1 5498722 584978 Papilio 21/07/09 Pale Swallowtail eurymedon 1 5493791 585272 Papilio 22/07/09 Pale Swallowtail eurymedon 1 5477570 589736 Papilio 22/07/09 Pale Swallowtail eurymedon 1 5477347 589867 Papilio 22/07/09 Pale Swallowtail eurymedon 1 5476429 590228 Parnassius 04/07/09 Parnassian spp. spp. 1 5467873 566305 Neophasia 19/07/09 Pine White menapia 1 5484633 589841 Neophasia 21/07/09 Pine White menapia 1 5488972 590216 Neophasia 21/07/09 Pine White menapia 2 5489007 590245 Neophasia 21/07/09 Pine White menapia 1 5488919 589941 probable Hydaspe Speyeria 19/07/09 Fritillary hydaspe 1 5483006 589981 Lycaena 18/07/09 Purplish Copper helloides 1 5479060 593790 Rocky Mountain Parnassius 05/07/09 Parnassian smintheus 1 5477209 582311 05/07/09 Swallowtail Papilio 1 5477263 582198 03/07/09 Swallowtail spp. Papilio spp. 1 5461235 575735 04/07/09 Swallowtail spp. Papilio spp. 1 5466675 567923 04/07/09 Swallowtail spp. Papilio spp. 1 5468841 564773 Unknown (not caught- small brown/orangy 04/07/09 butterfly) 1 5467894 566254 18/07/09 Unknown Anglewing 1 5480244 591098 Unknown 18/07/09 Anglewing/Tortoiseshell 1 5480135 590529 Unknown 18/07/09 Comma/Tortoiseshell 1 5482053 589711 19/07/09 Unknown frittilary 1 5483259 590121 Unknown Frittilary (not 22/07/09 caught) 1 5477230 590045

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 48

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Celestrina 05/07/09 Western Spring Azure ladon 1 5471704 583068 Papilio 04/07/09 Western Tiger Swallowtail rutulus 1 5467778 566688 Papilio 04/07/09 Western Tiger Swallowtail rutulus 1 5468910 564574 Papilio 05/07/09 Western Tiger Swallowtail rutulus 5469894 584327 Papilio 05/07/09 Western Tiger Swallowtail rutulus 1 5472180 582780 Papilio 05/07/09 Western Tiger Swallowtail rutulus 1 5471720 583047 Papilio 05/07/09 Western Tiger Swallowtail rutulus 1 5470198 584235 Papilio 05/07/09 Western Tiger Swallowtail rutulus 1 5470200 584234 Papilio 05/07/09 Western Tiger Swallowtail rutulus 1 5471482 583395 Papilio 17/07/09 Western Tiger Swallowtail rutulus 1 5470867 591239 Papilio 17/07/09 Western Tiger Swallowtail rutulus 1 5470553 591180 Papilio 17/07/09 Western Tiger Swallowtail rutulus 1 5469023 592098 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5481725 589765 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5481038 589866 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5480748 589912 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5480194 591602 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5480054 592133 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5482053 589711 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5482021 589737 Papilio 18/07/09 Western Tiger Swallowtail rutulus 2 5481818 589804 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5481539 589852 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5481432 589970

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 49

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5480781 589915 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5480255 590865 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5480057 592030 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5480056 592144 Papilio 18/07/09 Western Tiger Swallowtail rutulus 1 5479951 592267 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5484317 589585 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5484135 589725 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5483603 589830 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5483481 590071 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5483087 590051 Papilio 19/07/09 Western Tiger Swallowtail rutulus 3 5482541 590049 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5482104 590288 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5484450 589311 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5484378 589530 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5484268 589658 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5484595 589916 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5484454 590007 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5484264 590115 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5483725 590164 Papilio 19/07/09 Western Tiger Swallowtail rutulus 2 5483507 590080 Papilio 19/07/09 Western Tiger Swallowtail rutulus 2 5483251 590147 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5483184 590082

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 50

Appendix

Incidental Butterfly Observations UTM Zone 10U; UTM Scientific Number of Datum 83 Date English Name Name observations UTM UTM Northing Easting Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5482799 589899 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5482991 590076 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5483021 590165 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5483062 590191 Papilio 19/07/09 Western Tiger Swallowtail rutulus 2 5482970 590205 Papilio 19/07/09 Western Tiger Swallowtail rutulus 1 5482690 590176 Papilio 20/07/09 Western Tiger Swallowtail rutulus 1 5488545 590336 Papilio 20/07/09 Western Tiger Swallowtail rutulus 1 5488033 591463 Papilio 20/07/09 Western Tiger Swallowtail rutulus 1 5487608 592526 Papilio 20/07/09 Western Tiger Swallowtail rutulus 1 5487347 592843 Papilio 20/07/09 Western Tiger Swallowtail rutulus 1 5486868 593269 Papilio 20/07/09 Western Tiger Swallowtail rutulus 1 5485452 594758 Papilio 21/07/09 Western Tiger Swallowtail rutulus 1 5498848 584950 Papilio 21/07/09 Western Tiger Swallowtail rutulus 1 5498917 584953 Papilio 21/07/09 Western Tiger Swallowtail rutulus 1 5494716 585229 Papilio 22/07/09 Western Tiger Swallowtail rutulus 1 5475850 590180 Papilio 22/07/09 Western Tiger Swallowtail rutulus 1 5487908 590285 Papilio 22/07/09 Western Tiger Swallowtail rutulus 1 5476946 589401 Papilio 22/07/09 Western Tiger Swallowtail rutulus 1 5477347 589867 Western Tiger Papilio 04/07/09 Swallowtail? rutulus 1 5467874 566307

Surveys for Dun Skipper (Euphyes vestris) in the Harrison Lake Area, British Columbia, July 2009 51