A Cladistic Analysis of Graecopithecus
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Dryopithecins, Darwin, de Bonis, and the European origin of the African apes and human clade David R. BEGUN University of Toronto, Department of Anthropology, 19 Russell Street, Toronto, ON, M5S 2S2 (Canada) [email protected] Begun R. D. 2009. — Dryopithecins, Darwin, de Bonis, and the European origin of the African apes and human clade. Geodiversitas 31 (4) : 789-816. ABSTRACT Darwin famously opined that the most likely place of origin of the common ancestor of African apes and humans is Africa, given the distribution of its liv- ing descendents. But it is infrequently recalled that immediately afterwards, Darwin, in his typically thorough and cautious style, noted that a fossil ape from Europe, Dryopithecus, may instead represent the ancestors of African apes, which dispersed into Africa from Europe. Louis de Bonis and his collaborators were the fi rst researchers in the modern era to echo Darwin’s suggestion about apes from Europe. Resulting from their spectacular discoveries in Greece over several decades, de Bonis and colleagues have shown convincingly that African KEY WORDS ape and human clade members (hominines) lived in Europe at least 9.5 million Mammalia, years ago. Here I review the fossil record of hominoids in Europe as it relates to Primates, Dryopithecus, the origins of the hominines. While I diff er in some details with Louis, we are Hispanopithecus, in complete agreement on the importance of Europe in determining the fate Rudapithecus, of the African ape and human clade. Th ere is no doubt that Louis de Bonis is Ouranopithecus, hominine origins, a pioneer in advancing our understanding of this fascinating time in our evo- new subtribe. -
Human Evolution: a Paleoanthropological Perspective - F.H
PHYSICAL (BIOLOGICAL) ANTHROPOLOGY - Human Evolution: A Paleoanthropological Perspective - F.H. Smith HUMAN EVOLUTION: A PALEOANTHROPOLOGICAL PERSPECTIVE F.H. Smith Department of Anthropology, Loyola University Chicago, USA Keywords: Human evolution, Miocene apes, Sahelanthropus, australopithecines, Australopithecus afarensis, cladogenesis, robust australopithecines, early Homo, Homo erectus, Homo heidelbergensis, Australopithecus africanus/Australopithecus garhi, mitochondrial DNA, homology, Neandertals, modern human origins, African Transitional Group. Contents 1. Introduction 2. Reconstructing Biological History: The Relationship of Humans and Apes 3. The Human Fossil Record: Basal Hominins 4. The Earliest Definite Hominins: The Australopithecines 5. Early Australopithecines as Primitive Humans 6. The Australopithecine Radiation 7. Origin and Evolution of the Genus Homo 8. Explaining Early Hominin Evolution: Controversy and the Documentation- Explanation Controversy 9. Early Homo erectus in East Africa and the Initial Radiation of Homo 10. After Homo erectus: The Middle Range of the Evolution of the Genus Homo 11. Neandertals and Late Archaics from Africa and Asia: The Hominin World before Modernity 12. The Origin of Modern Humans 13. Closing Perspective Glossary Bibliography Biographical Sketch Summary UNESCO – EOLSS The basic course of human biological history is well represented by the existing fossil record, although there is considerable debate on the details of that history. This review details both what is firmly understood (first echelon issues) and what is contentious concerning humanSAMPLE evolution. Most of the coCHAPTERSntention actually concerns the details (second echelon issues) of human evolution rather than the fundamental issues. For example, both anatomical and molecular evidence on living (extant) hominoids (apes and humans) suggests the close relationship of African great apes and humans (hominins). That relationship is demonstrated by the existing hominoid fossil record, including that of early hominins. -
Unravelling the Positional Behaviour of Fossil Hominoids: Morphofunctional and Structural Analysis of the Primate Hindlimb
ADVERTIMENT. Lʼaccés als continguts dʼaquesta tesi queda condicionat a lʼacceptació de les condicions dʼús establertes per la següent llicència Creative Commons: http://cat.creativecommons.org/?page_id=184 ADVERTENCIA. El acceso a los contenidos de esta tesis queda condicionado a la aceptación de las condiciones de uso establecidas por la siguiente licencia Creative Commons: http://es.creativecommons.org/blog/licencias/ WARNING. The access to the contents of this doctoral thesis it is limited to the acceptance of the use conditions set by the following Creative Commons license: https://creativecommons.org/licenses/?lang=en Doctorado en Biodiversitat Facultad de Ciènces Tesis doctoral Unravelling the positional behaviour of fossil hominoids: Morphofunctional and structural analysis of the primate hindlimb Marta Pina Miguel 2016 Memoria presentada por Marta Pina Miguel para optar al grado de Doctor por la Universitat Autònoma de Barcelona, programa de doctorado en Biodiversitat del Departamento de Biologia Animal, de Biologia Vegetal i d’Ecologia (Facultad de Ciències). Este trabajo ha sido dirigido por el Dr. Salvador Moyà Solà (Institut Català de Paleontologia Miquel Crusafont) y el Dr. Sergio Almécija Martínez (The George Washington Univertisy). Director Co-director Dr. Salvador Moyà Solà Dr. Sergio Almécija Martínez A mis padres y hermana. Y a todas aquelas personas que un día decidieron perseguir un sueño Contents Acknowledgments [in Spanish] 13 Abstract 19 Resumen 21 Section I. Introduction 23 Hominoid positional behaviour The great apes of the Vallès-Penedès Basin: State-of-the-art Section II. Objectives 55 Section III. Material and Methods 59 Hindlimb fossil remains of the Vallès-Penedès hominoids Comparative sample Area of study: The Vallès-Penedès Basin Methodology: Generalities and principles Section IV. -
Chapter 1 - Introduction
EURASIAN MIDDLE AND LATE MIOCENE HOMINOID PALEOBIOGEOGRAPHY AND THE GEOGRAPHIC ORIGINS OF THE HOMININAE by Mariam C. Nargolwalla A thesis submitted in conformity with the requirements for the degree of Doctor of Philosophy Graduate Department of Anthropology University of Toronto © Copyright by M. Nargolwalla (2009) Eurasian Middle and Late Miocene Hominoid Paleobiogeography and the Geographic Origins of the Homininae Mariam C. Nargolwalla Doctor of Philosophy Department of Anthropology University of Toronto 2009 Abstract The origin and diversification of great apes and humans is among the most researched and debated series of events in the evolutionary history of the Primates. A fundamental part of understanding these events involves reconstructing paleoenvironmental and paleogeographic patterns in the Eurasian Miocene; a time period and geographic expanse rich in evidence of lineage origins and dispersals of numerous mammalian lineages, including apes. Traditionally, the geographic origin of the African ape and human lineage is considered to have occurred in Africa, however, an alternative hypothesis favouring a Eurasian origin has been proposed. This hypothesis suggests that that after an initial dispersal from Africa to Eurasia at ~17Ma and subsequent radiation from Spain to China, fossil apes disperse back to Africa at least once and found the African ape and human lineage in the late Miocene. The purpose of this study is to test the Eurasian origin hypothesis through the analysis of spatial and temporal patterns of distribution, in situ evolution, interprovincial and intercontinental dispersals of Eurasian terrestrial mammals in response to environmental factors. Using the NOW and Paleobiology databases, together with data collected through survey and excavation of middle and late Miocene vertebrate localities in Hungary and Romania, taphonomic bias and sampling completeness of Eurasian faunas are assessed. -
Origin of Language
Origin of language The origin of language in the human species has been the topic of scholarly discussions for several centuries. There is no consensus on the origin or age of human language. The topic is difficult to study because of the lack of direct evidence. Consequently, scholars wishing to study the origins of language must draw inferences from other kinds of evidence such as the fossil record, archaeological evidence, contemporary language diversity, studies of language acquisition, and comparisons between human language and systems of communication existing among other animals (particularly other primates). Many argue that the origins of language probably relate closely to the origins of modern human behavior, but there is little agreement about the implications and directionality of this connection. This shortage of empirical evidence has led many scholars to regard the entire topic as unsuitable for serious study. In 1866, the Linguistic Society of Paris banned any existing or future debates on the subject, a prohibition which remained influential across much of the western world until late in the twentieth century.[1] Today, there are various hypotheses about how, why, when, and where language might have emerged.[2] Despite this, there is scarcely more agreement today than a hundred years ago, when Charles Darwin's theory of evolution by natural selection provoked a rash of armchair speculation on the topic.[3] Since the early 1990s, however, a number of linguists, archaeologists, psychologists, anthropologists, and others -
Full Issue 114
South African Journal of Science volume 114 number 5/6 Decolonising engineering in South Africa The Grootfontein aquifer: Governance of a hydro-social system Household food waste disposal in South Africa South African behavioural ecology research in a global perspective Volume 114 EDITOR-IN-CHIEF Number 5/6 John Butler-Adam Academy of Science of South Africa May/June 2018 MANAGING EDITOR Linda Fick Academy of Science of South Africa South African ONLINE PUBLISHING Journal of Science SYSTEMS ADMINISTRATOR Nadine Wubbeling Academy of Science of South Africa ONLINE PUBLISHING ADMINISTRATOR Sbonga Dlamini eISSN: 1996-7489 Academy of Science of South Africa ASSOCIATE EDITORS Nicolas Beukes Leader Department of Geology, University of Johannesburg The Fourth Industrial Revolution and education John Butler-Adam .................................................................................................................... 1 Chris Chimimba Department of Zoology and Entomology, University of Pretoria Book Review Towards human development friendly universities Linda Chisholm Merridy Wilson-Strydom .......................................................................................................... 2 Centre for Education Rights and Transformation, University of A new look at Old Fourlegs Johannesburg Brian W. van Wilgen ................................................................................................................. 4 Teresa Coutinho Department of Microbiology and Commentary Plant Pathology, University of Pretoria Decolonising -
The Mysterious Phylogeny of Gigantopithecus
Kimberly Nail Department ofAnthropology University ofTennessee - Knoxville The Mysterious Phylogeny of Gigantopithecus Perhaps the most questionable attribute given to Gigantopithecus is its taxonomic and phylogenetic placement in the superfamily Hominoidea. In 1935 von Koenigswald made the first discovery ofa lower molar at an apothecary in Hong Kong. In a mess of"dragon teeth" von Koenigswald saw a tooth that looked remarkably primate-like and purchased it; this tooth would later be one offour looked at by a skeptical friend, Franz Weidenreich. It was this tooth that von Koenigswald originally used to name the species Gigantopithecus blacki. Researchers have only four mandibles and thousands ofteeth which they use to reconstruct not only the existence ofthis primate, but its size and phylogeny as well. Many objections have been raised to the past phylogenetic relationship proposed by Weidenreich, Woo, and von Koenigswald that Gigantopithecus was a forerunner to the hominid line. Some suggest that researchers might be jumping the gun on the size attributed to Gigantopithecus (estimated between 10 and 12 feet tall); this size has perpetuated the idea that somehow Gigantopithecus is still roaming the Himalayas today as Bigfoot. Many researchers have shunned the Bigfoot theory and focused on the causes of the animals extinction. It is my intention to explain the theories ofthe past and why many researchers currently disagree with them. It will be necessary to explain how the researchers conducted their experiments and came to their conclusions as well. The Research The first anthropologist to encounter Gigantopithecus was von Koenigswald who happened upon them in a Hong Kong apothecary selling "dragon teeth." Due to their large size one may not even have thought that they belonged to any sort ofprimate, however von Koenigswald knew better because ofthe markings on the molar. -
A Unique Middle Miocene European Hominoid and the Origins of the Great Ape and Human Clade Salvador Moya` -Sola` A,1, David M
A unique Middle Miocene European hominoid and the origins of the great ape and human clade Salvador Moya` -Sola` a,1, David M. Albab,c, Sergio Alme´ cijac, Isaac Casanovas-Vilarc, Meike Ko¨ hlera, Soledad De Esteban-Trivignoc, Josep M. Roblesc,d, Jordi Galindoc, and Josep Fortunyc aInstitucio´Catalana de Recerca i Estudis Avanc¸ats at Institut Catala`de Paleontologia (ICP) and Unitat d’Antropologia Biolo`gica (Dipartimento de Biologia Animal, Biologia Vegetal, i Ecologia), Universitat Auto`noma de Barcelona, Edifici ICP, Campus de Bellaterra s/n, 08193 Cerdanyola del Valle`s, Barcelona, Spain; bDipartimento di Scienze della Terra, Universita`degli Studi di Firenze, Via G. La Pira 4, 50121 Florence, Italy; cInstitut Catala`de Paleontologia, Universitat Auto`noma de Barcelona, Edifici ICP, Campus de Bellaterra s/n, 08193 Cerdanyola del Valle`s, Barcelona, Spain; and dFOSSILIA Serveis Paleontolo`gics i Geolo`gics, S.L. c/ Jaume I nu´m 87, 1er 5a, 08470 Sant Celoni, Barcelona, Spain Edited by David Pilbeam, Harvard University, Cambridge, MA, and approved March 4, 2009 (received for review November 20, 2008) The great ape and human clade (Primates: Hominidae) currently sediments by the diggers and bulldozers. After 6 years of includes orangutans, gorillas, chimpanzees, bonobos, and humans. fieldwork, 150 fossiliferous localities have been sampled from the When, where, and from which taxon hominids evolved are among 300-m-thick local stratigraphic series of ACM, which spans an the most exciting questions yet to be resolved. Within the Afro- interval of 1 million years (Ϸ12.5–11.3 Ma, Late Aragonian, pithecidae, the Kenyapithecinae (Kenyapithecini ؉ Equatorini) Middle Miocene). -
8. Primate Evolution
8. Primate Evolution Jonathan M. G. Perry, Ph.D., The Johns Hopkins University School of Medicine Stephanie L. Canington, B.A., The Johns Hopkins University School of Medicine Learning Objectives • Understand the major trends in primate evolution from the origin of primates to the origin of our own species • Learn about primate adaptations and how they characterize major primate groups • Discuss the kinds of evidence that anthropologists use to find out how extinct primates are related to each other and to living primates • Recognize how the changing geography and climate of Earth have influenced where and when primates have thrived or gone extinct The first fifty million years of primate evolution was a series of adaptive radiations leading to the diversification of the earliest lemurs, monkeys, and apes. The primate story begins in the canopy and understory of conifer-dominated forests, with our small, furtive ancestors subsisting at night, beneath the notice of day-active dinosaurs. From the archaic plesiadapiforms (archaic primates) to the earliest groups of true primates (euprimates), the origin of our own order is characterized by the struggle for new food sources and microhabitats in the arboreal setting. Climate change forced major extinctions as the northern continents became increasingly dry, cold, and seasonal and as tropical rainforests gave way to deciduous forests, woodlands, and eventually grasslands. Lemurs, lorises, and tarsiers—once diverse groups containing many species—became rare, except for lemurs in Madagascar where there were no anthropoid competitors and perhaps few predators. Meanwhile, anthropoids (monkeys and apes) emerged in the Old World, then dispersed across parts of the northern hemisphere, Africa, and ultimately South America. -
Class 2: Early Hominids
Earliest Hominins CHARLES J VELLA, PHD JULY 25 2018 This is latest theory of how Lucy died! We are Mammals 3 defining mammalian traits: hair, mammary glands, homeothermy Mammalian traits show an adaptation for adaptability Miocene era: 23 to 5 Ma, Warmer global period • Ape grade: Planet of the apes • Over 30 genera and 100 species of ape – compared with 6 today • Location: Africa and Eurasia Proconsul: 25 to 23 Ma, during Miocene; arboreal quadruped Primates • Larger body size • Larger brain • Complete stereoscopic vision • Longer gestation, infancy, life span • More k-selected (tend towards single offspring) • Greater dependency on learned behavior • More social Great Apes Bonobos and Chimps split ~1 Ma Superfamily: Hominoidea Gibbons, Gorillas, Orangutan, Chimpanzee, Human Greater encephalization (brain to body ratio) = smarter larger body, brachiation, social complexity, lack of tail Why did Newt Gingrich recommend this book to all new politicians? Detailed and thoroughly engrossing account of ape rivalries and coalitions. Machiavellianism: political behavior is rooted at a level of development that is below the cognitive and is as much instinctive as it is learned. de Waal 1982 De Waal: Machiavellian IQ Machiavelli's The Prince: Frans de Waal introduced the term 'Machiavellian Intelligence' to describe the social and political behavior of chimpanzees Social behaviors: reconciliation, alliance, and sabotage Tactical deception in primates: Vervet monkeys use false predator alarm calls to get extra food Chimpanzees use deception to mate with females belonging to alpha male Chimpanzee cultures • Chimp Cultures: shared behaviors in different communities: • pounding actions • fishing; • probing; • forcing • comfort behavior • miscellaneous exploitation of vegetation properties • exploitation of leaf properties; • grooming; • attention-getting. -
Fossil Primates
AccessScience from McGraw-Hill Education Page 1 of 16 www.accessscience.com Fossil primates Contributed by: Eric Delson Publication year: 2014 Extinct members of the order of mammals to which humans belong. All current classifications divide the living primates into two major groups (suborders): the Strepsirhini or “lower” primates (lemurs, lorises, and bushbabies) and the Haplorhini or “higher” primates [tarsiers and anthropoids (New and Old World monkeys, greater and lesser apes, and humans)]. Some fossil groups (omomyiforms and adapiforms) can be placed with or near these two extant groupings; however, there is contention whether the Plesiadapiformes represent the earliest relatives of primates and are best placed within the order (as here) or outside it. See also: FOSSIL; MAMMALIA; PHYLOGENY; PHYSICAL ANTHROPOLOGY; PRIMATES. Vast evidence suggests that the order Primates is a monophyletic group, that is, the primates have a common genetic origin. Although several peculiarities of the primate bauplan (body plan) appear to be inherited from an inferred common ancestor, it seems that the order as a whole is characterized by showing a variety of parallel adaptations in different groups to a predominantly arboreal lifestyle, including anatomical and behavioral complexes related to improved grasping and manipulative capacities, a variety of locomotor styles, and enlargement of the higher centers of the brain. Among the extant primates, the lower primates more closely resemble forms that evolved relatively early in the history of the order, whereas the higher primates represent a group that evolved more recently (Fig. 1). A classification of the primates, as accepted here, appears above. Early primates The earliest primates are placed in their own semiorder, Plesiadapiformes (as contrasted with the semiorder Euprimates for all living forms), because they have no direct evolutionary links with, and bear few adaptive resemblances to, any group of living primates. -
Enamel Distribution Patterns of Extant Human and Hominoid Molars: Occlusal Versus Lateral Enamel Thickness
Bull. Natl. Mus. Nat. Sci., Ser. D, 34 pp. 1–9, December 22, 2008 Enamel Distribution Patterns of Extant Human and Hominoid Molars: Occlusal versus lateral enamel thickness Reiko T. Kono1 and Gen Suwa2 1 Department of Anthropology, National Museum of Nature and Science, Hyakunincho, Shinjuku-ku, Tokyo, 169–0073 Japan E-mail: [email protected] 2 The University Museum, The University of Tokyo, Hongo, Bunkyo-ku, Tokyo, 113–0033 Japan E-mail: [email protected] Abstract Enamel thickness and distribution of the molar crown are important in the study of hominoid and hominid evolution. This is because the 3-dimensional distribution pattern of enamel within the molar crown has been recognized to have significant functional, developmental, and phylogenetic information. In order to analyze aspects of enamel distribution patterns, we quantified average enamel thicknesses separately in the occlusal basin and lateral molar crown, and compared extant great ape and modern human patterns. Interspecific comparisons show that, among modern apes and humans, chimpanzees and bonobos share a unique pattern of molar enamel distribution, in which the occlusal basin enamel is unexpectedly thin. Other taxa share a common enamel distri- bution pattern, greater occlusal than lateral crown average thicknesses, despite the wide range of absolute and relative whole-crown thicknesses. We hypothesize that thin occlusal enamel is a shared derived feature of the genus Pan, possibly an adaptation to relatively soft-fruit frugivory. Key words : Enamel thickness, Hominoid, Molar, Occlusal fovea increase in the application of micro-CT technolo- Introduction gy, a series of investigations has now been pub- Average enamel thickness was initially defined lished that report comparable 3D measures of in 2-dimensional cross sections of molars as AET (Smith et al., 2006; Olejniczak et al., enamel area divided by enamel-dentine junction 2008a, b, c, d).