a -lt"~±~T~ ~~·!c.f
Transactions and Proceedings of the Palaeontological Society of Japan
New Series No. :~2
Palaeontological Society of japan Novl:mber 10, 1958 CONTENTS
TRANSACTIONS Page
348. On the !VIiocene Pectinidae from the Environs of Sendai; Part 12. On Pecten
kagamianus Y oKoY Al\IA ...... 1\oichiro iVlASUDA 271
349. Molluscan Fauna from the Tertiary Formation of Chirai. 1\amisaroma.
Kitamino·kuni, 1-Iokkaido ...... Wataru ll."-SIMOTO and Saburo KANNo 285
350. Late Triassic Palaeoplwms in Japan ...... Akira ToKUYAMA 291
351. An Interesting Crustacean Remain, Ctenocheles sujalmi n. sp. from the
Pakogene of Kyushu. Japan ...... Rikizo IMAIZUMI 299
PROCEEDINGS ...... 305
Systematic Index 307
Index of Fossils ...... 309
President : Hisakatsu Y AnE
Councillors: Kiyoshi AsANo. Riuji ENoo (Business and Accountant). Seido ENoo, 1-Iaruyoshi FuJII'.IOTo (Publication). Shoshiro HANzAwA. Kotara HATAI, Ichiro HAYASAKA (General Affairs), Kazuo HuzioKA, Teiichi KonAYAsHI (Editor), Jiro l\iAKIYAMA, Tatsuro MATSU~IoTo, Masao l\iiNATo, Tokio SHIKA:o.tA, Fuyuji TAKAI (General Affairs). Ryuzo ToRrYAMA
Secretary: Tetsuro I-IANAI
Assistant Secretary: Takeo IcmKAwA
All Communications relating to this Journal should be addressed to the PALAEONTOLOGICAL SOCIETY OF JAPAN Geological Institute, Faculty of Science, University of Tokyo, Japan Trans. Pror. Palaeont. Soc. japan, N. S .. No. 32. pp. 271- 21'\'l. pis. 40~1 I, J'\ov. 10, 1958
:348. ON THE ;\fiOCENI:: PECTINlDAE FROl\1 'rHE ENVIRONS OF SENDAI : PART 12. ON PECTEN KAGAMIANUS YOKOYAMA*
KOICHIRO rviASUDA
Departnwnt of Geology, Faculty of Education. 'Tohoku University
ltittl'llf.t.i/i:rl~f!i*fedf,' Pcctinidac: -'CO) 1~. J>ec/elr kagamimms YoKOYAMA :c ..:n,-c: f.lOI!: Pecten kagamianus YoKOYAMA t t. ft 'L$t.:: r. *?-r.!t!!:l.l· t:, CD~ t~O)~)j>::~ '?~ 'l'M'itH..t.:f.li\1!:. Patinopecten kagamianus lmgamiamrs (YoKOYA:IlA), Pat. kagamiamrs monirmensis MASUDA. n. subsp., Pat. kagamiamrs nimaensis !viASlJDA. n. subsp., Pat. kagamimms permirus (YOKOYAMA) 0trniffifili:-:.J}tt J? ttJ., c. c v:bl'J·? t.:., Ut ~:, v)~ -'< r:::-:>t-L'Jcl~~ rrP. t'rHWHO)OOf*'· it!!'t'l".f.f!'~'"'
Among the fossil Pectinidae knO\Yn Introduction and Acknowledgements from the l'vliocene formations of Japan. Pee/en lwgmnianus is of particular in PPc!en lwf.[amianus was first described terest because of its confusing characters. by YoiwYAMA from the Kimachi forma limited geological range, more or less tion at 1\agami. Kimachi·mura. Yatsuka similar lithology of the formations in gun, Shimanc Prefecture in 1923. Sub which it occurs. and rather wide geo sequently it was reported by him as graphical distribution. Pecten pennirus and Pecten PPrmims Numerous specimens from the Nanao var. paucicostata from ncar .1\anao, Noto formation in Ishikawa Prefecture. and Peninsula, Ishikawa Prefecture (1926). from the J\1oni\\'a and Oido formations Later. the same author (1939) included in Miyagi Prefecture. those preserved Pecten permirus and P. permirus var. in the Institute of Gcologv and Palcon· paucicostata in the synonym of Pectrn tology, Faculty of Science. Tohoku Uni· (Patiuopn·ten ?) lwgamianus YoKOY c\MA, versity, and in the Saito [-lo-on Kai ?vlu· with the statement that it sho\vs con scum. both in Sendai City, and the mate· siderable variation in the number and rials in the Institute of Geology, Faculty size of the radial ribs. l'viATsuMoTo (1930) of Science. University of Tokyo in Tokyo, described Pecten p!icicostu!atus as new were studied by the writer. The results to science from the l\ioniwa formation of examination lead the writer to con in the environs of Senclai City, but this sider that !wgamianus may be classified species was included into the synonym into four subspecies. of Pecten !~agmnianus YoKoYAMA by No In this article the \\Titer describes the MURA (l9AO). These different names sho\Y morphological characteristics of the that the present species exhibit variable kagamianus group. and discusses their characters. relationship with related forms. The * ReceiYed Feb. 5, 1958: read at 66th meet geological significance, so far as can be ing of the Society at Akita. June, 15, 1957. judged from the field data, is also given.
271 272 Koichiro MAsuoA
Acknowledgements are due to Dr. Ko a fe\\· longitudinal striae which are pre tora liATAr of the Department of Geology, sent also on the latter. Shell margin Faculty of Education, Tohoku University. coarsely crenate. The ears seem to be for his supervision. Thanks are due to subequal. though the anteriqr is a little Prof. Sotoji IMAMURA of !he Geological broken. Length and height 105 mm. and ivlineralogical Institute. Faculty of Thickness 25 mm.'' Science, and l\lr. Kazuo 0~-:AllloTo of the Remarks :-This species is characteriz Department of Geology, Faculty of Edu ed by the large. orbicular. compressed. cation. both of the Hiroshima University. subequivalve. equilateral shell. flatly for their kind offer of !he specimens rounded radial ribs which are broader which they collected from the 1\'Iiocene than their interspaces and divided into deposits in Shimane Prefecture, and also a varying number of longitudinal riblets to l\liss Yoko SuzuKI and Mr. K6ki Nr on their backs in the right val \'e. and HEI, students of the Faculty of Educa by the left valve having the rounded tion, Tohoku University, for their assis radial ribs narrower than the interspaces tance in the collection of some specimens. and diYicled into a few riblets by shal low longitudinal furrows. Description The present species has been reported as Pecten. Patinopecten, and Vola and Family Pectinidae HATAI and NisiYAMA (1952) referred it to Lyropecten. However. the genus Lyra Subfamily Pectininae pecten is characterized by having the Genus Patinopec/en DALL. 1898 valyes with a hump or wave on the umbo and with distinct intercalary Patinopecten lwgamimms threads in the interspaces, and by the (YoKOY A~IA). 1923 hinge furnished with three distinct. obli que. divergent cardinal crura. 1923. Peel ell kagamiaiii/S YOKOYAMA, japan. Although the generic position of this jour. Geol. Geogr .. Vol. 2, l'·io. 1. p. H. species is doubtful. the writer refers it pl. 1. figs. la-b. to the genus Patinopecten with query. YoKOYAMA's original description is as The specimens of the present species follows: "Shell large. moderately thick. from the yarious mentioned localities much compressed (the right ,·alve a lit show variable characters, and based upon tle more than the left), orbicular, sub several features they can be classified equivalve and equilateral. Both valves into four subspecies. They are closely radiately ribbed. The ribs on the right related with one another. but may be valve sixteen in number, broad and flatly distinguished in their number and nature rounded, straight. broader than the in of radial ribs. the apical angle and the terspaces and with a varying number of depth of valves (Table 1, and figs. 1. 2, longitudinal striae on them; the inter 3:'. The distinguished ones are Patinopec spaces shallow, usually a little narrower ten kagamirmus kagamianus (YoKOYAMA). than the ribs and also longitudinally Pat. lwgamianus moniwaensis !'vlAsuoA. n. striated. The ribs on the left valve subsp .. Pat. lwgamianus nimaensis .\IAsu about equal in number to those of the DA, n. subsp.. and Pat. lwgamianus per right, straight, rounded. much narrower mirus (YoKoYAMA). respectively. These than the interspaces and furnished with are considered to be geographical or On the l'vliocenc Pectinidae from the Environs of Scndai 273 allopatric forms (lVlAYR, 1942). ly lVIiocene. 1~vpe locality, Geological formation and Depository :-1-Iolotype. Geological Insti Age:- Kagami. Kimachi-mura, Yatsuka tute. Faculty of Science. University of gun. Shimane Prefecture. (Lat. 35'25 N .. Tokyo. Long. 13~·58'1-:.) Kimachi formation. Ear-
,...., E E
~ -= ~ -l
115 8 1u5· 1f1R -v-rus- -uo- 12 :!3 I~ 113 I ~~ 11 100' 16 R F 107 llO ·10 II 105' 2-1 ~ 113 10 100' 17 'I R F 59 60 22 4 100' 26 R 78 I ~~ 7 too· 16 R F 57 56 22 4 100' 23 R 26 11 2.5 I OW lfi R F 38. 5 36 15 3 IOU -19 11 100' 16 I. F 36 3·1 : ~~: ~~ ! l~ I 9!i ·15 10 100" HI L F 32 32 2. 5 lllO • 26 I~ 88 10 I ou' 16 I. I F 93 92 12 !Ott• 21 L S!i 38 10 100· 17 L I F 87 85 ·Hi 21 ·I 100' 16 F 71 70 ' 30 I I }: II 1 Ll 1 5 : ~. 1 ~~: ~r 8!i 8 I 0 0" 1 7 R F ·13 41 ' 17 5 I 05' 24. I. 70 I :lO 5 I 0 0' 18 R F 42 38 15.5 4. 5 100' 2·1 I. f>ll 7 IUO' 17 f 16.5 15.6 ~/,,, r: 2 lliU' 21 L ------5-f' Ll ------'-·oov-1'1-f< G: 127 I:l6 50 I 05' 28 R II 0 39 I too· 12 R G 121 1:.!8 50 J:i 110· 29 7·1 8.5 IIJO" 10 t
ffl.fO. Pecten kagamians YoKOYAMA, NoMURA 100 ,: o A and ONJSJ, japan. jour. Geo/. Geogr., B 0 c Vol. 17, Nos. 3-4. p. 190, pl. 19, fig. 4. " D 1950. Pecten (Lyropecten) kagamianus YoKO· YAMA. KUBOTA. Shinseidai·IIO·I\enkyti, No. 6. p. (.!, pl. 9. tlg. 6:!.
Shell large. moderately thick. com ~0 pressed, orbicular. equilateral except for auricles: right valve slightly less convex than the left or nearly equal; both valves radiately ribbed and forming an angle of about 100, at apex. Right vah·e with 15 to 18, stout, round lopped radial ribs and fine concentric growth lines; radial ribs broader than Text-tlg. l. Graph showing the frequency their interspaces, divided into a number of radial ribs. of fine riblets by shallow longitudinal A-PatinojJecten kagamianus moniu•aensis furrows and rarely dichotomous near MASUDA, n. subsp. from the l\loniwa for the beak with narrow interspaces: longi· mation at Moniwa. Scndai Citr. Miyagi tudinal riblets on the backs of radial Prefecture. ribs appear near upper half of disc and B-Pat. kagamianus kagamirmus (YoKOYA· tend to become distinct towards the MA) from the Moniwa formation :ll Ka· ventral margin; radial ribs at extreme nagasc, Ogawara-m:~"hi, Shibata-gun. Miyagi Prefecture. lateral portions nearly flat, slender and C-Pat. kagamimms permims (YOKOYAMA) weak: interspaces between the radial from the Oiclo formal ion at Iwafune. ribs deep and smooth in the central part and Koganebasama, Wakuya-machi. Tfida of disc. but shallow near the submargins gun, Miyagi Prcfcdurc. and with a single or a few weak inter D-Pat. kagamianus permirus (YoKOYAMA) calary threads: anterior auricle sculp from the Nanao formation at Iwaya, Na tured with several radial threads and nao City. Ishikawa Prefecture. concentric lines. furnished with rather wide and shallow byssal notch. and rath· Patinopecten kagamianus kagamiamts er narro\v byssal area: posterior auricle similar to the anterior in sculpture: (YOKOYAMA), 1923 hinge with wide and shallow resilial pit Plate 40. figures 4, 5. provided with distinct lateral ridges, rather simple cardinal crura, and ill· 1923. Pecten lwgamianus YoKOYAMA, japan. developed ctenolium in young shell. jour. Geol. Geogr., Vol. 2. No. 1. p. 8, Left valve with elevated, round-topped pl. l. figs la-b. radial ribs which are narrower than 1930. Pecten p/icicostulatus MATSUMOTO. Sci. their interspaces in breadth and con· Rep., 'Tolzoku Imp. Uniz• .. Ser. 2, Vol. 13, No. 3, p. 105. pl. 40, fig. 15. centric fine growth lines. ornamented by 1940. Pee/ell (Pecten) kagamianus YoKoYAMA, obtuse network: radial ribs rather sharp NoMURA, Sci. Rep., Tolwku Imp. Univ., near the beak but tend to become round· Ser. 2, Vol. 21, No. 1, p. 16, pl. 2. llg. ed towards the ventral margin. and 15. divided into several, fine riblets by shal· On the l\Iiocene Pectinidae from the Environs of Sc!l!lai 275
llO t2':' 0
0 0 0 i 100 100 ' A " . "' ,..., so •o go • I o • l 00 ...., 2: . :; "' .... A 1: . . ~ 50 • niGHT VALVE X w LEFT VALVE :L ,,.'• "<> 0 6 •• ' {j 20 0 ~
0 10 15 20 •o IS 20 DEPTI1 I MM.) DEPTH ( r-1"'.) Text-figs. 2, 3. Graph showing the relationship between the height and depth of shell. low longitudinal furrows on lower half right valve which has low radial ribs of disc; radial ribs near the submargins carrying two or three longitudinal shal· nearly flat, weak and slender. and with low grooves on their tops. l'atinopecten a single or a few. faint intercalary yamasakii (YoKoYAMA) (19:W) which was threads; lon~itudinal riblets on the backs described from the Miocene Shigarami of radial ribs tend to become distinct formation at Shimosoyama, Shigarami towards I he ventral margin; auricles mura. Kami-Minochi·gun. Nagano Pre sculptured with sF-vera! radial threads fecture. also resembles the present one. and concentric lines. the anterior one a but it possesses a less number of low little larger than the posterior. Interior and rather fiat radial ribs which show surface of both valves gently folded cor tri- or bifurcation. responding to the external sculpture and Pecten (C/zlamys) kagamianus miyagi coarsely crenated at the ventral margin. ensis described by NAKAMURA (l!J.lO) from Dimensions :-Shown in Table 1. the Miocene Otsutsumi formation at 6- Comparison and Affinity:-This species tsutsumi. Taiwa-machi, Kurokawa-gun, resembles Patinopecten iwasakiensis (No l\liyagi Prefecture in the northern border MURA) (EI35l which was described from of Sendai is considered to be a synonymy the Miocene Tanosawa formation at Ho of Nanaochlamys notoensis (YoKoY A lilA) tatezawa. Jwasaki·mura, Nishi-Tsugaru (HATAI and MAsuuA, 1953). gun. Aomori l'refecture. but iwasakiensis In "A Note on Pecten lwgamimws YoKo can be distinguished from the present YAMA IL"-TAI (1938) mentioned that one by the left valve which has the whether Pecten sp. of the luzl[amimzus radial ribs gradually increasing to type reported from the Miocene 1\unnui wards the ventral margin by division formation developed in soutlnvestern and intercalation. no longitudinal riblet Hokkaido by NAGAo and SAsA (1934) is on the backs of radial ribs, and by the identical with 0TuKA's (1936) questionable 276 K6i<.:hiro MAsuDA
kagamiamts from the ivliocene Sugota for mation at Ukibuta. Higashi-Yuri-mura. Yuri-gun. Akita Prefecture. or with No PatinojJccleu kagamianus moniwaensis MURA'S (1935) s-hutaii from the Miocene 1\'lASUDA, 11. SUbSp. Tanosavva formation at 1-lotatezawa. Iwa saki-nmra, Nishi-Tsugaru-gun. Aomori Plate H, figures 3, 4. 5, 6a-b. Prefecture. is a question. The writer 1929. Pecten (I'alinopecten ?) kagamianus Yo considers that 0TuKA's P. aff. kaJ.;amimms KOYAliiA. Imp. Ceo!. Sun'. japan. Rep. may be identical with NoMuRA's species. c\lo. 104. p. 2. pl. 1, tig. 1. Furthermore. 1-IATAI has discussed the 1936. Pee/en (Vola) kagamianus YoKOYAMA, relationship between lwgamianus and s NoMURA and ZINBO. Saito Ifo·Oil f(ai hataii, and has inferred that the rei a! ion ,lfus., Res. Bull.. No. 10. pl. 20. fig. 3. existing between them is an interesting Shell large, moderately thick. com case of adaptation or lateral localization pressed, orbicular. equilateral except for produced as a result of migration. auricles: right Yalve nearly equal to or Remarks :-This subspecies is charac a little less convex than the left; both terized by its large. orbicular, compress valves radiately ribbed and forming an ed shell with 15 to 18. stout, round-topped angle of about 1000 at apex. radial ribs bearing a number of close-set Right valve with about 10. rather low. longitudinal riblets on their backs. by round-topp<.'d radial ribs. a single or a the hinge with rather sirnple cardinal few intercalary threads between radial crura. by the ctenolium in young shells. ribs near the submargins. and line con by the wide and shallow resilial pit centric growth lines: radial ribs much provided with distinct lateral ridges in broader than their interspaces, and di the right valve, and by the left valve vided into numerous. fine riblets by shal v.'hich is ornamented by elevated radial low longitudinal furrows; longitudinal ribs bearing several longitudinal riblels riblets on the backs of radial ribs vari on their backs. and by the interspaces able in strength and number, and first being wider than the radial ribs and appear at about half of disc length and the obtuse network. The right valve is tend to become dis! inct tmYards the ven· usually a little less convex than the left I raJ margin: radial ribs at the lateral one or sometimes subequal to it, but it extremities very low and slender, and seems that the right v:tl vc is sometimes with inlerspaccs variable in breadth: a little more convex than the left as interspaces between the radial ribs shal pointed by YoKoYAMA (19:23). low and smooth in the central part of Described specimens :-1\.anagase. C>ga disc: anterior auricle furnished with wara-machi. Shibata-gun. l'vJiyagi Prefec wide and shallow byssal notch and rathc ture. Calcareous sandstone of the 1\lo er narrow byssal area: posterior auricle niwa formation. DGS. Reg. No. 3518. sculptured with several. weak radial Distrilmtion:-1\.imachi formation in threads and concentric lines: hinge with Shimane Prefecture, Moniwa formation simple cardinal crura. and wide and in Miyagi Prefecture. and Nanao forma shallow resilial pit provided with lateral tion in Ishikawa Prefecture: all Early ridges. and furnished with ill-developed Miocene in age. ctenolium in young shell. Left valve with stout, elevated, round-topped radial ribs and concentric growth lines, and On the Miocene Pectinidac from the Environs of Scndai 277 ornamented by obtuse network: radial probably had a common ancester. ribs rather sharp near the beak, tend to Remarks:-This new subspecies is become rounded towards the ventral characterized by its large, rather thin , margin, and divided into a few riblets shell. 9 to 13. low. round-topped radial· by shallow longitudinal furrows on lower ribs which are much broader than their half of disc: interspaces much broader interspaces and divided into numerous than radial ribs, rather deep and smooth, fine riblets by shallow longitudinal fur and with a single or a few intercalary rows. very slender and low radial ribs at threads in the lateral portions of disc: the lateral portions of disc in the right anterior auricle a little larger than the valve, and by the left valve in haYing posterior. and sculptured with several elevated radial ribs which are much nar weak radial threads and concentric lim·s. rower than their interspaces and divided Interior surface of both valves gently into a few, fine riblets by shallow long-i folded and coarsely crenated at the ven tudinal furrow~. and a few. fine radial tral margin. threads on the surface of auricles. Dimensions :-Shown in Table 1. Type locality. Geological formation and Comparison and :lfji.nity- Pal inopeclr'll .'lf.,'e :-~loniwa. Senclai City, 1\liyagi Pre kagamimzus kagamimws can be distin fl'cture. Lat. :38o13'N .. Long. l-t0',17'E. guished from the present one by having l\loniwa formation. Early 1\liocene. rather thick sl1ell. more numerous ele Depository:- Holot ype, DGS. Reg. No. vated radial ribs with more numerous 3!iOG, Paratypes. DGS, Reg. No. :{S07 and and more distinct radial threads on the No. 1.!76. surface of auricles than that of IIIOilill'a !Jistribution :-1\loniwa formation in l\li· ensis. yagi Prefecture, Nanao formation in Ishikawa Prefecture. and Yanagawa and This subspecies is related with PatiJzo Futatsugoya formations in Fukushima pecten kimurai (YoKoYAMA) (l9~!i) which Prefecture: all Early Miocene in age. was described from the !'vliocene Kame two formation
13. Pt. -1, p. 8118, pl. !)5, tigs. 138, 14·1. than the posterior. Interior surface of both \·alves gently folded and coarsely Shell large, thick, compressed, orbicu· crenated at the Yentral margin. Jar. equilateral except im· auricles; right Dimen:;;ions :-Shown in Table 1. valve a little less con\'eX or subequal Comparison and Affinity:-This sub· to lhe left; both valves radiatelv ribbed species can be distinguished from luz· and forming an angle of about. lOW at gamianus kagamiauus, kagamianus moni· apex. waensis ancllwgamianus nimaensis by the Right valve with 21 to 29. stout. thick shell, larger number of radial ribs, elevated. round-topped radial ribs and characteristics of radial ribs and auri· fine concentric growth lines: radial ribs cles. and the hinge area. broader than their interspaces. and eli· Remarks:- This subspecies is charac· \·ided into several. line riblets bv shal· terized by· its rather thick. compressed low longitudinal furrows; rib lets. on the shell. 21 to 29, elevated. riJund·toppecl backs of radial ribs usually appear on radial ribs which are diYided into seve upper half of disc. and tend to become ral. fine riblets, conspicuous cardinal distinct towards the ventral margin; cura. deep and wide resilial pit provid· radial ribs at the lateral extremities verv ed with distinct lateral ridges, and rath weak and slender: interspaces betwee;l er wide byssal area. the radial ribs deep and smooth. but shallow near the submargins and some· l)pe locality. Geological formation and times with a single or rarely a few, .-'l;,;e:-·fwaya, NanaoCity. Ishikawa Pre very weak intercalary threads; anterior fecture. Lat. 37'111'03'' N .. Long. 136"57' auricle sculplUJed with several. line ra· 0.1·' F. N;Jn:-to formation. Early l\Iiocene. dial threads and concentric I i nes. and DepositOJy:- Lectotype. University of furnished with rather wide and shallow Tokyo (YoKoYAMA. 1926, pl. 2, fig. 1. here byssal notch and rather wide byssal area: designated). posterior one similar to anterior in sculp Described specimens: -lwaya, Nanao ture: hinge with conspicuous cardinal City. Ishikawa Prefecture. Calcareous crura, rarely with rather flat hinge plate sandstone of i'\anao formation. DGS, Reg. \vith very faint striae parallel to the No. 1888. hinge line in full adult specimen, deep Distribution :-Nanao formation in Ishi· and wide rcsilial pit provided with dis· kawa Prefecture and Oido formation in tinct lateral ridges, and ill-developed l'vfi yagi Prefecture: both Early Ivliocene ctenolium in young shell. Left valve in age. with elevated, slender radial ribs and concentric growth lines. and ornamented J\lode of Occurrence and by obtuse network; radial ribs much Geological Significance narrower than their interspaces. divided into a iew, tine riblets by shallow longi The l'vloniwa formation at Moniwa. Sen tudinal furrows on lower half of disc ; dai City ..Miyagi Prefecture where abun radial ribs at lateral extremities verv dant specimens of kagamia nus mouhw weak and slender. and with a single o.r r'nsis occur. consists of granule conglo· a few. line intercalary threads in their merate wit I! some rounded pebbles to l:On· interspaces: auricles sculptured with glomera! ic very coarse-grained sandstone about ten radial threads and concentric containing also Nanaoch/amys ltoloensis lines, and anterior one somewhat larger l YoKoYAMA), Placopecten nomurai J'vlAsu· 280 K6ichir6 1\IASUDA
oA. CryptopecfeH yallagmcaensis {No!\IURA Chlamys kumanodoensis l\IAsuoA, Placopec and ZtNBo). etc .. other pelecypods. gas ten allilzoensis (MATSUMoTo). etc., other tropods. echinoids. balanids. brachiopods pelecypods. gastropods, brachiopods, and bryozoans. Among the pectinid fos corals. echinoids and bryozoan-fragments. si Is. a rather large number of kagamiamts These pectinids occur as isolated valves, moniwaensis consist of somewhat water more or less water-worn, and arranged worn and some \Vith more or less broken parallel with the bedding plane with shells, both of which arc arranged paral 1he convex sides upwards or very rarely lel with the bedding plane. Though do\\'11\vards. some specimens have their concave side Abundant specimens of lwgamianus upwards. or irregular. the majority are /?agamiml/ls from the calcareous sand arranged with their ronvcx side up stone of the l\loniwa formation at Kana wards. Among the pectinids from tl:e gase. Ogawara-machi. Shibata-gun, about :\loniwa formation at l\loniwa. the thin 18 km. south of lVIoniwa, were found as and fragile Placopecten J/01/lll!'lli is sociated with some J>lacoPecten akihoensis, abundant. rather well pi·eservcd and Nanaoclzlamys notoensis, a few Cryptopec occur as isolated or sometime as intact /en yanagmcaensis. and gastropods. bra valves. Nanaochlamys notoensis and chiopods, balanids. echinoids and bryo C1)'jJ!OjJecfen _\'illlagmcaf'l/sis both of which zoan-fragments. These occur as rather occur in association with the mentioned well preserved isolated or very rarely ones occur as isolated valves. but are intact \·alves. arranged parallel with the better preserved than lwgamianu.~ moni bedding plane and with the convex sides waensis. From the lield data at 1\t.>n i wa, upwards. In the case of specimens with it is considered that the shells of kagami intact valves, it is considered that they anus monizcaensis were transported from were probably buried i11 situ. because a distance greater than the other men of their nat ural orientation. tioned species of Pectinidae. From 1he field data. it is probable that Some specimens of lwJ.[amia11us IIJOitiz!'ll the shells of kagamianus kagamiamts at ensis were cullectecl from the ~ranule 1\umanodo were transported from the conglomerate to very coarse-grained site of living to that of burial, while sandstone of the l\-Ioniwa formation at those from 1\anagase lived in the envi Kita-Akaishi. Sendai City, about 3 km. ronment in which they were buried. or west of lVIoniwa. in association with at least they were not carried from a Co•PI oPel en )'111/flgmcaensis, 1'lanaoclzlam_l'S remote place. no/oensis, etc. They occur as isolated From the available data concerning valves. with the convex sides u pwarcls the !\Ion i wa format ion, it is considered or rarely downwards. and are arranged that the depositional conditions at :Moni parallel with the bedding plane. These wa and Kita-Akaishi were somewhat may have been transported from the different from those of Kumanod6 and site of living to that of burial. Kanagase. as can be judged from the Several specimens of kagamianus kaga rock facies and the associated fauna. mianus were collected from the granule Thus it is evident that Patinopecten ka conglomerate of the 1\iloniwa formation ~,;amianus muniwaensis and Pal. l
City, Ishikawa Prefecture from where at conditions at Iwaya, I-Iosoguchi and numerous specimens of lwgamianus per Kokubu were different from that at mints occur. consists of richly calcareous Hiuchidani. sandstone containing Nanaochlamys nuto Several authors (NEWELL. 1948, BuRMA, ensis. Placopecten akilwensis, Chlamys l!l-18, SYlvEsTER-BRADLEY, 1951. et al.) ingeniasa (YoKoYAMA'. etc., foramini maintain that closely related subspecies fers. brachiopods, echinoids. bryozoans, arc not found together in the same bed balanids and sponge spicule:>. All speci at the same place, FLEMING (1957) states mens of kagamianus pennines so far as that different species of Pecten s. str. observed occur as isolated valves ar are seldom sympat ric and there is no ranged parallel with the b2dding plane such record in Australasia. However, and with the convex sides upwards. SnuTo (1957) has discussed the occur Some water-worn shells or fragments arc rence of sympatric species of Paplzia found in association. from the i\Iiyazaki group developed in The specimens from th
Explanation of Plate 40
Figs. Ja-b. Patinopecten kagamiamts permirus (YoKOYAMA), a, Right valve, x3/5. b, Hinge area of fig. Ia, x l. DGS, Reg. No. 1888. Loc. lwaya. Nanao City, Ishikawa Prefecture. Na n;to formation. Fig. 2. Patinopecten kagamia11uspermirus (YoKOYAMA). l{ight valve. x2/3. DGS. Reg. No. 3253. J.oc. Iwafune. Wakuya-machi, Tiida-gun. 1\Tiyagi f'refccture. Oido formal ion. Fig. 3. Patinopeclen kagamianus permirus (YoKOYAMA). Right valve, xl. DGS, I~cg. No. 3333. Loc. Oido, Wakuya-madti, Toda-gun, l\·Iiyagi Prefecture. Oido formation. Fig. ·1. Palinopecten kagamianus kaKtiiiiianus (YoKOYAMA), Right vah·c, ·.: -l/5. DGS. Reg. No. 351S. Loo.:. Kanagase. Ogawara-machi. Shibata-gun, l\liyagi Prefecture. Moniwa forma· tion. Fig. 5. Palinapecten kagamianus ka)(amianus (YoKOYAMA), Left valve, X4/5. DGS. Reg. No. 35Hl. Loo.:. Same as above. MAs uDA: Miocene Pectinidae Plate 40
K. MAsuDA photo. On the Miocene Pectinidae from the Environs of Sendai 283 consisting of sand or shelly sand quite 3) (in Japan<:sc). Shinseidai-no-Kenkyu free from muddy materials and inf'luenc (Cenozoic Research). lVo. 6, pp. 12-18. 2 ed by warm thermal conditions. pis. MASUDA. K. (19a5), Fossil Scallops from the The geological range of the mention Yasawagi-mura. !Iiraga-gun. Akita Pre ed subspecies of kagamianus is rcstri· fecture. Northeast Honshu. Japan. Saito ccd to Early l\liocene in a two-fold cliYi Ho·oll Kai 11fus., Res. Bull.. 1\'o. 2'1. pp. sion. 18-22. -- (1!J56). Some Fossil Pectinidae from the Heferences Oido Formation, Wakuya-machi. Toda gun, Miyagi Prefecture, ?\ortheast Japan. BURMA. B. II. (194S). Studies in Quantitative Ibid.. No. 25. pp. 22-26, 1 pl. Paleontology. I. Some Aspects of the lviATSUMOTo, H. (1930). On the ;-.,[arine Fauna Theory and Practice of Quant ita th-e of Three Fossil Zones of the Upper l\fio Paleontlogy. jour. Pal .. Vol. 22. No. 6. cene of Natori District, ProYince of pp. 725-761. 22 text-figs. Rikuzen. Sci. Rep .. Tolwku Imp. Unit• .. DAt.L, \V. H. (1898), Tertiary Fauna of Flo Ser. 2, Vol. 13. No. 3. pp. 96-109, 2 pis. rida. 1'ra11s. Wagner Free Ins/. Sci .. Vol. HAYR. E. (1942). System'ltics and the Origin S. Pt. 4, pp. 571-916. 13 pls. of Species. Columbia Unit•. Press. pp. 3- FLEMING. C. A. (1957). The Genu:; Pr•clcn in 33·1. New Zealand. N. Z. Gcol. Sun· .. Pal. Bull. NAGAO. T. andY. SASA (1931), Latest Recent 26. pp. 5-69, 15 pis .. 13 text-figs. Geologic History and Cainozoic Forma liANZAWA. s.. K. IlATAI, J. IWAI, N. KITA· tions of the Southern Part of Hokkaiclo MURA and T. SmBATA (1953). The Geo (4) (in Jap:mese). lour. Geo/. Soc.fopan, logy of Sendai and its Em·irons. Sci. Vol. 41. iVo. 488, pp. 211-260. Rep., Tolwku Unit• .. Sf'r. 2, 1-o/. 2.r;, PJl. NAKA~IURA, i\1. (1940). On Sam(' Pectinidac: 1-50. 2 maps. 4 charts. Fossils from the l\lioccnc Deposits of HATAI, K. (1938), A Note on Pecten lwgami- the Tomiya Block. Jviiyagi-ken. Northeast anus YoKOYAl\IA. Bull. Biol{eogr. Soc. Honsyu, Japan. japrm.]our. Ceo/. Geogr .. japan. Vol. 8, No. 6, pp. 103-110 Fol. 11, ·"''os. 1-2. pp. 1-15. 2 pis. -- and S. NrsiYAl\IA (1939), Palaeontological NEWELL. N.D. (1948). Infraspecitic Categories Notes on Certain Japanese Scallops. jour. in Invertebrate Paleontology. jour. Pal., Geol. Soc. Japan, Vol. 46, :Vo. 64,1. pp Vol. 22, No. 2, pp. 225-232. 37-46, 3 texst -figs. NoMURA. S. (1935). Miocene Mollusca from -- (19·11), On Some Fossils from the Oido the Nisi-Tugaru District. Aom·Jri-ken, Shell-Beds developed in Toda-gun, Riku· Northeast Honsyu, Japan. Saito Ho-oll zen Province, Japan. japan. jour. Ceo/. Kai :Hus .. Res Bull. 1Vo 6, pp. 19-81. 'i Geogr., Vol. 18. No. 3, pp. 109-118. l pl. pis. -- and S. NrsiYAMA (1952), Check List of -- and N. ZINBO (1936). Additional Fossii Japanese Tertiary Marine l\lollusca. Sd. l\Iollusca from the Yanagawa Shell Bed;; Rep .. Tolwku Unil•., Ser. 2, Special Vol. No. in the Huku~ima Basin. Northeast Hon 3, pp. l-46-l. syu, Japan. Ibid .. No. 10. pp. 335-3·15, -- and K. MASUDA (1953). On the Miocene 1 pl. Pectinidae from the Etwirons of S.:n lecture. Japan. japan. jour. Geo/. l;eogr .. the United States. Jllim. 11!11s. Roy. Ilist., Vol. 17. J.Yos. 3-4. pp. 181-194. 3 pis. Be/g., Ser. 2, Fasc. 13, pp. 3-76, 6 p!s. OTUKA. Y. (1935). The Oti Graben in South· YoKOYAMA, M. (1923). On Some Fossil l\lol· ern .l\oto Peninsula, Japan. Part 3, Bull. lusca from the Ncogene of Izumo. japan. F:arlhq. Res. Ins!., Vol. 13. Pt. ·I, pp. 8·16- jour. Geol. Geogr .. Vol. 2, No. 1, pp. 1- 909. 5 pis .. 21i tcxt·ligs. !1. 2 pis. (1936), The Takasegawa Grcl'll Tuff -- (1925). 1\Jolluscan I~emains from the Up· Beds of the Yuri District. Akita Pref.. permost Part of the Joban Coal-Field, Japan. and Correlation of the Lower four. Col/. Sci., Imp. Uniz•. Tokyo. Vol. Neogene in Japan. Ibid., Vol. f./. Pt. 3, 45, Arl. 5, pp. 1-32. 6 pis. pp. 483-452. 1 pl. (1925). Tertiary Mollusca from Shinano SmJTo, T. (1957). Fossil Paphia from the J\fi. and Echigo. fou·r. Fac. Sci .. Imp. Univ. yazaki Group. japan. jour. Ceo/. Geogr .. Tokyo, Sec. 2, Vol. 1. Pt. 1. pp. 1-23, 7 Vol. 28. Nos. 1-3, pp. 139-160, I pl. 9 pis. text.figs. (1926). On Some Pliocene Shells from Sn.vF.STER·BRADLEY. P. C. (1951). The Sub. Kaga and 1\;oto. four. Geo/. Soc. Tokyo, species in Palaeontology. Ceo{. Mag., Vol. Vol. 33, No. 3!J I. pp. 9-11, 1 pl. 88. pp. 88-102. (1929). Pliocene Shells from ncar Nanao. TUCKER· HowLAND. I I. I. (1938), The At I antic Imp. Geol. Sun• .. japa11, Rep. No. JO.J, pp. and Gulf Coast. Tertiary Peclinidae of 1-7, G pls. Explanation of Plate 41 Fig. I. Palinopecten kagamiamt!; 11imaeusis MASUDA, n. subsp. Holotypc. Reg. No. T. NM. 26. Geological and Mineralogical Institute. faculty of Science. Hiroshima University. Right valve, x7/8. Loc. Akazaki. Nima-machi, Nima-gun, Shim K. MAsuDA photo. Trans Proc. Palaeont. Soc. Japan. N. S., No. 32, pp. ~85-290. pl. 42, No\'. 10, 1958 :3-Hl. l\lOLLUSCAN FAUNA FROi\'f THE TERTIARY FORMATION OF CHIRAI. KA.?IHSAROMA. KITA!vTINO-KUNI, HOKKAIDO* WATARU HASIMOTO and SABURO KANNO Geological and !vlincralogical Institute, Tokyo University of Education :lt&if!:lt~FlHJt,'WU. J11Jl(H.i'io:>:';i'Lc-~;f:!'IEfUtbltf.r::-?t '1:': :1ftff·o:> I A t~*LUUfiJill:ltQ[]I J:{lr.glll.J, nl*H.iii::J(!i)'i-tt "!> m.::::::>f.tJ. •:.rwt·Ji.l.. l!t1 i '!'t..J:tiH., t.:c i:J·--; 1:' 4>:fl!lhH· G Pecten kanehami t~:lifHt;-~:ttt.: c. cil;zn 'J. :ro)n/tiL:!M!u-+:'1:> t Ul-cl't~oJ-:<. HH'i·l, Ltoc0f;.tft6ltt'i'J;J{.Ht.... u1 1-", t.t lt'fillf0.lt!!Z'{Il!jf'U.: ~~ i:S :::}:'t1fi U.:fY:Icl ;,:, litlllii i' ~'1i t. .lliyagipedcll 0) 1 ~iHl. :1:5 J: C Introduction and Acknowledgements deye]oped in an adjoining area, the senior author visited the above mention In 1951. TAKEDA and KITAr.AwA briefly ed locality**. and he made a collection wrote on the limestone resources in the of the molluscan fossils including some Chirai area. Saroma-machi. 1\itamino interesting scallops described in this kuni. Hokkaido. and referred io a article, though he failed to discover marine Tertiary formation yielding such "Pecte11 lw11eharai YoKoYA:..IA ". molluscan fossils as Pecte11 fumelwmi The writers wish to thank Prof. Ko YoKoYAMA, Pecten spp., and Cardium sp. tora IlATAJ of our Institute for his According to this paleontological data. valuable suggestions. Acknowledgements the senior author correlated this forma arc clue to I'vlessrs. Akitoshi INoMA, tion with the Kawabata which is deve Hisashi Ivl!zuMoTo, and 1-likosuke WATA· loped in Central IIokkaido when he NAnE. students of our Institute. and l'vir. compiled the geological map of !Iok Rikio YAMADA. a farmer residing in kaiclo in the scale of 1: 200.000: this was 1he area of the fossil locality. for their published from the Geological Survey kind assistance in collecting the fossils. of Hokkaido in 1957. Since then. particularly from the in terest of the first report of " Pecten lwneharai '' from the Okhotsk side of Hokkaido and also to its stratigraphic significance in relation with the cold water fauna of the Tokoro formal ion * Received Feb. ~6. 1958; read Feb. ~. 1958. ** Financed by the Science Research Ex penditures Fund of the Ministry of Educa Text-fig. 1. l\laps showing the geographic tion position of the fossil locality. 285 286 Wataru l-TAsJ!It<>To and Saburo KANr;o die to upper Miocene in japan. The fauna Geology and Fauna taken as a whole apparently represents The Tertiary formation in the Chirai a rather warm water assemblage like area is very limited in its development that of the .Nanakita formation in illiya and unconformably overlies the Saroma gi Prefecture. where G!ycymeris matsu group of so-called Mesozoic age: it clips moricnsis J-IAT.~.J and KAKAI\tURA, IHiyagi very gently to the south. pecten matsumoriensis 1\IAsuoA, Chlamys The lithologic succession of the forrm kmtehami (YoKoYAMA). Crene!la I omiyaen tion is as foll.ow: the lo\\·est part con sis I-hTAr ancl NAKAMURA. and Dosinia sists of granule conglomeratic porous lumelwrai YoKOYAMA are characteristic. sandstone of reddish brown tinge main!~· lt may be added that PatinoPecten due to the cernentin:_~ material of iron t:higanouraensis NAKAMURA and Turritella oxide: this is superposed with an irre s-lwtaii NoMURA. have been reported gular alternation of brown sandstone and from the Shiogama 1\Iiocene which is reddish brown conglomerate as a transi considered to be a correlative of the tional part to the next younl!er massiw Nanakita. Patinopecfen clziganouraensis brown sandstone which is in turn con 1\AEA!IIURA is allied to P. lwbiyamai KA formablv o\·erlain by a bed of yellowish MADA. though precise comparison of the light grey tuff. The tirst and the last two is difrlcult O\\·ing to the poor pre· mentioned beds yield molluscan fossils. senation of the former. hut the former is superior to the latter From the abm:e consideration. the in number oi individuals. writers conclude that the Chirai fauna The lowesl. the reddish brown con is a correlative of the Nanakita and the glomeratic sandstone (Loc. I) yielded Shiogama. Therefore, the geological abundant individuals of Glycymcris cf. age of the Chirai fauna may be conside idensis KANNo, Chlamys tSn·iftopedeJI) red as middle Miocene. at least, not Slcijtii (BERNARDI). "PI'Cft'll'' sp., and younger than the upper 1\Iiocene. il1iyagipecten saromensis I-IAst r.toTo and The Chirai formation is distributed in KANNo, n. sp.. among which the last an isolated srrwll area patched in the mentioned is the most predominant. The Toyokoro-hitami tectonic zone in broad pelecypods are largely represented by sense (1-L~sJMOTo. 19S8J \\·here there is isolated valves arranged horizontally no development oi the typical green tuff with the convex surface above. The facies. regardless of that it is situated shell materials of them are hardly pre bet\n.•en the ·· green tuff regions" of the sen-eel. and almost :1!1 are represented Okhotsk side of central Hokkaido and as mou I cis. the Inner zone of the Chishima (Kuricl) The uppermost. yellowish tuii iLoc. 2) geo-tectonic structure. yielded Gfvcymeris d. idf'nsis KA:- NA,;Ao, 1956). Kitami region. In this case, the problem In the east of the Chirai area, there arises as to how was introduced such a develops the oil-bearing Neogene Terti warm fauna into 1he Okhotsk region. ary group which is superposed uncon Presently we have no information on a formably on the late r..Iesozoic deposits. fauna equivalt:'nt with the Chirai from The following geological succession has anywhere in the eastern part of Hok been reported: namely, the Tokoro kaido. formation (or the Kurumatomanai) of clastic sediments is oyerlain by the Systematic Der.cription Abashiri format ion of pyroclastic nature. The former was once considered to be Family Pectinidae of the Oligocene Poronai stage owing Subfamily Pectininac its faunal elements, but recently it is Genus Miyagipeclf'll l\lAsunA. 1952 correlated with the cold water type deposits of the Takinoue stage ISAsA and J\Jiyagipecfell saromensis HAs I :-.toTo INoUE, 1939; Sui MALl A and YAsAKI. l9fi7), and KANNo. n. sp. and is middle and lower l\Iiocene in age. The latter is correlated with the Kawa Plate 42, Figures 1-5. bata formation in central Hokkaido. Shell moderate in size. orbicular. sub The cold water type of the Takinoue equivalve. equilateral. much compressed. stage is represented by the Chikubctsu Right vah-e smooth, sculptured with fauna which includes se,·eral Poronai Yei~Y fine conct:'ntric Jines of growth and elements, while the soul hern one is indistinct radial threads: auricles sub characterized by the warm water Taki equal in size, sculptured with concentric noue fauna which has been correlated growth lines and wide and shallow with the lower Kadonosawa fauna of byssal notch; interior surface inacccs Iwate and l'viiyagi Prefectures. sable. Left valve ornamented with radial According to FuJIE and Uozu~>n (1957), striations and faint concentric growth the Kawabata fauna (in their revised lines; radial striations numbering about meaning) is said to be distinguishable 20, tend to become obsolete on lower from the Takinouc fauna in broad sense half of disc, and associated with an being the younger one, but no descrip intercalary between them: intercalary tion of the faunal assemblage has been radials disappear on lower half of disc published to date. in adult shell, while the main radials Since the Chirai fauna, as already extend to its base, though they are stated, is compared with the Nanakita rather indistinct: radial striations sepa fauna, it is younger than the Takinoue rated bv intcrspaces much broader than fauna and older l han the Togeshita their \\:idth: auricles subequal. though fauna of the upper 1liocene Wakkanai the anterior one is slightly larger than stage (llAsiMOTo, 1950). accordingly it the posterior, marked with about five is considered to be of the Kawabata radial ribs and closely arranged con stage. cent ric growl h lines: internal surface If the foregoing remarks are accepted. slightly grooved in accordance with the it is possible to consider that the surface ornamentation. Height. 66mm.: warmer Kawabata fauna overlaps the length. 67 mm.: hinge-length. 25 mm.: cold Chikubetsu fauna in the southern depth, ca. G mm.; apical angle. JU5° 288 \Vataru HASIMOTo and Saburo K.~NNO measured on a plaster-cast of the halo type (right valve). Patinopecten aff. kobiyamai Comparisoll :-Jlliyagipecten matsumori KA MADA. 195"1 ensis MAsunA, 1952 (~lAstmA, 1952. pp. 252-253, pl. 24. figs. 4-7), is allied the Plate -1~. Figures 8-9. present new species. but the former differs from the latter by having a Remarks :-An incomplete right and larger apical angle. closely arranged left \'alves were examined. Though the and larger number of radial ribs, and present specimens resemble 1\AMAD.••.'s fainter radial threads on the lower half type species in the general features. of the shell of the leit valve. and fur· some differences exist bet ween them. ther. the interspaces between the radial Namely. the shapes of the ribs are threads in the former are narrower than closely similar. especially in that minor in the latter. The surface of the right striations are developed in the interspa valve of 1\IAsunA'S species is ornamented ces of the .radial ribs in the marginal with closely arranged concentric growth part of the left vah·e, however. the lines while in the present new species number of radial ribs of the writers' they are hardly developed. specimens exceed those of the type Remarl?s:-The distinction between :-;pecies: there are more than nine radials the genus Miyagipecfen and Placopecten in the left and right vaJyes whereas has been discussed by 1\IAsuoA (1952, pp. there are only five or six radials on the 251-252). The present new species some· type specimen. Considering the number what resembles Placopecfen allilwensis of radial ribs, the present specimens (IVL~T5UMOTO), (MASUDA, 1952. pp. 2:-i0-251, seem to be a species different from pl. 24, fig. lll3l. in possession of radial KAMADA's. and from general morphology, threads on the right valve. though they a descendant of that species. The pre are hardly developed, and the radial sent specimens resemble Patiuopecten threads on the left which are weakly kimurai (YOI logical !\lap (Jf llokkaido in the scale of Res. /Jull., no. 26. pp. 31-39. pl. -1. 1 : 200. OOIJ (part 5). MATSUMOTO, II. (19:{0), On the Marine fL'I.NZAWA. S .. K. IL>\TAI et al. (1953). Geo· Faunae of Three Fossil Zones of the logy of Sendai and its Ell\·irons. Sci. Rep .. Upper 1\liocene of Xatori District. Pro· Tohoku Unir .. ser. 2, rol. 25. pp. 1-50, vincc of l~ikuzcn. Sci. Rep .. Tohoku Imp. with 2 maps. Unit• .. st·r. 2. l'ol. 13. no. 3. p. 106, pl. 40, HASIMOTO, \V. (1%0). The Geolog~· of the figs. 'i-S. :\l:trginal Part of the Kabato J\lassif (in NAKAMURA. 1\1. (1940). On Some l'cdinidac Japanese). Bull. Geol. Comm. Flokkairlo, Fossils from the l\lioccne Deposits of 110. 13. pp. 1-7. the Tomip Block. l\liyagi·kcn. North· -- (1958). Considt-ration on the Geological e:~st Honshu. japan. fapall. ]our. Geof. History of the Yczo·Saghalien Geosyndi· Geogr .. ro/. 17. 11os. 1-2. pp. 1-15. pis. 1-2. nal :\rca (in Japanese). ]ub. Pub!. Cm11111 .. KoMURA. S. (1935). Miocene :\lolluscs from l'rnf. H. fvJr~roTo's Sixtieth Birthrl. pp. Siogama. 1\ortheast Honshu. Japan. Saito 101-112. Ho·on 1\ai Mus .. Res. Bull.. no. (j, pp. \93 --and S. NA.cAo (1956). The Oil-Indication -231. pis. IG-17. and !\at ural Gass from l\lonlwtsu·gu n, -- (1940). l\Iolluscan Fauna of the 1\loniwa Kitamino-lwni. Hokkaitlo (in japanese). Shell 1\cds Exposed along till' Natori· ]our. japan. Assoc. Pelr. Tech.. rol. 21. no. gawa in the Vicinity of Scndai. Miyagi 3. pp. 102-\0:i. Prefecture. Japan. Sci. Rep .. Tolwku Imp. KA~IADA. Y. (19:i·l). Patinopecten kobiyamai, a l.,'llil' .. SCI'. 2, 1'0/. 21. 110. J. pl. 1, figs. 2, 3: ~ew i\Iioccnc Scalop from the Johan pl. 2. tig . .J. CoaJ.Field, japan. Trans. Prnc. l'alaMnl. SAsA., Y. and T. INOUE (1939). The Strati· Soc. japan. N.S., 110. 15. pp. 173-l'i.S. pl. graphy of the Tertiary Formations on 23. the Environs of i\bashiri. llokkaidn (in KuBOTA.. K. (\!150). A New Species of !'l·clen Japarwsc). jour. ]aj>rm. :lssoc. 1'1'/r. Tech .. (P/acopecten) from the :\eogc·nc: of rol. 7. no. 6, pp. 1-11. Southwesh:rn llokkaido. japan. ;1/in. and SmMADA. T. and K. YAZAKI (I 956). The Geol .. ro/. 3, no. 5. pp. 183-185. pl. 7. figs. Report of the Geological Sur\'cY on the 1-1. Environs of Abashiri·city. Holtkaido (in 11ANSFIELD. W. C. (1936). Stratigraphic ~igni·. ..JapancSL'). ]our. japan ..-lssoc. Pl'lr. Tech., ti Explanation of Plate 42 Figs. 1-5: Miyagipecten saromensis HASIMOTO and KANNO, n. sp. 1. Right valve of a plaster·cast of the holotype. Reg. No. 6195. 2-5. Left valves of a plaster·cast of the paratype. Reg. No. 6196. Fig. 6: Crenella tomiyaensis IIATAt and NAKAMURA. Reg. No. 6199. Figs. 7a-b: Spisula sp. indet. Reg. No. 6201. Figs. 8-9: Pati11oj>eclen aff. kobiyamai KAMAPA 8. Right valve. Reg. No. 6200. 9. Plaster·cast of a left valve, showing the external sculpture. Figs. 10-11: Glycymeris cf. idensis KANNo 10. Plaster·cast of a right valve. 11. Inner mould of a right valve. showing the taxodont teeth, Reg, No. 6l!l8. All figures in natural size liAsi Mo To & KANNo : Mol luscan Fauna from liokkaido Plate 4~ S. .t\oKI photo. Trans. Proc. Palaeont. Soc. Japan, N. S .. No. 32, pp. 291-298, pl. 43, Nov. 10, 1958 350. LATE TRIASSIC PALAEOPHARUS IN JAPAN* AKIRA TOKUY AMA Geological Institute, University of Tokyo *1llr$.=:!ll;'f-O) Palaeopharus: til!¥ • P:l$h~iil~IC:.J: f), i::. O)~:IJ: Cardiniidae 11:.1/.ill,.,, preheterodont (:heterodont >ei.'lll'f;tt:t;..~fifllJ~L' Palaeoplrarus KITTL is an important ported from Maritime Province of member of the Upper Triassic faunas of Siberia. may be a subspecies of oblonga Japan, e. g. the Kochigatani fauna at tus. Sakawa and Sakuradani, the Nabae at 3. ,lfineplwrus triadicus ToKUY AMA repre sents a new subgenus. linking the so Heki. Nabae and Kyowa and the l'vline ca lied Triassic Pleurophorus (\'l AAGEN'S at Mine. It is often accompanied by sense. 1907) with Palileopharus. JV!inetrigortia and "Bakevellia," and indica tes the shallow water facies of an em Table 1. Palaeopltarus in Japan. bayment, inland or open-sea. KoBAYASHI and IcHIKAWA (1951) were the first to Area find this genus in japan at Nabae and Heki. Subsequently another 2 species and 2 varieties were added from Heki Palaeoplwrus by KoBAYASHI and·fcmKAwA (1952), from Nabae by NAKAzAwA (1955) and from oblrmgatus (KOBAYASHI & * • • lclliKAWA) Kyowa by NAKA!'lo (19571. Here the writer describes the following tllree oblongatus buriji KIPARISOVA * - - -- - maizuruensis KonA v ASH! & forms from the l'vline series: ICHIKAWA - * * * 1. obfongatus (KOBAYASHI and IcJIIKAWA) maizuruensis n. subsp. by which is widely known from Sakawa NAKAZAWA - * (KoBAYASHI and ICHIKAWA, 1950), Sa maizuruensis var. inwmurai kuradani (ICHIKAWA, 1954) and Mine NAKANO - - * (KATAYAMA, 1939),- and was referred 11wizuruensis var. flexicoslalus to Pleurophoms, but the Mine form NAKANO - - * proves its being a Palaeopharus. paucicostatus NAKAZAWA 2. buriji KIPARISOVA (1954) which was re- - * (Mi11epharus) triadicus ToK~- '* YAMA, n. sp. * Received March 22, 1958; read at the 67th Meeting of Palaeontogical Society of See the Table for Palaeophams in japan, April 26, 1958 at Niigata. Japan. On this occasion the develop- 291 292 Akira ToKUYAMA ment of the "pseudocardinal" teeth in The tirst group is the true schizodont the Triassic "schizodont" pelecypods is in which Oil(! triangular media•1 tooth also discussed. ~ exists ·on · lhe · left vaiYe. and the The writer is greatly indebted lo Pro dental arr:1ngement is as shown in fig. fessor T. KoBAYAsm for his kind. guid 1. In -the second group. the triangular ance and encouragement. median tooth is seen on each ntlve (3 and ~; awl tooth :~a becomes obsolete. According to CHAVAN !1954. in NEwELL, Hinge De,·elopment in the Triassic 1957) the hinge formula of "Permo "Schizodonts '' pi/Oms" is :2, (•Jb:. PIJ-(:)a;, 3b. PUl. but The hinge apparati of the Triassic NEwELL CO\lld not confirm the existence Schizodonts can be· classified into the of the tooth 3a in the Ame1:ican "Permo following groups: p/wrus ". As shown in the above for- .1. True $chiwdonts or :llyop/wria group mula. posterior lateral teeth are de\·elop including Trigo11odus. Pachycardia. lfemi ed in this group. Another Permian najas, Myop!wria and illilwlri),;ollia. genus Ce!toiclcs KEWELL (19:17) has 2. (Jb). 2. Prc-llcterodonts Pll in the left and 3b. (['II. PIII in the 2A. "Pleuroplwrus "* group including right yalve. Triassic .Cardinioides KonA. "1'/eump!wrus" and Cardinioides. YASHL and lclllKAWA ~}95:2; has the same 2n. Cardinia group including Pa/aeopha· kind of hinge, although jts cardinal ih a rus. Jsoprisles and Cardinia. pseudocardinal. In the third group the tooth ~a become.s obsolete and"' is sepa R 3o 3b Myophoria rated from the nymph in some forms; L 4~• 2' • laterals arc developed in the anterior -as P.!.J..!!I) 3 well as in the posterior -J~art. although "Pteurophorus l~> " anterior laterals are insignificant in 1• 41 Pll Alii p.!t!!.ll some pri rn iti ve forms. The so-called i Palaeopharus pseudocarclinals are often seen in the All )~~t Pll A!.J.!!_I) 3 P.!J.!!.I) three groups. In the first group the Cardinia tooth ~ (or ~b) is crenulate as in \ z;;;> -All 2• Pll Trigonodus and Pac/Jycardia .. The Unioni Text-figure 1. Hinge arrangement of pre- dae belong to this group in vvhich the hdcrodonts and schizodonts. position of pseudocardinals is variable. In the 2A group the left cardinal tooth takes the aspect of the pseudocardinal * As Cox pointed out (MARWICiz, 1953). the generic name. Pleurophorus K!NG (184-1). as in Cardinioides. In the 2B group it is preoccupied by 1'/europ/wrus. MuLsANT is not a cardinal but a lateral tooth. (18-12). a scarab _Coreoptera (S<;HUI{T7.E, KonAYASIII and lciiiKAWA (1951) empha KOCKENTHAL und IlEIDER, 1935). CHAVA."i sized the resembl:cutce·of the pseudocar 195,1) proposed Permophorus for its substitu· dinal tooth between Unio and Ptilaeopha te, but it is open to question whether Pe/'1110· rus. but the resemblance may be super p/torus is valid or not, because Pleurop!wrella ficial. because the pseudorardinal in GmTY (190-1) or Pleurop!wrina I.rcnAREW unionicls is really a kind of cardinal (1925) may be its synonym (NEWELL. 19-10. tooth. while in Pa/aeoplwrus it is genera 1957). For the time being the writer uses " Pleurophorus" for the pelecypod genus in ted from an anterior lateral tooth. The question. writer thinks that the presence or Late Triassic Pah'!'•.•Pharus in Japan 293 absence of a " pscudocardinal tooth " is opisthodetic. external and supported by not so essential for the pelecypod clas well defined nymphs in anterior half of sification as I hey arc evaluated. Palaeo· shell: area fairly large, elongated and plzams should be separated from Uuio stria ted transversely by growth lines. group which is perhaps a descendant of Anterior addctor scar stout. sl rongly the Myoplwria group. The three groups impressed; posterior one lar~er and less are all closely allied in the muscle scars distinct; pedal retractor scar very deep, and ligament characters. although their just behind anterior adductor scar. One hinge apparati are different genetically. cardinal tooth t:~b) on right valve and The oldest is the Myoplzoria group from two 1~b and -lbl on the left: one pos which the "Pleuroplwrus" group was terior lateral tooth on each valve strong presumably derived in the Carbonifer and massive. Anterior lateral tooth ous; subsequently the Cardinia group wide and its groove finely crenulate and from the second group in the late pseudocardinal-like. Ornanwnt variable: Triassic period and attained its acme in test smooth or concentrically striated the Jurassic. This is the link between and often radially costate. the " pre-heterodunts " and I he true Remarks:- Bt)HM (1914. S. !i!ili) consider· heterodonts which the latter t1ourished ed andersoni to be a Pleuroplwms but in the late l\1esozoic and Cenozoic. lt is andersoni as well as japanese species of interesting to see a transitional form Pa!aeop/zarus are separable from "Pie· between the "1'/europlwrus" and Cardinia urophorus" KrNG in the generic rank groups in the !\line collection'>; 1llincplw· as KoBAYAsHI and IcumAwApointed out. rus. new su bgcnus. may be the most They laid stress on the pscudocarclinal primitive and earliest member of the tooth in this genus. In the typical Cardinia group. maizuruensis from Nabac and lleki. it looks like a cardinal tooth and similar to a pseudocardinal in Unio. NAKAZAWA ne~cription of Species (1955) noticed extensive variation in its Su perfami I y Cardiniacea shape and called it a "pscudocardinal area " which does not mean a tooth. Family Cardiniidae ZrTTEL because the area in each valve does not interlock each other. In the Kyowa Genus Palacup/wrus Kr-rTL. emend. specimen. tNAKANo. 1957. pl. !1. fig. Ill. KonAYASllt and lcmKAWA the .. pseudocardinal " tooth is elongated l!!Oi: Pal apparati of Myophoria-type. Its pseudo this in cardinal teeth. nymph and pos cardinal is a kind of cardinal tooth and terior lateral tooth, but an invaluable the crcnulatc tooth on the right valve distinction is the absence of the "Pseudo interlocks with the crenulate socket on cardinal" in MARw 1cK'S. The hinge the left valve. Therefore. so far as the apparat i of the Upper Triassic "Pleuro "pseudocardinal " is concerned, Palaeo p/wrus" curionii BITTNER disagree with pharus is essentially different from Unio those of Palaeozoic "Pleurop/wrus" s. and the resemblances are superficial. str. (BiiHM, 1914). BtTTNr:R's may be According to NEwi·:LL (1957) Permian congeneric wit l1 MARW tcK's. Jurassic '' Pleuroplwrus" has the following 1\alentera !\IARwtcK (1953) is also a characters*: relative of Triaphoms. 1\.a/en/em ap Lunule and escutcheon well develnped. pears to be related to Myoconcha rather ligament opisthodetic and parivincu lar, than Palaeoplwrus. After all. Cardinia supported by well defined nymphs which AGAssiz may be the closest ally to are limited lo the anterior one-fourth Pa!aeoplzarus, although AGAssiz's is of the dorsal margin: dentiton consist distinguishable from this by its surface ing usually of one postero-lateral tooth ornament and anterior lateral tooth. (Pili) in the right valve along the shell The two genera are so closely related margin which passes over a single in interior and ligament characters that lateral (PII) of the left valve: one they belong to the same family. l'vlore cardinal tooth in each valve. that of the precisely. they have similar arrange right (3b} passing abO\'e that of the left ment and shapes of cardinal. posterior (2): rarely. there is an addilinnal poorly lateral Ieeth and equally large and developed. inner postero-lateral tooth elongated ligament area. Moreover. in (PI) and an outer cardinal (4b) in left Minepharus, a new subgenus of Palaeo valve of a few individuals. p!wms, the anterior lateral groove is not This agrees with Permian ·· Pleuro crenulate and the lateral is though phorus" in the characters of lunule, smalL similar to the one in Cardinia. escutcheon, ligament, nymph and hinge The strong muscular impressions and arrangement. but the teeth 2 and ·Ia entire pallial line are the features com and nymph are more developed in this mon between the two genera. Therefore. than in that genus. The tooth 2 is l'alaenpltarus 111 ust be referred to the very small and elongated in this, while Cardiniidae. it is a triangular boss of moderate size in "1'/europ/wrus ". Furthermore, um-. bones are pointed a little more posterior PalaeojJharus oblongat us (KonAY ASHI ly in it than in "Pl!'uroplwrus ". Tria and lcHIKAW A) plzorus l'vL~RWICK (Hl5:~) from the Upper Triassic of New Zealand agrees with Plate ·13, figures 8-11. * KING (1850) illustrated two prominent 1!l39: Pleurophorus cfr. Perlo11gatus BoHM II}' cardinal teeth in each valve and a marginal KATAYAMA. f. Geo/. Soc. Japan, t•o/. lateral tooth in the left valve. but BoHM -J{j, p. 381 (listed). (1914) and NEWELL (19·10) pointed out his 19511: Ple!lroplwrus oblongatus KoBAYASHI erroneous observation. Its dental formuln is and lcHJF;AWA, ]. Fac. Sci. Unit•. Tokyo, 2, (4b), PII- (3a), 3b, Pill according to set. 2, mi. 7, p. 212, pl. 1, fig. 10. CHAVAN (1954). 1954: Plettroplwrus cfr. oblongatu;; ICIIIKAWA• Late Triassic Palaeopharus in Japan 295 ]. lnst. Polytech. Osaka City Unit•.. t•ol. ribs. The latter may be no more than ], 110. 1. p. '10. varietal forms. The anterior portion is a little acuter in the !\line form than DescrijJtion :-Shell three to four times in the typical Kochigatani form. as long as high: umbo subterminal. anterodorsal margin almost straight: Comparison:- The !\line form is indis anterior part roundly trigonal : dorsal tinguishable from the typical form if the pseudocardinal is not preserved. and ventral margin~ almo~t straight and parallel in median part: posterior because the two forms are similar in margin more gradually rounded than the outline. muscular scars. pallial line, anterior. Lunule small and triangular: ligament features and in the hinge ligament opisthodetic, external: lig<.. apparati. especially in the anterior hinge ment area elongated. and striated trans area. They arc almost identical in the versely by growth lines. Anterior general outline and surface ornaments. adductor scar ovate, strongly impressed: The preservation of the typical form in posterior one subrounded. large and less coarse sandstone suggests that the distinct than anterior one: p~dal retrac "pseudocardinal" is eroded out in the tor scar small and pitted. just behind holotype. Therefore the writer thinks anterior adductor. Pallial I inc entire. that "Plem·op!wrus '' oblongatus should Left hinge consists of a cardinal (3b). be referred to Palaeop/rarus. Carnic "pseudocardinal" and posterior lateral Pa/aeopharus buriji KJPARJsovA from tooth: 3b \\·elded to an elongated nymph. :Maritime province of Far East may be Right hinge consists of a triangular wnspecific with this. According to her cardinal tooth (2) and a socket just pos (1954, p. H) the surface is feebly marked terior to 2. "Pseudocardinal" wide. and by radial striae. Faint radial striae and corresponding to one on the left valve, pseuclocardinals are sometimes recogniz but they are not interlocked with each able also in this species. That form other: they are so to speak "cardinal differs from this in the more anteriorly areas". Surface ornamented with con pointed umbo and more inflated posterior C tani and Siberian forms is \'cry widely ted from margin of lunule; 4b elongated. distributed in northeastern Asia in the slender and welded to a depressed tri Carnic age. angular nymph; a socket 3b' deep. fairly short. Socket PI' profound and remark Pa!aeop!wrus ob!ougatns buriji able. Pill less distinct than PI. A narrow K!l'ARISOYA furrow divides anterior lateral (Alii). Ornament consists of stout pseudo-radial Plate 43. figure 1::!. costae and concentric lines of growth. 1951: Palaeopflarus buriji KlPARISOVA, Field Remar!?s :-In this subgenus the an illustrated hand booh etc .. p. -!4, pl. 34, terior cardinal 2b. is not so remarkable figs. 5. 6. as ·· Pfeurop!toms" in which. according to Bt>HM and NEWELL 2b is stout and Characterized by very weak pseudo triangular: posterior cardinal 4b slender radial striae in postero-dorsal area. The hut distinct in this subgenus. while in striae are widel~· spaced. angulated on Permian .. Pleuroplzoms" it is not sepa the top and crossed by concentric rated from the nymph. WAAGEN (1907), lines of growth. The posterior outline is however. described 2b. 4b and 3a of slightly more inflated in Siberian (KIPARI· "P." wrionii from Seiser Alm. There sov A. tlg. 5) than the common 1\Jine iore. except for its rudimentary anterior form. Hinge norm:d for Palaeop/unus. lateral. it agrees with Triassic ·· P!euro· Occurrence :-Common in sandstone of plwrus ··. This rudimentary lateral is the upper I-lirabara formation and mid cleYeloped into a pseudocardinal of dle !\so formation (mid.- up. Carnic) Palacoj>lwrus. Therefore the presence and upper Carnic of Maritime Province of the anterior lateral groove is an of Siberia (T; of KIPARrsovA). important distinction of Palaeoplzants from "P/europhorus ": the groove is Subgenus J11im•pharus ToKt:YAr.-tA. crenulatc in Pa/aeoplzarus (s. str.). but non-crenulate in lllilleplzants. If it is new subgenus crcnulale. there is no distinction of this T_lj'Je species:-Palaeopltams (:11inrplw· subgenus from the typical Palaeopllarus. nts) triadicus ToKUYA:l.IA. ne,,· subgenus The writer opines that "pseudocardinal" and species. of Pa!acoplwrus is derived from the Diagnosis:-Shell elongated. elliptical. lateral tooth ontogenetically. This sub a little compressed. :1t1d slightly taper genus is closely related to Cardinia in ing anteriorly; umbo prosogyrous. at a the musculature and hinge, although fourth from anterior extremitv; liga the anterior lateral is a little weaker ment opisthodetic. subinternal: ligament in the former than the latter. They area triangular and depressed; I unule agree with each other in the arrangement small but distinct. Internally. anterior of cardinal teeth, elongate-trigonal nymph adductor scar very strong; posterior one and massive lateral teeth. general fea subcircular. larger and less distinct than ture of hinge area, prominentopisthodetic anterior; pedal retractor scar small. and external ligament. stout muscular prominent. Pallial line entire. Hinge impression and pallial line. Thus .. Ple consists of a right and two left cardi· urophoridae" are linked with the Car nals, and a lateral tooth in anterior and diniidac by this subgenus. posterior. Tooth 2b not clearly separa· Cardinia is said to be related to Late Triassic Palaeopharus in Japan 297 Trigonodus (WAAGEN, 19117). but its hinge 111easurcmeut iu 111111. Length Height is ncit her of Trigonodus- nor Jl!yoplioria tvpc. and agrees with that of l'alal'o 1-lolotypc (Left) 50. 5 21.4 p/zarus. especially this subgenus. There Paratype (Right) "15. 3 15. () fore Minep/zarus is a primitive form of \·:~rietal form (Left) <15. 5 21. :! the Carcliniiclae. \·ariatal form (Right) ~4.G 17.0 ."ipecies:-P. (Minepharus) triadicus. monotypic. Obsen~tllirm :-!\ small and rounded Distribution :-Carnic: Japan. proclissoconch is preserved in an im mature shell tlig. 5). In the subsequent stage the radial rib starts on the postcro Palaeophams i MincP/wrus: triadic us vent raJ side \Yhere it is originated from ToKUYAMA. new subgenus and the concentric rib or striation. This new species con(entric line curves down to the posterior corner and develops into a Plate 43. ligures 1-'i': rcxt-ligure ~- radial rib. The same process of costa lion was found by B,~mM (l!l03) in ander Description:-- Shell moderate in size. snni. The umbo lies at or ncar the but small for Palaeop!tams. elongated. middle in the juvenalium. but is slightly elliptical. a little compressed and slight :mteriorly in later stages of growth. A ly taperin.~ anteriorly: umbo prosogy varietal form (fig. c!) is very short. Its rous. at a fourth from ant:.>rior extremity: internal mould closely resembles that hinge lirie a little longer than a half of in Cardiuia. shell: ligament opisthodetic. external: Comparison:-This species is shorter ligament area wide. distinct. located than Palaeoplwrus and resembles Cardi rJear middle and transversely striated llia in outline. Except for radial costae this is close! y related to Cardinia. both internally and externally. The surface ornamentation of this species is similar to those of Palaeopharus andersoni, !'. verlongus. " Pleuroplwrus ., curionii and Palaeozoic "PleurojJ/mrus ·• radial us. From all or them. however. this is easily distinguishable by its hinge ap Text-figure ::!. Palaeoplmrus (.lfinepllarus) parati. It is a linking form between I riadicus. the so-called Triassic ·· Pleuroplwrus" and Palaeoplwrus and also between by growth lines. Lunule small· but Palaeop/wms and Cardinia; the earliest distinct. Hinge and other interior member of Cardiuia is coexh;tent with characters of Mineplwrus. Surface cover this. ed \vith several stout and rounded pset d) Ocwrrence:-Common in 1he transitic radial costae on dorsal side. They begin nal zone from the inland to open-sea in the earliest stage and becon1e stout facies of the middle llirabara stage (low. and widely spac~d. GrO\Yth striae fairly Carnici at Ilirabarazaka. west of Omine. strong, concentric and somewhat irregu l\Iine in Kagato. larly. 298 Akira ToKUYAMA the lleki Formation in the Province of References cited Tamba (Kyoto Prefecture), Japan. ]apa11. BtinM, j. (1903), Ubcr die obcrtriadische f. Geol. Geogr., l'Ol. 22. pp. 55-81, pis. 1-3. Fauna der Bareninsel. K. S1•e11slw VeteiiSk. MARWICK, J. (1953), Divisions and Faunas Akad. !/mull.. Bd. 37, Nr. 3, 76 S., 7 Taf. of the llokonui System. Geol. Sun•. N. Z., -- (191·1), Zur Gattung Pleuroplwrus KING Pal. Bull., no. 21. 141 pp., 12 pis. und !Uyoconclla SowERBY. ]b. k. k. Preuss. NAKANO, M. (1957), Carnian Fossils from geo/. L-:1., Bd. 35. S. 5·19-561, Taf. 21. Kyowa, Okayama Prefecture, japan. ]. IcnmAwA, I<. (1954), Late Triassic Pelecy· Sci. lliroshima Unit•., scr. C, m/. 2, no. 1, pods from the Kochigatani (;roup in the pp. 63-67, pl. 9. Sakuradani and Kito Area, Tukushima NAKAZAWA, K. (1955), A Study on the Pele Prefecture. Shikoku. Japan. Part. 1. f. cypod-Fauna of the Upper Triassic Ins/. Poly/ec/1. Osaka City Univ .. ser. C, L'Ol. Nahal' Group in the Northern Part of 1. no. 1, pp. 35-37, pis. 1, 2. Kyoto Prefecture, japan. Pt. 3. f. Col. KATAYAMA, M. (1939), Stratigraphical Study Sci. U11il•. Kyoto., ser. B. MI. 22, 110. 2, on the l\Iine Series in Prov. Nagata. f. pp. :!·13-260, pis. 13-Hi. Geol. Soc. japan. mi. -16. pp. 217-234, pl. 8. NEWI,LI,, N.D. (19·10), Invertebrate Fauna of KING. vV. (ISH), On a New Genus of Palaeo· the late Permian Whitehorse Sandstone. zoic Shells. Ann. Mag. Nat. !list., l'Of. 14. Bull. Geol. Soc. rlm.. t•ol. 51, pp. 261-325, pp. 313-317. pis. 1-10. -- (1850), A Monograph of the Permian -- (1957), Notes on Certain Primitive He· Fossils of England. vol. I. pp. 1-253, pis. terodont Pelecypods . •-!m. ,\Ius., Nol'itates, 1-23. Palaeon/ogr. Soc. London. 1/0. /857. pp. 1-14. KoBAYAsHI, T. and K. IcHIKAWA (1950). Late NicoL. D. and W. T. ALLEN (1953), A new Triassic J\lytilus, Volsella. /'lt:urophorus pelecypod genus from Upper Triassic and !1/yoconcluz from the Sakawa Basin strata in Peru. f. Wasltin~:ton Acad. Sci., in Shikoku. Japan. f. Fac. Sd. Univ. r•o/. 13. no. 11, pp. 34·1-46. Tokyo, sec. 2, 1'01. 7. pt. 3, pp. 208-215, ScHUR Explanation of Plate :13 Figs. 1- 7: Pa/aeopharus (Mineplwrus) triadiws ToKUYAMA, new subgenus and new species ...... 297 1: llolotype, left valve: 2-7: paratypes: 2.4: short varieties: 5: immature left valve showing prodissoconch. x 3: 6, 7: internal moulds of both valves showing hinge and interior. Loc. Hirabarazaka, west of Omine, Mine city, Prov. Nagato. Figs. 8-11: Pa/acop!wrus oblmrga/us (KoHAYASIII and ICIIIKAWA) ...... 294 8: Internal mould of immature left valve, showing hinge apparatus. Loc. West of llirabara, Mine city. !l: Elongated form. Loc. Hirabarazaka: 10: surface of left valve, loc. ditto. 11: ;t: right internal mould showing hinge apparatus and interior: h: clay cast of the same specimen. Loc. ditto. Fig. 12: Palaeopllarus ob/onga/us buriji KIPARISOVA ...... 296 External clay cast showing weak ribs in the posterior area. Loc. North of Omine in Mine. All figures except fig. 5 are in natural size. All specimens illustrated here are deposited in the Geological Institute, University of Tokyo. ToKuYAMA: Late Triassic J>alaeopharus Plate 43 1 5 3 6 7 8 9 12 10 Trans. J>roc. Palaeont. Soc. Japan, N. S., No. 32, pp. 299-30-1, pl. 4-1, Nov. 1o. I !l:i7 351. AN INTERESTING CRUSTACEAN REMAIN. CTENOC!JELES SUJAKUI N. SP. FRO.M THE PALEOGENE OF KYUSHU, JAPAN* RIKIZO HviAIZUMI Institute of Earth Science, College of Arts and Sciences, Tohoku UniYersity :hmr~im.=:*C'/EJJIJ: vi" e:.H Ctenocheles sujakui n. sp.: {f{:~~rNJ~/J1jffi~!AtH.!~~ljr,fiJ=:~ ~ftil.tal:~ifr 11 lj-Jl', f,/!1.\t 249.68 m :toJ: U 309m O)}llili!fffiliff"f~~-/)·1; **11111i'li:.J: ~ t~H~"' H t:. Ctenocheles sujakui n. sp. ~ifG""L.,t.:, 'CO)*iVCrt:JrPf.::.r;r:.~mtf.fifliir:f:lt1i:E,'{I<:J: 1Jft'lf.11~1 O)j.;:llfftJ•I;tlU.f!cHt, W.l:jfni<:J: ~lfHJ-~ttThiJ, relict -c'JJ~, 1 ;..t.:rn!li .:e- O)*!!Rir.t!;?f<:6:J:> IJ • {t1iO)W:!U1-c' t t>tJ. ~ ~"9 (, :ltl!f!"f::: ~ The crustacean remains here descri· the Crustacea at his disposal. l-Ie is bed, were collected from the Kishima also grateful to Emeritus Professor formation in the boring cores of the Hisakatsu YABE, Professors Sh6shiro Well. No. 11 of the Kogayama Coal I-IANZAWA and Kiyoshi 1\sANo of the Mines, Nagao, Minami·Taku-mura, Ogi· Institute of Geology and Paleontology of gun, Saga Prefecture by Mr. Tomosuke Tohoku University, Professor Kotora SuJAKu of the Mitsubishi i'·.'Iining Com· liATAI of the Department of c;eology. pany and were submitted to the writer College of Education, Tohoku University with other remains of a new genus of and Professor Kenzo YAGt of the Insti· the Goncplacids. They proved to be a tute of Earth Sciences. College of 1\rts very rare genus of the Family Callianas· and Sciences, Tohoku University for sidae, of which descendant is a relict in their continuous encouragt'mcnt s. the Japan Sea, Ctenoclzeles balssi Krsm· NOUYE, 1926. Description Although the C/enocheles remains are fragmentary, they have some importance Tribe Thalassinidea from the stratigraphic view point by Family Callianassidae their well-defined characters. Genus Ctenocheles KtsHtNouvE, 1926 The writer is indebted to l'vlessrs To· mosuke SuJAKu and Y 6ichiro MoRINAGA, KISHINOUYE, K., 1926, Two Rare and Remark Yasuhiro KAMADA and Toshio KrHARA for able Forms of Macrurous Crustacea from kindly placing the fo:osil specimens of Japan, A1mot. Zoo/. japon., Vol. 11, pp. 63-66. Fig. 1. * Received April (j, 1!!58: read at the 55th GLAESSNER, M. F., l!H7, Dcl'apod Crustacea Meeting of the Palaeontological Society of (Callianassidae) from the Eun:nc of Vic· Japan, Oct. 10, 1955, at Scndai. toria. Proc. Roy. Soc. Victoria, Vol. LIX. 299 300 Hikizo I MAIZUMI (Yew Series). Par. 1. pp. ·l-7. Pl. l. Figs. lures of Ctenocheles. K. BEURLE!'; drew 8 and 9, Text-fig. 1. attention to their striking resemblance Genotype: Clenocheles halssi KISHt:-;ouYE. to the cheliped described hy BALSS as Pentachles sp. ? In his description of Ctenoclteles z•ictor from the Eocene Rivernook House, on The fragmentary han1s and fingers of the coast about one and a quarter miles Jsdmodactylus also correspond with southeast of the mouth of the Gellibrand Ctcnocheles in their shape and weak River. Victoria. i\1. F. GLAESSNER has ornamental ion. though the dent iculat ion already stated that Cteuoclte!es was of most of these fingers is incompletely established for a living Thalassinid preserved. It should be noted that their corresponding in essential features with identification with Ctenoche!es does not Callianassa. but distinguished by the affect other species of lsclmodactylus, excessive deYelopment oi the right chcl Isdmodactys macrodaclylus (ScHLl'TER) liped. which resembles that of the and lsclmodacty!us esocinus (FRITSCH}, in lobster-like deep-sea Decapod Tlummas which long spiny claws are associated with lobstcrlike remains of the carapace torfw!es. Specimens of Tliaumastoc/ie!es or abdomen. Long fingers with long japonicus Cu.MAN arc collectccl from pointed teeth arc obsened in more than depth of about -100m. of the Sagami one family of Decapod Crustacea.· but Bay. Shizuoka Prefecture and I\umano nada. by dragnets. the shapes of ornamentation oi the hand Only a right cheliped of this species and fingers make it possible to distin guish them. was collected by DoFLEtN from Japanese waters and was described by BALss as Ctenoclieles has an asymmetrical anrl ? Pentacheles nov. sp. Therefore. the unequal first pair of the pereiopocl and specific name of the genotype was de very specialized large cheliped. The nominated as ba!ssi by KisHINOUYE. only specimen of the genotype. Ctcnoche The genotype. Ctenoche!cs balssi K1sm· !cs balssi has first pereiopocL of which Noun: was collected at Osu. ncar Kashi the right one is larger than the left wasaki, Niigata Prefecture. This speci and is specialized in the shape of the men. measuring 10 em in length. is finger like a comb. fcmaL::, and is more or less mutilated. Syntype No. 1. the manus with the This deep water dweller is blind. Any lixed finger of the present species of the other specimen of Recent species has Ctenocheles is determined as the right never been known to be collected there one. by the swelling of both surfaces of after. the manus and the vertical furrow 1\I. F. GLe\EssNER included Jschnodacty behind the articulation with the dactylus lus (lsclmodactylus coolwi RATIIntrN. lll35: of one surface. The surface with a Jsc/modactylus Cltlfellus I~ATHBUN. 193:-i: vertical furrow behind the articulation [srfmodrtctylus? dentatus RATII11UN, l!l35) with the dactylus. is inner one and from the Paleocene and Eocene of therefore the manus is determined as Alab:una anci l\lississippi and Thaumasto the right one by the orientation of the dw!es rupeliensis lh:uRLEN. 1939 from the inner surface of the manus. This right lVIiddlc Oligocene of Hungary. in the manus of the present species of the genus Ctenod1eles. The incomplete chcli Ctenocheles (syntype Ko. 1 ), belongs to peds of Tlwumastodwles rupeliensis the smaller cheliped of the first pair of BEURJ.EN. 1939 possess distinctive fea- the perciopoda. The other four syntypes An Inter..;st ing Crustacean f of the present ~pecies of the Ctenoclzelcs, rous irn:gularly polygonal pavements (syntypes. ?\o. II- V). are isolated. long. apparently belonging to the lower layer thin and denticulated fingers. They are cover the whole surface except the no doubt bel on.~ to the larger one of till' distal port ion. Manus of the left chela. fin;t pair of the pereiopocla. The left decorticatecl, outer surface covered with manus (syntype No. VIII) of the present polygonal pavements: palm much swol species is larger type. len: proximal end lacking: articulating It is not always the case that in tlw cond~·Ie with the movable finger is large: same taxonomic unit of Crustacean De· the immovable finger is strongly dc!lcxed. capuda, the one cheliped. whether it is fixed finger imperfect! y preserved. right or left. is longer than the other. perhaps shoncr than the palm. and oval Though it has never been a~sured in the in cross section at the base. with blunt living or fossil species as the Ctenoche!cs. and imperiectly prescrvC'cl teeth along have commonly asymmetrical chelipeds the prehensile edge. fingers of the especially in the male. The present large chela very thin. platy and long species of the C/enoche/cs has the left attenuate abruptly in the distal part. larger cheliped. while the known speci with regularly altcrnating longer and mens of Ctenm:lteks vir:tor and Ctenod~t· shorter pointer! teeth: longer teeth of les ba!ssi. have the right larger chelipeds. t'1e fln!-(cr are regularly spaced and gradually shorten their length to the anterior part of the finger. the posterior Ctenochl'll!s sujalmi n. sp. part a~1d the base of the finger unknom1: bet\n•en the longer teeth about three Plall' ·1-1. figures 2-5. or four shorter teeth ~rranged, in these Based on the right manus with the shorter teeth there are differences in fixed finger (syntype Xo. I), four frag length. and the one near the longer mental fingers (Syntypes No. II-V.i of teeth shorter than the others: the curva the larger chela (perhaps. left one). !he ture' of the teeth slightly directed back right immovable finger of the second wards in most cases: the distal ends of pereiopod (syntype No. VJ,, the fragmen the teeth curved. The base of the right tal posterior part of the chela (syntype immovable fingers, (syntype No. VI\ No. V!I). and the anterior part of tlw showing the inner superior surface with left manus with the stump of the fixed the ill-preserved small teeth along the finger (syntypc No. VIII). prehensile edge. perhaps smaller chela Palm of the right manus elongate of the second pereiopod. The fragment about one and a fifth times as long as of the posterior part of the manus hi.~h: swollen. thickness half of the (synt ype No. VII), perhaps belongs to width: superior border blunt, almost the left larger chela of the first pcreio straight but gently swelled in the middle pocl. co\·ered with irregular pavements. part: inferior border also blunt, but thin Dimcnsio11s :-:1\'lanus 1syntype No. I): ncr than the superior: raised and swollen length of palm: 16.:! mm: width of palm in the middle and the posterior part. at distal part: llnm1. maximum width with 7-8 pits along the inferior margin. in the middle part of palm: lcl mm. A vertical furrow behind the articula width of proxim::d part of palm: 12.5 tion with the dactylus on the inner mm: thickness of palm: 6.8 n1111; length s.1r.' 1: e is broad and shallow. Nunw- of fixed finger: 10 mm: basal section of 302 Rikizo IMATZUMI fixed finger: 2.5mmX1.8mm. c/wles victor is more than 14 111111 and Fingers (syntypc No. II): length. 29 the length of its long teeth are over 2 mm+. width, 2.5 mm±: (syntype No. 111m and the length of its short teeth Ill); length, 26 m m +. width. 2.fi mm ± : are less than l/2mm. while the length syntype No. IV. length. 20mm+. width. of the anterior part of the finger of :~ mm±; syntype No. V. length. ll mm-1-. Ctenocheles sujalwi is more than 29 mm Distances between the longer teeth. 5 and the length of its long teeth arc over mm and 5.8 mm in syntype No. ll at 2mm and the length of its short teeth the anterior part; 4.2 mm and 5 mm in are less than 1 mm. Morphologically syntype No. III at the anterior part, the they resemble very much, but the speci distances between the longer teeth be men of the finger tlgured by M. F. come longer forward. (syntype No. VII): Gt.AESSNER in his figure 9. Plate I. (M. fragment of thC' posterior part of the \V. G. K. No. 1925) shows the long teeth manus of the left larger chela. width, bending slightly forward and the long 8.5 mm+. length. 14mm+. (syntype No. teeth of the fingers of the large cheli VIII); the left manus, the width of the pcds of Cteuoche/es sujalwi leaning anterior part. 12-13 mm: sect ion of the slightly backward. except the anterior stump of the immovable finger. height. long teeth of the syntype No. II. 2.7 mm. width, 0.7 mm. C/Pilocheles sujakui resembles closely Geological J/orizon. Locality and Re· to the living Ctenocheles ba!ssi. but dif positm:v:-Kishima formation, Ashiya fen; from the latter by the robust shape group: from the boring core of the of its finger, and hy the shorter palm well, No, 11 of the Kogayama Coal of the smaller cheliped of the first mines, 1\litsubishi !\lining Company, at pcreiopod. the western extremity of Nagao. ivlinami lsclmodactylus culte!lus RATHBUN, 1935 Taku·mura. Ogi-gun. Saga Prefecture*; is described by the two specimens as IGPS Joe. no. Sa-002; lat. 33°17'18'' N., follows, the one is the dactylus of the long. 130°7' E., T. SuJAKU coli.. 1953, left cheliped, hololype (this specimen is syntype No. I. sample number of the mistaken for the right one by 1\l. j. boring core No. 59, 2·19.6 m in depth. RATHBUN, in the description, p. 14, but dark siltstone, IGPS coli. cat. no. 79567: stated as the left one in his explanation syntypes No. II-VII. sample number of of the figure. p. 139), preserved in the the boring core no. 68. 309m in depth. Alabama Museum of Natural History, dark siltstone. IGPS col!. cat. no. 7!:1568. from the Eocene Sucarnoochee formation Syntypc No. VIII. without sample num· at Estelle, Wilcox County, Alabama and ber of the boring core. but seems to be the other is the right propodal finger. near to No. 68, syntypes No. II-VIII. 309 paratype, preserved in the John I Iopkins 111 in depth. dark siltstone, IGPS coil. University from the lower Eocene at cat. no. /9:168. with a molluscan frag· Dry Creek, Jackson, Hinds County. mcnt (Crassatcl/iles? sp.). Mississippi. These two specimens are Remarks :-Cienoche!es victor is larger also smaller type. Among these two in size comparing with Ctenodwles su specimens. paratype. the right propodal jakui, the length of the finger of Cteno- finger is 8.6mm in length (end lacking). 1.·1 mm in height at articulation, 0.5 mm * Jtd"llliVJ ':VJ£ 1n11tH~ ~ wftt~Srri.v:IJ. ~.~rill~ ll1 in width of the teeth. The propodal ~mwr. f!i.l111i!lllftr-l:.~li!i_, finger of the left smaller chela, (syntype, An lnkrcsting Crustacean J No. h of Ctenocheles sujalmi is more distinctive features of Ctenoche!es. but robust than the propodal finger of the Ctenocheles sujakui is easily distinguish· right smaller chela of Jsclmodactylus eel from the preceding species by the culte!lus. lschnodactylus cooltei RATHBuN. large number of the short teeth of the 1935 is collected from the Eocene finger. The living Australian Ctenoclzeles Sucarnoochee formation. Midway in collini i\·1. \YARn seems to be Yery robust. Prairie Creek region. Wilcox County, Alabama (holotype and paratype) and from the Sucarnoochee formation. Estel· References cited le. Wilcox County, Alabama (other spe· cimens). the type locality of lsc/modacly· BEURLEN. V. K. (HJ:N). ~eue lkcapoden-Krc· Ius cultellus. The specimens of !sc/mo· h!ic a us dem ungariscllt'n Tert iii r, 5 1\bbil.. Taf. ;-. Pa/aeo11/. Zeilsdt .. Fo/. 21, dactylus cookei are the right manus only. pp. 135-160. which belongs probably to larger cheli· GLAESSNJo:R, M. F. (1929), Fossi lium Cat:1logus. peel. It is more smooth in its su rfacc I : Animalicr. Pars 4 1. Crustacra Decapoda. and smaller than Ctenoche/es sujalwi. -- (19·11). Decapod Crustacea (Callianas The larger cheliped of Ctenocheles suja· sidae) from the Eocene of Victoria. Proc. kui is easily distinguished from Ctcno· Roy. Soc. Victoria. l'o/. LIX. New f"a .. cheles w/tel!us by its larger articulating i'ar. 1. pp. '1-7. Pl. 1. Figs. il and 9. Text condyle with the movable finger. Jig. I. lsclmodactylus (? \ de,rtalus RATHBUN. KISHINOUYr:. K. (1926). Two Rare and Rc· 1935 are composed only of the right nurkable Forms of J\lacrurous Crustacea propodal fingers and collected from the from Japan . .'\mwt. Zoo/. japon., Vol. 11. pp. 63-6ti. fil~· 1. same locality as the other specimens of ~AGAO. T. (HI:!S). A Summary of the !'a leo· RATtHiul'l, Dry lsclmodactylus cultellus gene Stratigraphy of Kyushu. Japan. Creek. Hinds Country. These three with some Accounts of tht: Fossilifl'rous species of Isclmodadylus. Isdmodacly/us Zones. Sci. Rep. Tolwlw Imp. Unit• .. Seco. cultel!us. I sclmodartylus cool~ei. I sclmo· Ser. (Geology). Vol. X II, No. 1. pp. 1-10. dactylus ( ? ) dentatus may actually 1'1. I. Table I. represent the different part of the same -- (19·11). On Some Fossil Crustacea from species. respectively, the dactylus and japan.]our. Ntc. Sci .. 1/oldwido lmj>. Unil• .. the propdal fingers of its left smaller ~er. IV. Vol. VI. No.2. pp. 85-100, 1'1. :?6. cheliped. the manus of its right larger RATIInUN, M. J. (1935). Fossil Crustacea of the cheliped and the right proporlal finger Atlantic and Gulf Coa:;tal Plain, Geol. of its larger cheliped. Soc. America. Sp. P .. 1\'o. 2. pp. G3-66, 1'1. 14. Figs. 1-12. 19-22 and 2-1. The incomplete chelipeds from the YAMASAKI, T. and MORINAGA, Y. (19 50), l\liddle Oligocene ( Kisceller Tegel) of Stratigraphical Relation bctewecn the Obunda near Budapest. Hungary which Sasebu and Karatsu Coal-fidds. Kyushu. K. BEuRLEl'l 09:l~l) clescribcd as Tlw· jour. Geol .."'oc. ]apall. Vol. LX. 1Vo. 710. wnastocludes mpe!iensis possess the pp. 473-486. 5 Text-figs. 304 J~ikizo IMAIZUMI Explanation of Plate 4-l Fig. I. Ctenoc/ieles balssi 1\tsHtNOUYE, collcctccl :11 cJsu, ncar, Kashiwasaki. Niigata Prefecture, rc1>roduced from Fig. 1 of KtsmNoUYE's paper, nat. size. Figs. 2-1. Cte11ocheles sujakrmi n. sp., Kishim:~ formation and Ashiya group, from the boring core of the \V ell, No. ll of the Kogayama Coal mines, Mitsubishi !\lining Company, at the western extremity of Nagao, ivlinami-Taku-mura, Ogi·J.(Un, Saga Prefecture, T. SuJAKtr, coli., 1953, syntypc No. I. sample number of the boring core No. 59, 249. 6m in depth; syntypes .:-\o. II-VII. sample number of the boring core No. 68, 309m in depth; syntypc No. VIII. without sample number of the boring core, but seems to be ncar to No. 68. with a molluscan fragment (Crassatellilcs? sp.). Fig. 2. Syntypc No. I, right manus with the fixed finger, external vil'W, '<. 8/3. Fig. 2a. Manus of the same specimen as fig. ~. external view, x 7/3. Fig. 2b. Manus of the same specimen as ftg. 2, internal view, x 7/3. Fig. 2c. Manus of the same specimen as fig. 2, internal view, x 5/2. Fig. 2d. Manus of the same spedmcn as fig. 2, internal view, x 1~/G. Fig. 2c. Fixed finger of the same specimen as fig. 2, uppcr view, X 12/5. Fig. 2f. Fixed finger of the same specimen as fig. 2, lower ,·iew, x 2. Fig. 2g. Fixed finger of the same specimen as fig. 2, antt'l'ior Yiew, x 3. Fig. 3. Synlypcs No. !1-V. four llngcrs of the: larger chela (perhaps, left onl') and the right immoYable fingl!l' of the second pereiopod. syntype No. VI, x 2. Fig. ·1. Syntype No. ~VIII, anterior part of the left manus with the stump of the fixed fin. ger, >< 23/12. Fig. ·Ia. Same specimen as fig. '1. X 13/6. Plate 44 I MAIZUMl: Ctenocheles su}·akuz . n. sp. 305 PROCEEDING OF THE PALAEONTOLOGICAL SOCIETY OF JAPAN r13 4:?11:111''(0 m'iO [OJ ~~J 1958$ 6 ,E! 6 II ...... Rokuro MoRIKAWA Jtifijif!);:~ Jli!•'f.ffl; Jibfi•'jl; li:~'T=f.i~IC.t5 1• •""'C!#Hill -1. Foraminifera from the Akahira Group U:o (~0 r.· 13 il) o ~~I::H ~t ~ ~iiii1H!i0' in the Northeastern Corner of the Chi- K.lMif~MEIIltA:O.>ii!i ~ -c ,n ~ o chibu Basin. Central Japan ...... 1. Pamfusulhw and Pseudodolio/ina found ...... Hiroshi UJIIE in the Base of the Kamiyoshida Group. 5. bllliF.!t:IKTi~i'!liJ>till'!O)!~!lll'Jilr. "?l ''L . the Uppermost Strata of the Chichibu ...... ftiti.Hn.::.-. System, Kanto l\·lassif ...... 6. Plica/ounio of the Wakino Formation ...... Rokuro MoRIKAWA ...... Yoshihisa OHTA 2. On somt.: Fusulinids from the Higashi 'i. 1'\liocene Pero/rochus and its Associated zawa Valil'y. l\1amiyoshida-mura, Chi Fauna from Tochigi Prefecture. Japan chibu-gun, Northern Part of Kanto ...... ~~roK~~ l\!assif ...... J~okuro l\IQRIKAWA 8. On Some l\lorinc 1'\lioccnc Mollusca 3. Fusulinids from Shiroishi-yama, North- from Mic Prefecture, japan ...... ern part of Kanto Massif ...... Yoshio ARAI r114~~i~l'.¥-1··r:i-~m n rnJl~lJi' 15. ftlrE!t-f.i!iJIJ.\'li*lfilf Pectinidae ( ;,:- OJ 15), 19. Elcctron·l\Iicroscopic Fine Structure of Pecten cosibeusis YoKOYAMA &V: ;_:-v)W.:J Fossil Diatoms. VI. Observation on iillfL '"l (f~ih'D ...... lt'JW :1f:·-f!~ Some Diatoms Found in Dicalitc ·• DL- 16. On some l\Iiocene P• .IIloPe in japan (('\. 72~3 .... ' ...... Haruo OK UNO M;) ...... Kiyotaka CH!~ZE! 20. Palynological Notes on the Cenozoic 17. **'41'il- t!JQl~fl fflj!!l;~(l) r11 ffr IJJ: i :..-- ::1 ;t 7f =- Rocks from Japan I ...... Carci1lop/ax (('\.;\%) ...... -0-J.jUJit ...... 1\-lisaburo SH!MAKURA 18. Studies on the Ostracoda from japan 21. m=~·lfa~ rtE ~ :c O).i!illl!ll!tm! t mt'l!'.< IV. Family Cytheridcidae SARS 1925 ((\: t (Note: Every reference is to the number of artil:il') PALAEOZOOLOGY Protozoa KANllMA. M. and SAKAGA~Il. S.: J/esoschub(•rlella. a new Permian Fusulinid Genus from Japan ...... 31~ YosHIDA. S.: Palaeogene Foraminifera from Kombumori. Eastern Hokkaido. J;:tpan ...... 315 Cue len terata MtNATO, M. and KATO. l\1.: N.:!w Spe«.:it:s of Sipho11ode11dro11 frl)m Japan ...... 313 --- and Two Carboniferous Corals from the Kitakami Mountains, North· east Honshu ...... 3~8 ---and A short Note on /.mJ.wialeoidr:s loriyamai l'vhNATO ...... 333 ---and NAKAZAWA, K.: Two Carboniferous Corals from Okayama Prefecture... 308 YosHIDA.. T. and KATO. i\1.: .. Onimaru type" Coralg newly found in the Northern Kita· kami Mountain Region. japan 3?~_;:) 1\Julluscoidea KATAOKA, j.: llryozoa from the Daishaka Formation (Pliocene). 1\·linami·Tsugaru.gun. Aomori Prefecture 329 KoTOH. ]. : Brachiopoda from the Daishaka and Tsurugasaka Fossil Zones. l\Iinami Tsu· garu·gun. Aomori Prefecture ...... 306 ToKUYAMA, A.: On some Upper Triassic Spiriferinoids from the Sal\:twa Basin in Prov. Tosa. Japan ...... 3~~ ---: On some Jurassic Hhynchonellids from Shikoku. Japan ...... 327 l\Jollusca 1\J 307 ...1 308 Systematic Index lTOIGAWA, ]. : On the Miocene Lucinidac irom the l\Iizunami Group, japan ...... 305 ----: On some Cephalaspid Opisthobranchia from the japanese lVIioccne ...... 334 KANNo. S.: Fossil and l~cccnl. Species of the Genes Panomya from japan ...... 307 KoBAYASHI. T.: Nipponiirigonia and Rutitrigonia in japan ...... On an /lturia from the Poronai Shale in 1-Iokkaido ----: An Aturia from (?) Kusu Count~·. Oita Prefecture. Kyushu...... ----: A Gotlandian Nautiloid from the 1-lida Plateau in japan MASUDA. K.: On the l\Iioccne Pectinidae from the Environs of Sendai: Part 10, On Pee· ten (:-lequipeden) yauagau·acnsis No~rURA and ZINBO...... 336 Ibid.: Part 11. On Pecten (CI!lamys) crassi11enius YoKOYAMA and its related Species ...... 341 ll>itl. ; Part 12, On Pee/ell ka;;amianus YoKOYAMA ...... 3-18 l\IATsur. 1\I.: Species of the Genus Nej>funea from the Palaeogene Formations in the Ku- shiro Coal Field, llol:kaido, japan ...... 338 illATSUMoTo. T.: A Turonian Danwsifes from Ilokkaido, Japan. (Studies of the Cretaceous Ammonites from I-lokl;aido and Saghalien) ...... 319 NAKAZAWA, K.: "Monophyllites ·· aralwrensis sp. nov. from the i\laizuru Zone ...... 340 SHIBATA. 1\-1.: Some Molluscan Fossils from the Eastern Part of the Tanzawa l\lountain- land...... 309 SHUTO, T.: Gra11uli.titsus from the l\liyazaki Group (Palaeontological Study of the l\liya. zal Arthropoda IMAIZUM!. ]{.: A Miocene Fossil Crab, Faratymolusyabei. n. sp. from Nagano Prefecture .. 310 ---: Three new l\Iiucene Species of Callianassa from Nagano Prefecture ...... 318 An interesting Crustacean 1\emain, Ctenocl;e/es sujakui n. sp ...... 351 KoBAYASHI. T.: Some Cambro-Ordovician Fossil~ from the Tan'gyang or Tan yo District, South Korea ...... 339 PALAEOBOTANY ENuo, R.: Stratigraphical and Palaeontological Studies of the Later Palaeozoic Calcare- ous Algae in Japan. X Ill. A Restudy ot the Genus Physoporella...... 3-17 IcmKAWA. \V.: Some Notes on a rare Species Triceratium simp/ex j. BRUN ...... 323 ---: Once again on a rare Species Triceratium simp/ex]. BRuN ...... Short notes 2 Kir.IURA. T.: i\ksozoic Plants from the Tctori Series, Central Honshu. Japan ...... 331 OKuNu. 1-1.: Electron-Microscopic fine Structure of Fossil Diatoms, V; Observation on some Diatoms found in Celatoms ...... 3-13 SuzuKI. K.: On the Occurrence of Cercis in japan ...... 332 TAKAHASHI, K.: Sabalites aus den \·\Takata Schichten von Tsushima, Nordkyushu ...... 335 l\Iiscellanea I-lATA!. K.: Interesting Sand-pipes from the Miocene Toyoda Formation, Yamagata Pre- fecture, Japan ...... 321 INDEX OF FOSSILS Notes: Every reference is to the number of article: words in it:1lics arc names of genera and species: words in hea\·y type, names of new families, genera and species. Cncis Endoi ...... 332 A -- sp...... 332 Cltlwnys in).;eniosa tanakai ...... 33·1 ··Adeon" sp...... 334 kolakiensis ...... 326 Alderiua lum::awai...... 329 kurumensi.Y ...... 326 Amussiopec/en iifomiensis ...... 311 cf. kurmJu'usis ...... 326 -- p!anicoslulatum ...... 311 obe.~a ...... 3-14 -- praesignis...... 311 sp...... 326 -- yabei ...... 311 Cladophlebis sllins!mensis . . :n1 Am.lf[da/ophyllum giganteum ...... 308 c:ampltysoporel/a conforma...... 347 A11tropora daishakaenRis .329 -- facet a ...... 3H -- elongata ...... 329 llurltwsn ...... 34i -- lwlaii .3~9 C/isaxopltyllum mm atetsuene ...... 30/l Arcomytilus sp...... 33(1 C/isiopltyllum :1ff. nl'coyanum ...... 328 Arcomytilus dairensis . 33(1 Coleopora tsu{lnruensis...... 329 As! rea (Pachypoma) omorii ...... 309 - tsugarueusis mn.•udai ...... 329 Aturia minoensis var...... 32·1 Co plot ltyris adamsi ...... 306 yokoyamai...... 317 Crassima rgi nate//a parvinl'ieular ia ...... 329 Cryptopecten yana;:awaensis ..... B ...... 336 C; euocheles ...... :~!)1 Berkeia shantungensis ...... 339 -- sujakui. .351 Boliz·ina k/einpelli ...... 315 Ctspidaria (?) a sp. .337 Burmirhynchia ...... 327 (?) b sp ...... 337 ·• --" eapillala ...... 327 Cydammina paci,lica ...... 315 -- japoni('(l .. ' ... ' ...... :~27 pacifica var. kushimensis ... .315 -- (oriiiOSI/l'IZSis ...... :l27 (~l'liclma sp...... 334 Cyliclmatys sp ...... 334 c Cymbella cistula ...... 3·13 Callianassa Shikamai ...... 318 -- Tanakai ...... 318 ...... 319 -- l'agii. . .. 311! Damesites ainuamts Decorifer ena ...... 334 .. Camptonectes •· oishii ...... 326 " subflabelfiform is ...... 326 Dipltyphyllum delicatum ... 3~8 " asp...... 326 " b sp. . . . 3~6 Cardiniidae ...... 316.350 £,/folium sp ...... 326 Cardinioides ...... 316 E•1tosolenia aff. t•asiformis ...... 315 -- ot•atus ...... 316 cocy/china aJfabilis...... 334 -- variclus ...... 31G - habei...... 334 -- sp. ··············· ..... 316 -- tokiensill ...... 344 Cassidulina margareta ...... 315 Eopecten (?) sp...... 326 Cavi/ucina (Monitilora) kitamurai . . .. 305 Eoscap!umdcr corpu/enta ...... 334 309 310 Index of Fossils Exodrdla sp...... 3~9 ;Uineplwrus ...... 350 Miyagipecten sarc11nen.~is ...... :l49 .. Jfmrop!zylfites " amlmrensis ...... 310 l\lytil idae ...... 330 Gforipalfiroll aassit•e11inm ...... 341 Jlyt ifus (Fafcimyl ilus) lzl•rcmirus ...... 330 -- iZIII't'IIHiH ...... 341 (-) - subsp...... 330 Grammalodmr (?) sp...... 337 (--) stricapi/lalus ...... 330 Gramtli[usus ...... 3-16 (-)- subsp...... 330 -- dna/is...... 3-16 (-) sp...... 330 -- koiJIUITI/IH ...... 3.(6 -- mntsrwwtoi...... 3·16 Gullulina cf. adhaerens...... 315 -- cf. problema ...... 3!5 Nep/zmea dispar ...... 338 -- d. r·arians ...... 315 -- ezoana ...... 338 -- lwruhalai ...... 338 J[ -- modesloidea : ...... 338 -- onohelsuensiR ...... 338 /la/iotis (Hulwfiolis) koikei ...... 309 -- slzoroensis ...... 338 J/,wzaslumia ...... :l39 -- sitakarensis ...... 338 ( ?) sp...... 339 -- sub1•m·inalu ...... 3:ls /Jemithyris psi/lacca ...... 306 -- sp...... 338 f!exaphyllia sp...... 325 t\'ipponitrigonia ...... 31·1 !fomomytl saloi ...... 337 1'111/l'l'X(I...... 314 sp...... 3:ll -- kikudziana ...... 314 -- t/171/IIUII/1/i ...... 3l-1 I -- sagall'ai...... 311 -- sakamofoensis ...... 314 lddingsiu orit•nlalis . . .. . 339 ]limzimr /mslzinJensl' ...... 315 -- soradriensc \'ar. lmulmmorit•nse ...... :l15 )( Kallirhyndzia sp...... 327 0 1\.inf!s/onia ...... 339 .. Oslrea" a sp...... 330 -- pm·a/ll•fa ...... 339 "-.. b sp...... 330 Oxy/oma (?) sp...... 330 L Laqueus rull(•l/us ...... 3<)6 p Lmzsdalenidt•s /oriyamai ...... 333 Lucinoma acutilinea/a .305 Pa/aeopharus ...... 350 -- rwkii ...... 34~ -- oblongatus ...... 350 -- oblongalus lmriji ...... 350 -- (Miuep/zarus) lri(l(/icus ...... 350 Panomya ...... 307 Mesoselwberl ella ...... 312 --amp/a...... 307 -- slzimtuluniensis ...... 312 -- beringiana ...... 307 -- /lzlllllfJHIIIIi .... 312 gigcml ea ...... 307 ;l-/e/osira distans \'ar. lira/a ...... 3-13 i Z/111/0 ...... •...... 307 grmmlata ...... :~13 simo/omensis ...... :{07 -- -- forma cun,ala ...... 343 Paralldodon sp...... 337 ;l/ichelimweras lzidense ...... 345 Parasmittina I rispinosa cwmtll'ii'IIHis ...... 329 Index of Fossils 311 Paralymolus yabei ...... 310 Saxoluci11a (J!egaxinus) k·hataii ...... 305 Pan·irh\'1/chia bella .327 Sip!wnodendron japonicum ...... 313 Patinopecten kagamimzus ...... 3·1!\ -- nakazau•ai ...... 313 ---- kagamianus ...... 34S Spiriferinuidf'8 ...... 322 ---- molliwae11sis ...... 348 IICIII(Ii ...... 322 -- 11imae11sis...... 3•18 -- sakmt'clli/IH ...... 322 -- permirus ...... 348 -- yeharui ...... 322 -- aff. kobiyamai...... 349 Pil!ucina (Sydlorina) yoko_ramai ...... 305 P/1'/hometopus lo11gispinus ...... 33~ 'I' Pleuromya lmshidoll'lzsis ...... 337 Tegu/a (Chlorostoma) nm·usei ... 309 Pro/acardia kurumensi,q ... . . 337 Tcre/lratuliua coreauim...... 306 Pseudog!am/u!iiiii obtusissima .315 -- gouldi...... 306 Pteria (s. 1.) kilalmm iensis ...... 330 -- japouiur...... 306 Punctospir~(er .322 Te/mcydus ('f'ialom ...... 343 -- triadil'IIS . .. .322 Tlzracia suhrlwmbica ...... 337 -- triadiws kasfri1miensi11 ...... 322 -- sp...... 337 Pupa hiyoslziensis ...... 33-1 Triccratium simplex ...... ' .. 323 ...... Short note 2 R Uadulonec/ites ...... 32ll -- japonicus ...... 3:!ll v com·exus ...... 320 ---- var. Va1'iam11ssium (?) sp ...... 326 Ret usa (Coe/opltysis) s/wkuborensis. 33·1 l'irl{ulina lwshiroenHiH. .315 - (-) sp ...... 33·1 Volsella baket'elloideH ...... 330 " Rhynchollella" aff. bouchardii ...... 327 -- sp. ····· ...... 330 "--" d. haradai ...... 327 Volmtlella m inoensi11 ...... 334 Ringicula ( Ringiculina) minomzsis ...... 33•1 yamcw!'h i i ...... 334 - (-) sp...... 33·1 sp...... 334 Rutitrigonia ... 31·1 -- yeharai ...... '3J.l '" s TValluci na lwbei ...... 305 Sabalites taislwensis ...... 335 -- okumumi ...... 305 a -lt"~f.~~~ -~·i~f Transactions and Proceedings of the Palaeontological Society of Japan .!\ cw Series No. ~!i ,..._No. :;~ 1957- 1958 Palaeontological Society of Japan November lU, 195H The heading in Japanese commemorates the handwriting of Prof. lVI. YoKoYAMA, father of japanese palaeontology, who was Professor of stratigraphy and palaeonto logy at the Geological Institute, Imperial University of Tokyo. The fossil on the front page is Argonauta tolmnagai YoKoYAMA. I!H3. CONTENTS Number 25 (Published April 30, 1957) Transactions Article Page Plate 30:>. Junji lTOIGAWA: On the l\Iiocene Lucinidae from the rvrizunami Group, Japan...... 1-6 1 306. Jiro KoToH: Brachiopoda from the Daishaka and Tsurugasaka Fossil Zones, Minami Tsugaru-Gun. Aomori Prefecture ...... 7- 10 307. Saburo KANNo: Fossil and Recent Species of the Genus Panomya from Japan ...... 11- 16 2 308. l'vlasao lVIJNATo and Kciji NAKAZAWA: Two Carboniferous Corals from Okayama Prefecture ...... 17- 20 3 309. Matsutaro SmnATA: Some l\Iolluscan Fossils from the Eastern Part of the Tan;..mwa Mountainland ...... 21-25 4 310. I Number 26 !.Published June 15, 1957) Transactions 312. Mosaburo KANUMA and Sumio SAKAGAMI: J11esosclmbertella, a new Permian Fusulinid Genus from Japan ...... 41- 46 8 313. ?vlasao l\hNATo and Makoto KATo: New Species of Siplwnodendron from Japan ...... 47- 50 9 314. Teiichi KonAYASHI: Nipponitrigonia and Rutitrigonia in japan ... 51-61 10-11 315. Saburo YosmnA: Palaeogene Foraminifera from Konbumori, Eastern Ilokkaido. Japan ...... G3- 68 316. ltaru I-L"-YAMI: On the Occurrence of Cardinioides in the Liassic Kuruma Group in Central japan ...... 69-73 12 Publications received ...... 62 Proceedings ...... t'l Number 27 (Published September 30, 1957) Transactions ~H7. Teiichi KoBAYASHI: On an .-ltnria frum the Poronai Shale in Hokkaido ...... 75-80 13 318. Rikizo h1AIZUMI: Three new Iviiocene Species of Callianassa ii from Nagano Prefecture ...... 81-85 H 319. Tatsuro l'viATsuMoTo: A Turonian Damesites from Hokkaido, .Japan. !Studies of the Cretaceous Ammonites from I-Iokkaido and Saghalicn) ...... 8G- 88 15 320. ltaru I-IAYAJ\11: Radu/onecliles. a new Pectinid Genus. from the Liassic Kuruma Group in Central Japan ...... 89-93 16 321. Kotora HATAI: Interesting Sand-pipes from the l\liocene Toyoda Formation. Yamagata Prefecture. Japan ...... 95-98 322. Akira ToKU\'AMA: On some Upper Triassic Spiriferinoids from the Sakawa Basin in Prov. Tosa. Japan ...... 99-106 17 323. Wataru lcmK"nvA: Some Notes on a rare Species Triceratium simplex ]. BRuN ...... 107-109 18 Publication received ...... 94 Proceedings ...... 110 Number 28 (Published December 15, 1957) Transactions Teiichi KonAYAsm: An _-{furia from (?) Kusu County. Oila l'reft•cture. Kyushu ...... 111-114 19 Takashi YosHIDA and Makoto K."-To: "Onimaru type" Corals newly found in the Northern Kitakarni Mountain Region. Japan ...... " ...... " ...... " .... " ... """ 115-117 326. Itaru 1-lAv AMI : Liassic C/llamys . .. CmnjJ!onectes " and other Peel i- nids from the Kuruma Group in Central Japan ...... ! 1!)-127 20 327. Akira Toi Number 29 (Published April 10. 1958) Transactions 330. Itaru HAYAMI: Liassic Vo/se//a, .Mytilus and some other Dysodont Species in japan ...... 155-165 23-24 331. Tatsuaki KIMURA: 1\lcsozoic Plants from the Tetori Series, Central Honshu. Japan ...... 166-168 25a :{32. K(·iji SuzuKI: On the Occurrence of Cercis in Japan ...... Hi!)-.171 25b 333. l\lasao l\IINATo and l\lakoto KATo: A short Note on Lonsda/eoides toriyamai l'vliNATo ...... 172-174 334. junji hoiGAWA: On some Cephalaspid Opisthobranchia from the iii japanese Miocene ...... 175-183 26 Proceedings ...... 184 Number 30 (Published june 20, 1958) Transactions 335. Kiyoshi TAKAHASHI: Sabalilcs am; den Wakata Schichten von Tsushima, Nordkyushu ...... 185-188 27a 336. Koichiro MAsunA: On the Miocene Pectinidae from the Environs of Sendai; Part 10, On Pecten (Aequipecteu) ymwgawaensis NoMU· RA and ZINBo ...... 189-192 27b 337. ltaru HAYAMI: Supplementary Descriptions of the Liassic Pelecy- pods from the Kuruma and Shizukawa Groups in japan ...... 193-200 28 338. Masaru MATsUI: Species of the Genus NePtunea from the Palaeo· gene Formations in the Kushiro Coal Field, Ilokkaido, japan ...... 201-210 29-30 339. Teiichi KoBAYASHI: Some Cambro-Ordovician Fossils from the Tan'gyang or Tan yo District. South Korea ...... 211-21<1 340. Keiji NAKAZAwA: ·· ll1onojJ/iyl/iles" arakurPnsi~ sp. nov. from the Maizuru Zone ...... 217-219 31 Proceedings ...... 220 Number 31 (Published August 1:1, 1958) Transactions 341. Koichiro I\fAsUilA: On the .Miocene Pectinidae irom the Environs of Sendai: Part 11. On PerfPII ICh/amys) rmssiveuius YoKoYAMA and its related Species ...... 223-229 :~2 342. Katsumi H1RAY AMA: On Luri!loma ao/{ii, a new Pleistucl'nc Lamellibranch from Chiba Prefecture. Japan. Notes on Japanese Lucinid l'vlolluscus. Part 2...... 231-236 :~3 343. Haruo 0KuNo: Electron-Microscopic fine Structure of Fossil Diatoms, V: Observation on some Diatoms found in Celatoms ... 237-242 34-35 344. Masahiko AKIYAMA: New Fossi I Ch/amys from the Environs of Nagano City. Japan ...... 243-247 36 345. Tciichi KoBAYASill: A Gotland ian Nautiloid from the Hida Plateau in japan...... 248-252 37 346. Tsugio SnuTo: Granulifusus from the Miyazaki Group (Palaeon- tological Study of the Miyazaki Group. V J ...... 253-264 38 347. Riuji EN Do: Stratigraphical and Palaeontological Studies of the Later Palaeozoic Calcareous Algae in Japan, XIII. A Restudy of the Genus PllysojJOrel/a ...... 265-269 3!l Short Notes 2. Wataru lcmKAwA: Once again on a rare Species Triceratium simPlex ]. BRuN ...... 230 iv Proceedings ...... :ao :\umber 32 1 Published November 10, 1958) Transactions 348. K6ichir6 rvL~sUDA: On the ~iliocene Pectinidae from the Environs of Scndai; Part 12, On J>ecten kagamianus YoKoYAMA ...... 271-284 40-41 349. Wataru HAsiMOTo and Saburo KANNo: Molluscan Fauna from the Tertiary Formation of Chirai. Kamisaroma. Kitamino-kuni, llokkaido...... 285-290 42 350. Akira ToKUYAMA: Late Triassic Palaeopharus in japan ...... 291-298 43 351. Rikizo hrArzu:o.n: An interesting Crustacean I~emain. Ctenocheles sujakui n. sp ...... 299-304 4,1 Proceedings ...... 305-306 Systematic Index ...... 307-308 Index of Fossils ...... 309-311 NEWS International Oceanographic Congress Lt. New York 0 United Nations Building "(' 1959 ~: 8 f] 30 Fl ;/Jd:, \I } J 12 1:J 1 "t'001J•H 7.> ., C. C. L'li (I) The History. (2) The Boundaries. (3) The Deep Sea. ('I) Dynamics of Organic and Inorganic Substances &V' (5) The Marine Lift Regime U> 5 symposia :~;:,Hiifli~tn::t>.Oil:f!i1 5 (i) symposium L'l.l~0 5 P.~mil:Mil&~tl.Oo (a) The palaeogeography of marine floras and faunas, (b) Biogeographical regions in the sea. (c) Evolution and adaptation in the sea. (d) The behavior of marine organisms as influenced by environmental factors, (e) Physiology of marine plants, (f) The culture of marine organisms as a means of understanding cvironmental inllucncc on populations. ~'f.l4~'irlltJ!C.IIIJ{t'ltH!i.t t •,. Dr Mary SEARS, Chairman, Woods Hole Oceanographic Institu tion. Woods llole, Mass .. U.S. A. ·lth Panafrican Congress on Prehistory (;t«Jv.;t:-f,U=":.--="·0 Lcopoldville "(' 1959:1f:8H 2-1 -.....29 ~w:.nHmt il n .o c..t !U.J: "? t.:o ~~LHf. t ~$jllJ~·f.OO{~. ~ 2 $tli~I.:~·Y.~II,t;. m:i $%~ ~-i'.l!'f: W:I(M) 3 HlltuH::.5HJ•tt1."[Jfh'J·~1 7.> :YJi:L'J.> 7.>" ~'f.AiiHi.rJCll::.llllfr-tt G~l U \, Secretariat du IVme Congres Panafricain de Prchistoirc Laboratoirc de Geologic Universite Libre de Bruxellcs 50. avenue F.-D. Roosevelt Bruxcllcs (Belgique) IJil nu ~-& I IIH Ill! s I ~iJi( $~tfw a 1958 !F *~·if§:; * * * ~ I 1958 ~f 12 .F.l 6, 7 f3 I 1958 $11 R~ m1 2 lill ~J ~ 1 a;. o * ~ 19ss &f. 2 R 14 El I 1959 !F 1 F.J zo a 1 9 5 8 if~ 11 }j 5 II ,:P .!ill! 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