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Ohio Shale Conodonts

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Ohio Shale Conodonts OHIO SHALE CONODONTS RALPH H. BOND, Department of Geology, Capital University, Columbus, Ohio Although conodonts are numerous in the Ohio shale, they have received very little attention. To my knowledge, Cooper's paper (1931a), in which seven species are described from the top of the shale just northeast of Columbus, is the only publication on the subject. The present study encompasses a large variety of species and is the forerunner of a comprehensive study of the conodont fauna. The material for this paper was collected from the lower portion of the Ohio shale where it is exposed along a few small tributary streams of the Olentangy River about two miles northwest of Worthington, in central Franklin County. Thirty-seven previously described species and eighteen genera are recognized. Twelve forms are unassigned because of doubtful relationships. Among these may be several new species and at least one new genus. It is hoped that these doubtful forms can be described when additional material is collected and studied. The species described in this paper are listed as follows: Ancyrognathus irregularis Branson and Mehl Metaprioniodus biangulatus Huddle Bryantodus commutatus Huddle Ozarkodina delicatula (Stauffer and Plummer) Bryantodus concavus Huddle Palmatodella delicatula Ulrich and Bassler Bryantodus germanus Holmes Palmatodella ? sp. Bryantodus inequalis Holmes „ . 1 *1 TT1 . , , _ , Bryantodus nitidus Ulrich and Bassler Palmatolepis glabra Ulrich and Bassler Bryantodus serrulus Huddle Palmatolepis quadrantinodosa Branson and Bryantodus subcarinatus Huddle _, Mehl . Bryantodus subequalis Cooper Palmatolepis regularis Cooper Bryantodus subplanus Huddle Palmatolepis subperlobata Branson and Mehl 77 j. • • -,- j, rr JJI Polygnathus nodocostata Branson and Mehl Buprwmodma prona Huddle Polygnathus pennatuloidea Holmes Hibbardella angulata (Hinde) Polygnathus sublatus Ulrich and Bassler Hindeodella aculeata Huddle Polygnathus sp. Hindeodella alternata Ulrich and Bassler Prioniodina separans Holmes Hindeodella germana Holmes „ . , , , ., TT , Hindeodella subtilis Ulrich and Bassler Pnonwdus alatoides Holmes Hindeodella sp. 1 Pnonwdus alatoideus Cooper Hindeodella so 2 Pnonwdus alatus Hinde TT. , , ,, ., ... t TT ,,, Prioniodus cultratus Ulrich and Bassler Hmdeodelloides bicnstatus Huddle i7 . , . ,TT , . ... ,. 1 Spathognathodus subrectus (Holmes) Lizonodina sp 2 Subbryantodus radians Branson and Mehl Ligonodina sp! 3 Subbryantodus sp. Lonchodina multidens Hibbard SynpHoniodina sp. Lonchodina perarcuata Ulrich and Bassler Distacods, 3 sp. Lonchodina perlonga Ulrich and Bassler Dermal plates, 2 sp. No additional evidence is furnished by the present conodont collection to aid in the solution of the general problem of the stratigraphic position of the Ohio shale. It is expected, however, that future studies will reveal data of value in determining the age relationships of the formation. The conodonts occur most commonly in very thin bone beds in the shale, with only occasional specimens scattered between the beds. Some of these bone beds are located at the contacts between the black shale and interbedded layers of gray shale. Others may be found as very thin layers along bedding planes in black shale of uniform lithology. In most places the conodonts are rather evenly dis- tributed laterally, but in one place at least they are concentrated in small discontinuous patches which consist largely of conodont fragments. This latter occurrence suggests that they were brought together by small eddies or currents in 21 22 RALPH H. BOND Vol. XLVII the water and that they were moved about considerably as they were being deposited, thus resulting in their fragmental condition. The state of preservation of the conodonts at the different exposures varies in a marked degree. In some places the fossils are largely broken, but the fragments are excellently preserved. In other layers the conodonts have been dissolved away by the sulfuric acid produced by the weathering of pyrite and marcasite in the shale outcrops and have left clearly defined impressions or molds which are usually sufficiently complete to permit identification. The conodonts are quite variable in color, some being white, some amber- colored and transluscent, and others are colorless and so transparent that the shale on which they lie may be seen through them. Many of the specimens examined have a high luster. In general, the fossils of any one bed are similar in color regardless of the species. Associated with the conodonts are small dermal plates, minute black spines, small carbonaceous fragments of wood, and a great abundance of plant spore cases which are in some instances so numerous that the surfaces of the shale are almost completely covered with them. In two of the bone beds which were studied a fine network of minute cylindrical tubes was observed. These are filled mostly with pyrite and marcasite crystals, but in places they are hollow. Their identity is unknown. For the most part the conodonts described in this paper had to be studied in place on the slightly weathered surfaces of the thin layers of shale, because when the rock was broken down the conodonts fell apart along minute fractures. Some fossil material was separated successfully from the shale by boiling in a solution of sodium hydroxide, and the conodonts were recovered by sifting and by using liquid separates. Material thus separated from the shale consisted largely of fragments, but some identifiable specimens of the simple cone type as well as many minute specimens of the platform type were recovered. These will be studied at a later time. The total number of fossils preserved sufficiently well for study was about 450. This is a rather small number considering that at least thirty-seven species are represented. Branson and Mehl have also noted a proportionately large number of species relative to the specimens examined in their collections from the Harding sandstone of Colorado (1933a, p. 22), and from the Grassy Creek shale of Missouri (1933b, p. 183). Acknowledgment is made of the assistance of Dr. Grace A. Stewart of the Department of Geology, Ohio State University, who directed the paleontological research. EXPLANATION OF PLATE I All specimens are from the lower portion of the Ohio shale, upper Devonian, central Ohio. Magnification as noted. Figs. 1-3, Distacodidae. Figured specimens: X50 (p. 23). 4, 6, Prioniodus alatoideus Cooper. Hypotypes: 4, an impression, X30; 6, X 25 (p. 23). 5, 7, Prioniodus alatoides Holmes. Hypotypes: 7, an impression. X25 (p. 23). 8, Prioniodus cultratus Ulrich and Bassler. Hypo- type: X25 (p. 23). 9, Prioniodus alatus Hinde. Hypotype: X20 (p. 23). 10, Ligonodina sp. 1. Figured specimen: an impression. X12 (p. 23). 11, 12, Euprioniodina prona Huddle. Hypo- types: impressions. X25 (p. 23). 13, Synprioniodina sp. Figured specimen: an impression. X20 (p. 23). 14, Ligonodina sp. 2. Figured specimen: an impression. X10 (p. 23). 15, Palma- todella delicatula Ulrich and Bassler. Hypotype: X20 (p. 23). 16, 17, Hindeodelloides bicristatus Huddle. Hypotypes: 16, an impression, X20; 17, X25 (p. 23). 18, Ligonodina sp. 3. Figured specimen: an impression. X12 (p. 23). 19, 21, Hindeodella aculeata Huddle. Hypotypes: X12 (p. 23). 20, Hindeodella sp. 2. Figured specimen: an impression. X25 (p. 23). 22, Hindeo- della germana Holmes. Hypotype: an impression. X15 (p. 23). 23, Hindeodella alternata Ulrich and Bassler. Hypotype: an impression. X24 (p. 23). 24, Hindeodella sp. 1. Figured specimen: an impression. X14 (p. 23). 25, Hindeodella subtilis Ulrich and Bassler. Hypo- type: X20 (p. 23). 26, Palmatodella ? sp. Figured specimen: an impression. X20 (p. 23). 27, Prioniodina separans Holmes. Hypotype: X12 (p. 23). Ohio Shale Conodonts PLATE I Ralph A. Bond 18 19 21 22 ?5 26 27 23 24 RALPH H. BOND Vol. XLVII SYSTEMATIC DESCRIPTIONS Family Distacodidae (Ulrich and Bassler, 1926) The specimens illustrated on Plate I, figures 1-3, represent a small group of distacods recovered from conodont bone bed material by boiling, washing, and screening through the 100-mesh sieve. They are somewhat smaller than many of the distacods described in the literature, but they compare favorably in size with several specimens from Pander's conodont material which was available for com- parison. An opaque white appearance is developed more or less completely in these tiny cones, a characteristic which they share with the blade or platform type conodonts recovered with them in the screenings. This opacity conceals the growth axes. The upper parts of the cones have been broken away, but otherwise the luster, striae, and thin edges of the basal cavities are well preserved. Pending the collection and study of additional material no specific nor generic assignments can be made with any assurance at this time. The illustration on Plate I, figure 1, represents a minute, simple cone having a base which is extended and flared both anteriorly and posteriorly. The basal cavity is moderately large. The cusp is thin, long, curved moderately, slightly compressed laterally, smooth, with a faint keel developed on the anterior edge. (Figured specimen, Ohio State University Geological Museum, no. 18367c.) Figure 2 on Plate I illustrates a simple cone with a small oval base which is very slightly expanded and has a rather shallow cavity. The cusp is oval in cross- section, smooth, and slightly curved, forming a high angle with the plane of the base. (Figured specimen, Ohio State University Geological Museum, no. 18367a.) The specimen
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