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ltt (9ll6t)6rf s tt :(flE'?rsr^fN Nullsia!o1. 5 No I (1986) of its closcrclirtivcs in Baeckeqwhilemaintaining Ilyltctcal.vmmuF,ndl. as a distinctgenus. H.vpotalvmma with its more nurnerousand exsertcdstamens with basifixed anthers is distinct, but its reniform seeds,anllrers opening in slits,and fruit morphologl', sho\\'that it is more closely related Io Rinzia than Rinzia is to Bentham's seclionsHclrmogia. Iluhingtonia and Sthidiomyrtus of Bae(ked.

Rir',zlais reinstatedhere as the typc spccies,Rinziajinlana, has beenfound to belong to a group ol specieswhich can be separatedfrom their nearestrelativcs on the basisof the followingcombination ol characters:

( l) Filamentsllattened rather than tcrcte:ind oltcn *'ith an emarginateapex (especialll, thc f ilamentsoi the antcpetalousslamens).

(2.)Anthers attached(but only at the mid-point) to thc adaxialsurface of the filament (scctcrminology and morphologysection below, and alsoFigure I parts2 and 3) rather tlran dorsifixed and versatiie(as in scction Euryomyrtus ol Buetk.eaL. and some related easternAustralian species)or basifixed(as y1 H.1,pocall,ntnta),but seethe note under R. (ornosa which is an cxception as it hasdorsilixed, versatileanthers.

(3) Stamens l0 (or l2 when tire I'lowr:rsare 6-mcrous),or when lesshaving one or morc of thc antcsepalousstamens missing.

(4) Flowersborne on a welldevelopcd anthopodium, but havingonl),- a rudimcntary pcclu nclc.

Thc first of thcsecharacters (llattened lilaments) separates Rl/?.zid from the other species of Baeckeinaethat have reDiform sccds.Therc arc parallel developmentsol flattcncd filarncntsin othcr groups of specieswithin the Baeckeinaebut the spcciesin tltesegroups have different seeds, anther attachmcnt and placcntatlon. For examplc, Baer'lrerr drurnmondii tyasincludcd in sectionRrrzl4 by Bentham( 1867).but whilc its stanens have bload filaments, its anthers.placentation and secdsindicate that it is not closelyrcliited Io Rinzia as the genusis deflned lrere.lt is felt that the combination of charactersgiven abovcquitc adcquatclyjustifies the reinstatementofRi ziato gencriclevel.

Systematicl'osition of Rinzra

Rlr-ula was placedbv Schaucr( I 843)in the Baeckeae.while Bentham( 1E67)treated the genusas a sectiono[ BaetkeaL. which he placedwith Hypctr:ttllrttmu,S(holtzia, Astartea "Baeckeaeae" a:.ldBalaustion as a subtribe of the tribe I.eplospermeae.Niedenzu (1898) apparentiy recognisedthat thc speciesplaced in section Rll?zl4by Benthanrtl'ere more closelyrelatcd to Hlpocel.|mmu as he removedthcm lrom Baetkea and placed them in Hlpotalynrttta (he perpetuatedthc error with Baetkeadruntmctnrlli however). While this placementrccogniscs the cLoserelationship between the speciesplaced in Rrn;la and those in Itypocalvnma it ignores the dillerencesbetween thcm. Briggs and -lohnson (1979) considcred that Bae(keo neededluthcr study to determine whether or ltot its scctions (including scction Rirzla) deservedgeneric rank and placed it in a Bqet:keosuballiancc ol their Chameldrclum alliance. subsequently(.tohnson and Briggs 1985) they have abandoncdsuballiances tn the Chamclautianralliance.

It is outsidethe scopeof this paper to considerthe relativemerits of thcse placements abovethe genuslcvcl. The more recentsystem o1'Briggs and .lohnsonis not usedbecause it has only beenproposed informalll'. '(SZI {aq dN , LUorJIIV) (0lx) saurltnor€tnSorrrpup,r\EIr 3ur^{oqs qBr3d o.{\l 9 (0fx) saln^oo^\l.qlJo luetuqreli?3ur^\otls eluor€ldJo uorl.os ()1 ssorr pu€ €tuar?td uo .tn^o Jo ^\ar^eprs E (ozx) uolltluar"ld 3t]] put unrqru?dtrqiql ,{r€^oaql Jo uolsnj oqt trrr^o aqt otur al{ts oqt Jo uorlrasuraql 8ut^loqsro^ olJ :qr qSnorqr uortrosIPurpnlr?uo'l t (0tx) puPIS .^r1rauuo.r' oqsotp.^oruarr.!llu€q1r.{\auespuesplsLuorluau€]ssnol€desslutJolurLUrlUJodrlpueraqluV t '(lno p.u:1lpu'0Zx) aer^ IErx"p€ surtutls Z (Pa^orur stelrd .LUos'0p.) ra,roll I r',/rrtr!.,rz,y I ornBtJ

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,rzury uiBpnrl tutortEt,,t LI' E 4lE NuylsiaVol 5. No. I (1986) Terminology and MorphologY

Anth(padiutll is uscd as defincd by Briggsand.lohnson (1979),("l-he intcrnode between ") a flower and the most distal node of the axis that it terminates to describepart oI the axis on which the flower is bornc. ln Ritlziatine anthopodium is the longcstpart of this axis, the proximal part of which is hcre reierred ro as lhe pedun(le and which is quite shortin all nembersof the . lril is usedto describethe outgrowth from the hilum which claspsthe seedof most specics ol the genus.This appearsto be the firsi time sucha structurehas been reportcd 1or the .lt is dry andquite thin in driedspecinens but su'cllswhen the secds are soaked in water,indicating that in fresltseeds it may be 11eshy.No infornation iscurrerltl-v available as to what role it might play in the dispersalof the seeds.See photographsof seedsin Figure2.

Hypanthiun has been uscd to reler to the cup or tube around the ovary. This term has bccn widely usedfor this structurein the literatureon the Myrtaccircand reflectsits most "floral probableorigin which is a devclopmentofthe receptacle.The tcrm tube"is rciected on thc basisadvanccd by Briggsand Johnsott(1979), i.e. thal it is olicn confusing.Briggs "perigynium" and.l ohnson ( 1979)used the tcrm but rejectedit in a laterpublication (Briggs and Johnson1985) in favourof hypanthiun.Where thc diameterol thc hypanthiumhas beengiven it was measuredat thc baseof the calyx lobes(this point usuallyhas the largcst diameter).

Floh,er size, which is quite valuable in dilferentiating betweenspecies, is given as the diameterof 1hecirclc that would includcthe petals(e.g. 5.0 8.0mm acrosscorolla).

Anther attathnten /. Exccpt in R. carnosathe anthersin Rinzia are attachedat their mid- point to the adaxialsurface ol the filamcnts(see Figure Ll) ratherthan beingdorsifixcd and versatile (i.e. attnched at the tip of the filament to the mid-point) as in sectiott Euryont.,-rttt.sol Baetkt'o or basifixed as in Hypcttalyntmn. The dorsifixed, versatilc condition with terelefilaments is thc most widespreadin the natural group to which Rinzla belongs,occuring in speciesgroups with diflercnt stamenarrangements, different sized and colourcd sccdsand dilferent mcchanismsoi fruit dehiscence.and as it is also widespread in the Myrtaceae as a whole is consideredto be the primitive condition. The condition in Rinzictisinterpreted as davcloping through the extensionol the lilamcntslrom the point behind the attachmcnt point to give the appcarancethat the anlhers have moved from thc tip to the adaxial surface,with the filaments appearingnot to end at the centrc ol the anthersbut to carry o n, oftcn cxtendingpast the top of anthcrs.Although still dorsilixed this constltutcsa characterstate diflerent to dorsifixed. versatileanthers and basiiixed anthers aDd is one of the charactcr stateswhich separalesRifzid lrom its rclativcs.R. catnos.thas dorsifixed, versatileanthers but has the broad lilaments of Rinzia. This may indicate that the adaxial position of the anthersin the other spccieshas cvolvcd within the genus.Thc basifixedattachnent \n Hl,potolymnta appearsto havedeveloped through the fusion of the filament to the lowcr pa of the anthers.

Slamen arrangemenl.As noted above thc stamensare typically very regularly arranged, with one oppositeeach petal and sepal,although there are variations.Johnson and Briggs (1985,Figure 8) haveproposed a schematicoutline of trendsin androecialevolution in the Myrtaceae,with the dcvclopment being from fasciclesopposite the pctals to single stamcnsopposite the sepals,or irregularly spaced,in the Bae(keasuballiance (Johnson and Bliggs 1985,Figure 8, J-M). Thc typical arrangenlentit Rinzia could be viewcd as "sequence" part way do\\,n this with the occasionalpair of stamensopposite the sepals tr'i! I sr rPq at'?rs (rruorsrl|tzr )rrf)s :) /,1J Luorl 'sp3s 'sPe.s 'Y Z),ror.r7.r y pup (uriurtp-) Jo ,rS urt 6 S6lZ u)B rJ t A uro.rJ Z) ".)/l\rrr') ,\\oi tuorrog (uptrp).,.^\otrNZ/trrfl]Y (7yruorj sp.rsZ) Dtro/intPq )s a (!|llnDuotultSolffur)aIxl lL'r-tilrl^J 06lo Jrutttsglt DxpnL 7 n LtIorJ\pris a) D.tr|t|.ty'(aiutx )r.d qlunql ltlr arlt))D s v rL'orl sp.rs a)eptD)a).i\o y 1{orpirql (.3utd SurtIt\'sr.rllrlsoHJo s.dolsuriqlrou r3$ol 6lll 1./,V,r,v/, tuoU '\peas 'ir^r uotu.totu y uoproD qrno\ t\n t.{P^\ttirH ,iunqlV rrN l ,,,vlrrl I N ual.l spaesa) ouDtnl a d 'r}t9 Jo y (ppor rirg supetio .)if(r :.tDu1;0 l/ ruorj spi.s Z) Dtt,tDqdtotury y :^\oi puor;S (.rrJJor\od Jo 'tgnl tur) ?)tr'1Jo lsci srt[rr 8t urpn4l :7 /4 ruo.rj sp.:s t)'nrr]'rl'll.,.r y (iro)l urrn-L CtSfl DullD!) V.) ur,rij spe:. a) \u1./.r1V :^\or do1 rru,or prirtl r}rt dor luorj rq8u or ni.l sarr3ds,rz,?y t(, 1p..S Z .rntr:l

6lt .r rr4l u,ipr I I,,rt'irtft\ 420 ,luyrsiaVol 5. No. I (1986) rn R. (arno.sapossibLy indicating this spcciesis lessdeveloped in this character.Those specieswhich losc somc or ali of the antesepaiousstamens would indicate a divergence in Rinzia from the the trend to singlestamens opposite thc scpalsonly.

Inflorescente:The most common inllorescencestructure is a monad with a very short peduncle and a relatively long anthopodium. Mctaxvtriads occur rarely in R. dintorphandrawhich usuallyhas d ichasia,the two flowersof which appear1o be at d iffcrent stagesoi development.The latter situation also occtrs in R. aflinis.

TestasurJa

Protesses;Hair-like structuresfound at the basc of the abaxial sidc ol the antepetalous lilaments oi somespecies (e.g. /1. .ie.r.rihs).They may be vcstigialstaminodia.

Materials and Melhods

This paper was based on thc study of the gross morphology of herbariun material borrowedfrom Australianherbaria supplemented by observationsand co]lectionsmade in the field of nine of the twelve species(R. carrutsu,R. bngiJblia und R. o-xytottoide.t were not obscrvcdin the {ield by the tluthor).

The material examinedincluded type specimensof all the previouslydescribed species.

The specimenscited were seiectcdro cover the geographicalranges of thc species,with preferencebeing given to collectionswith duplicateslodged in more than one institurion.

The measurementsgiven for the parts of flowerswere i]lade on material that had becn boiled in water with a small amount of dctergentin it. The mcasurementsol lcavesrefer to dry material.

Rinzrc Schauer

J. C. Schauer,I-innaea l7: 239 (1843\.-Baeckeasection Rinzia (Schauer)Bentham, Flora Austrai. 3: 74 (186'7).Type: RinziaJumana Schauer.

Subshrubs low and straggling to ercct, or with prost.ate branches,rarely enti.cly prostrate.Glabrous cxcept for onc species.Leavcs opposite, app.essed to spreading,shortl), petioiateor sessile(one spccies),linear to orbicular,ovate or obovate,flat or margins recurvedj or variously thickened (plano convex or concavo-convcx,abaxial surface shallowlyconvex to semi-tereteor semi-elliptic),entire, rarely with hairson the margins, glanduiardotted, occasionaily discolorous. Flowers l-12 nearthe tips of the branchlets, or alongthenl, small to medium(3- l9 mm diamcter),solitarv or 2-3on a commonpcduncle in axils oI upper leaves;pedu ncles very short wilh a pair of terminal bracteolcs;anthopodia eachwith a pair of bracteolesat thc basewhen more than one flower on a peduncle,0.g 27 mm long; bracteoieselliptic to cymbiform, persistcnt,olien coloureclor with a coloured edge.Hypanthium obconical to hemisphericalor shorllycylindrical. Calyx lobcshaif to as long as the hypanthium,triangular, deltoid, semicircular or semi-elliptic.petals suborbicularto obovate,or angular.Stamens 5- 12, usually l0 with oneopposite each petal andcaiyx lobe, when less than l0 all or someof the antesepalousmissing, l2 whenflowers 6-mcrous,or occasionally( R. (qrnosa)two stamensopposite some calyx lobes.Filaments all flattened,the antepctalousbroad, the antescpalousnarrower, the antepetalouslonger than the antesepalousand fused to them from shortly (near base)to up to their whole length, sidesstraight or incurveci,apices truncate, rouDded, or emarginate,antescpalous somctines pointed. Anthers usually attached to thc adaxial surface of the filaments, dorsifixed and versatilcin one spccies;loculi parallel,opening in straight or rarcly curved etr?llrJespeos :snlncol JJd Z seln^O / osoulD,-A g Buolurur E l-t J c speos:ssorJe 'passJjddr? ruru 9 a-S S srJ,{olJ:ra^uoc o,\EJuoJ se,\"e.I g D.tqn.tU 9 3uo1u.rur/lglJspoes:ssoroe 'Surpue:ds ruru II-9 / srJ,{olJ :xe^uoc-oue1d se^ea.l g 'elelluD tou spees:snlnJolled 6-t seln^O / 'slrx" Jeolur liJplrloss.re,roll g ' DipuDtldtotutp y l, 3uo1ruru 9.1-1 J spcf,s:otrq4\.{lalello lurd slJ,'!\oU:lredlxeu Surddplle^olsnf 'lpcruoJqo se,ltal:irnll ur IJrqt,{lrre1pue peurd,{letnuru tsourle 'leJupurl^c ,{1f,.!Er {luoqs ol I"JrJeqdsrtuaqrunrqluud,{g ' . 9 slu!/Jb A 8 8uol ulLug l-tl spe.s:)iurd^luouluoJ ss.l f,trq^\ sra^\oU :./t-s/'l,{q :red lxeu SurddellJ^o ,{lJEnsn sa,\EeJ:lrnrj ur snoeJeu"qopue qloours'l"JruocqoluntqluedFH .g 'sllxe ur s:redut sla,\\olC g '(ruru JEcl S 9-Z I) DtloflssDt)y pue (urur g g-92) Dkln|y roJ ldaJxaSuol ruru 'x.,\uoJ-oApcuoc .l t 8 0 €rpodoqtup :snoJolocuoJ :o xe,luoc-oue1dsoAer.l iluostl.lou) y Z iuol trlurg l9I suaLuEtssnoleledelue :a,\oqE se lou Jsr,rlJqto lnq (eiBllsold Sutpnlcur) rulol ut JIqeUe^ s'lu€ld . . t Dtlo/),iuol )1 t tuol uru t Z-6 1 sueuets sno1ul:de1ue:pa^eol ,{losuep ,{llensn'lloqs't.?.rosloJqJuplq puc elu.rlsordsrucls urEru qtr,r slueld .t '3uo1 uur 9 l g l spres:quprruuroJ] peqcrpse^eal ro'pe^rncer sutileru ot tpu li^outl"saleol :snlnJol:ed g-g se1n,rg.g " ' ' -...... 3uo1 srpto.))o.).1xoU t tulu g.l-1,.1 .t speos:pe^I]Jet {ltueururordsurSletu JDel:snlncol Jed ZI-g selnAO 3uo1uru g.l-9. I spees :Surpeordssle]Jd :pllolol sso.rJEluru 6l-g sjc,{\ou:snlnJol led seln^O " ZI-g U Dtlo/t.tallotl)sy I 3uo1ruu t l-t.l spres:,{lepr,u ?urpee.rds .U iorr .{llensn-c1u]cd :e11o:olt ssoJce tuu g-g sl.,rou lsnlnJolJJd g lo t seln^O '(DtlarLalp?s y Jo suoulnodsetrlos ul ,q ldeJxourut I u?ql Jalpelt) tu ol utut l7-y r,.rpoOoqtue :sno.Io1or"rp 't pru-rruo t.1 'pJ^JnJel peqr:t,{lyeuorsecco flluoulruoldo},{ltq8rls urSlELu lo tpu sel€o.l .l sarradsoq1 o1 f,ay

'urlErtsnV 'serceds .uoLlngub.to uiatsJy'AJo .rouroJtser\\ qtnos iqt ot lrulepue 11u e[e,'\ L '..JJuqJ,, ']urJnlsuell sp Eurdole,taptou .urqt sJIn^o p.uoqv elJrpel 'oluo !^rssrTruJLIt punol? pedd"lA\lo Iceq peploJIcau pup suopel[toc,!uaq1uJJ,{]eqeueld ')rf,u .qt oj lallpied paur]tPI., repualsE uo llpLussuopel,{toJ :pacs SurllrJ o,{rq!ug u,\\orq 'lsollrded 'snoeJetsnJc rirEpol u,r\orq ol rlloorus e']sal:oprs e,\pcuoc eql Jo Jlluot ur utnlrq 'ullo.ltuoJ .eJlu.J :(D.rqnl V p\tEDsou.tD) U ur td..ye) otellltE spe3s cqt tp ur peulnl ucuo 'padtqs-lacnes ritl?JuropunJqt ro pJu.]]pU !unrqluEd,{q:(srxe trnl] aql o1 sr18uelqBU le 'pepunol tsoluleIrlun) ,{1apr,tr Suruedo so^le^:ttnlj poJsrqo(lluepuad'esrq eql lpeu ^luo urnrqlued,{qo1 pasnl's3qol x,{lpo eq) o'} IEnbo T'rlnsder lecrrpurlfc.ro.relnqolS e llnrJ uls 'pe9ue:re yeurpnlSuolrqt Jo eprsraqlre uo eluarpld ?ql 01 peqoele ^llerper lsoutl" ^loter 'snlnJol 'tq8relts ,sJrcods ro s,\\oi ol\t ur a ro leretplloi lJd 6-Z eSpelprxEq" 3q1 [latel 'rulo.lrucl ,peIlels aruosur pclurodpue euo seln^O llls i(11uurpn1r8uo1 ]ou tnq ,setceds 'srrD eruosur orlueJiro,.lt J"eu peqJe]le,{luo lelou .ql uo seeject1drl1e pesru.r Jo SurtstsuoJ sEluaJrld :Jl!xe uor'lElueJeld 31,{ls asEq punotr pesse:dap.ptoAoqo ol lelnqolS 'Jelncol-t Jo ,{lJrrr ro -f ,{JeAO potlEluJ tou lo e}pJrdEJulu8tts :{1"^o eql otur ,{ldeapol 'lnols 'aterel Iltroqs p.tresur ot rcpuJls al,{ls pto^oqoot t"lnqolSpuelS e.lrtceuuoJ strls t.f ,r:r?ry ui;prrl :I urlortr t\ NuylsiaV01.5, No. I (J9E6)

9. Flowcrs I 12, not clusterednear ends oI branclllets; seedsverrucose or with endssomewhat pointed; leaves not regularlyfour ranked (i.e. not quadrifarious) 10. Seeds vcrrucose,ends roundcd; corolla 7.5- Il.0 mm across...... 12.R. craseilolia 10. Seeds smooth to colliculate.ends somewhat pointed;corolla 6.0-7.5 mm across...... 9.R.lumana Flowers I-5 clustcrednear ends of branchlets;seeds ncither verrucosenot with ends pointed; leavcs regularlyfour ranked(i.e. quadrifarious). I L Hypanthium hemisphelical:petiole absent or indistinct...... 1l ll sessilt I1. Hypanthium shortly cylindrical or rarely broadly hemispherical;leaves definitely pet- io1a1e...... 10.R.

l. Rinzia schollerilolia (Lehm.) Trudgen, comb. nov. Baetkettsthollerilblla Lchm, Pl. Preiss.2: 369( I 848). I-1'pe.Seven Mile Bridge,Plantagenct, , Preist2015 (iso:MEL).

Subshrubto 20 cm tall and 60 cm across,slctzs spreading to procumbent.or scrambling, or lower ones, or all, prostrate. Leave.tdistant, spreadingto divaricate,or occasionally twistedtowards each other: lamina lanceolate, acute 3-10 mm iong,l-2.5 mm wide,almost flat to margins recurved,smooth, discolorous(pale bclow). often reddish,especially near tips of shools.oil glands rarely visible. Flower.sl-4 on a slloot, solitary in axils ol upper leaves,crect; anthopodia 4 l6 nim long: bracteolesnarrow elliptic, concave,acute, 1.E- 3.0 mm long, oftcn reddish. H,t-panthittmobconical, 1.7 2.4 mm wide, green or reddish; calyx lobeserect, not keeled,semi-circular to oblong,1.0-1.7 mm long,petaline at least on margin, pink to white then with reddish blotches,entire to lacerated.Cotolla brrght deep pink to \\hite, 5.0-8.0mm across;petals sub-orbicular to orbicular.clawed, not spreadingwidely. SlamensI0, one oppositeeach petal and calyx lobe, or rarelyup to three antesepalousstamens missing; antepetalous filaments broad, flat, tapering slightly irom base.adnate to the clawsof the petalsand just equallingthe calyxlobes, 1.2- 1.7 mm long, 0.6 0.8 mm wide, shortly united to tlle antesepalousfilamcnts; antesepalous lilaments flattenedbut not much broaderthan thick,c. 0.8-1.1mm lorlg;anthers 0 3-0.4mm long; connectivegland small, globular to shortly cylindrical. Ovary 3locular, 2,/:length of hypanthium, lower r/r fused to hypanthium. Style lairly stout, reaching to betwecn antcsepalousand antepetalousanthers: stiSma capitate. Placentasoval, c. 0.4 mm long and 0.3 mm wide. Ovules 4-5 per loculus. Llndehistetlfruit: hypanthium hemispherical; capsuleequaling calyx lobes. Dehisced.fruit:hypanthium almost flat (disc shaped)with a shofl cylindrical edge; calyx lobes erect, valves opening very widely, obtuse Seedr reniformto stoutlyreniforn, 1.3-l .4mm long,0.7-0.8 mm broad,ariilate; testa d ark brown, neitherdull nor shiny,minutely coiliculate. r!-m 6rlo filling seed.radicle massive. cotyledons small on a sienderneck, both appressedto the radiclc.

Selectedspetinrens. WES IERN AUSTRALIA: Ncar King GeorgesSound, 1884,Miss Franklyn (M EL 76374):5 milcs (8 km) S ol Narrikup on the Albany road, R MelNille 4397(K, MEL. NSW, PERTH): King GeorgesSound, -/. R. Muir(MFl,76316); Gardincr Riverroad. 34' 2l'S, 119"23'F, G. Perry205 (PERTH)i LedgePoint road, 34" 59'S,I l7' 58'E. D. Whiblev 5 165(AD, PER l'H).

[)istrihution. Endcmic to the areafrom Albany to Bremer Bay, WesternAustra]ia Map l. r

'"csnJ'eteln.)rlloc 3]nunu le^ sr,\0eletsat :€lel|lE 'eIruro|uu-osserc '!unecErpull,{c IaA eruroJruer ?uRxeS rtupnbaEulnqol-ec,{lEc oue^o relrAeJq utnrqlued,{q :sn}JnJg utnlntol red g-E e1n,to lutn}€uuoc urnrqluedKq 'eJelnool-t 'snqrl"ulSi€rue p" aroualur eupd unr:e,t6 e8uol tutu 8 l-9 1 snqlclde 'elelsnAup rurlupe:3 eurlelcdelu" ElueurelIJ:€1"uuoJ rutpntrBuol 11,o:d eut1e1:de1ue '61 '.eql€ "turur€lrI :erorSuolsro 1en erlurnbee lletad euru:e1gulelp urul 8 01-0 8 Ie^ easol 'utl?rp 'urneceJpurl.{J €lloloJ eJruocqo rs"q urut 0 e-g Z :e1t,rerqruntqluldlH run8uol 'IIJElIlos ruru 0l-/ runrpodoqluu :rlrsod solnu€J sntas 9-l sorolC o,^.rnJrranbunrald '-rolocsrp'elel 'e8uo1 'eleloacuel eur8:eur tulu t Z-0 1 ruru7 6-6 y eurur€l:€lncp Ie^ EBuolqo 'elecu!Arp 'ptrsodsrp atsn8ue Ie^ erturled ocrpoLu"IIol snl"r'lsordle^ snsnJJasnlncllnJl ',tou ds'ue8pn:1 ;ruoslrJoruslzul5 Z

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,rz,ry uiripn 'j ! ruto.t'?ti{ 4)4 \uyllia Vot 5. \o. I 986) hpu.i. WesternAustralia, Stirling Range National Park. on StirlingRange Drive lo Red Gun Pass.c.9 krr.i bv roadNW of ChcsterPassRoadturnolf.L'.N.S.,tatkson3329(ho|o.. PER'lH: iso:AD. CANB, K). Pcrennialsubshrub to 60cn acrosssclambling or prostratc,or lowerbranches prostrate othersscrambling or spreaciing.L,cler. neither distant nor crorvded(cxcepl near tips ol shoots),somewhat spreading near lips of branches.divaricatc lower down. occasionally twisted.lamina narrow oblongto oblonglanceolatc or lanceolate,4.0-9.4 mm long, 1.0- 2.J nm wide. usuallv rounded (rathcr than llat). acutc, marginsrccurved, smooth. discololous.older leavesoltcn reddish.oil glandssmall but frequentL)visiblc. /rlowers crect.I 4 (6) pcr shoot.solitar], in lcaf axils;anthopodium 7-19 mm long:peduncle 0.1- 0.2mnr lo ng; bracleoles lanccolate, concave,2.2-3.2 mm longusuaily reddish. Ht'ponthiunt shortlycylindrical with an obconicalbasc,2.5-3.0 mm diameter,grecn or reddish;calyx Lobescrcct, not keeled.triangular to obtuseor scmi-circular,I.4-1.6 mm long. petaline on nrarrgin.cnlirc. Cotolla pink to white.8.0 l0.U nrm diameter,petals suborbicular to orbicular,spreading widcly. Stantans 10, onc opposite each petal and calvx lobe, cqualling or exceedingpetals; [ilaments flartened, broad, tapering Irom basc,fused to rjrlength of antepelalouslilamcnts, antepelalous filaments I.6 l.ll mm long,0.7-0.8 mm broad,adnate at baseto petiils.ilpices emarginate; antesepalous filamentsl.2-1.3 mm long 0.5-0.7mm broad: anlhersc. 0.4 mm long; connectivegland small. c,"-lindrical.Ov4rr' 3-locuiar, equalling stanrinophore. lowcr halIfused to thchypanthium. Stylc stout. Dot t:ipcring under stigma.equalling anthers of antcpctalousstanens; stigma capitate. Placel.ita oval, c. 0.5 nm long,0.z1nrni rvide.Ovules 5 ll per loculusin tro inrerlockingro\\s. Ljndehis(e.lfru ; hypanthiumshortl_v cylindrical, ovary expanded, cqualling calyx lobes; pendent. Dehisted lrull: vaLvesopening r,cry rvidely. Sceds rcniform to stoutlyreniform c. 1.5mm long.ar.illatc; testadark brown.neither dull nor shin1.smooth to minutclycolliculate. llnlbri.r_, similar to R. srhol/cri/olio.

Selettetl.s:1tt,tinreni. WESTERN AUSTITALIA: Stirling Rangc.National Park, ,4.M. ,1.rAb,r'368 I (A D. PER TH, NY. TI); Stirling[Langc Narional Park, Lower norrhern slopcs ol Hostcliers.G. Keigherrlll9 (l,ERIH); StirlingIi ngeNational f,ark. Pillcnorupfire break.bchindBlull Knoll. M.fi. lrudgen1730(PERTH. NSW) cast lron Solomon'sWell, StirlingRangc. 28 Scptember.1902. A. Morrison(PERTH).

Distrihuliotl.I-.ndemic to the StiflingRangc. Western Australia. Map L

LIqbitat. I Ltuitl in mallee hearh $,ith Eutalyptus (incLucling1,. m(rginatd, .larrah ), Allota.suorinu, Xunthorrhoeu, I)r)antlrq, Restionaceac,and Clvpcraccaeon lower hillslopeswith dun colouredsanclv or clayeysandy soils.

Note\. Rin:io nktrrisonii is closcll related to R. longi/olia and R. stholleri/oliu (see discussionunder the latterspecies).

Due to crorvdinglrom lhe iar!:crantepetalous stamens (which converge to form a cone around the st!le) the tops ol the antescpalousstamens are pushedinwards towards the style.AdVentiiious roots ltavebccn observed on prostratebranchcs of one collectionof R. tltoni.\onii (111.E. lrutlgen 17301.

(ons?r|ation.\tatus. Rinzia rnorritonr'l is apparcntlynot uncommon in thc Stirling Rangc NationaLI'ark. lt has beenobserved there in sevcralareas with differentfire histories includingareas that had bccn unburntfor 3-5 and greatcrthan l5 vears.Consequently, thc speciesis probablysecure and nccdsno specialattetltion for its consel\atlon. l|ltntologt'.Thc species was named for AlexandcrM or.rison( 1849 to l9l3) who mademany collcctionsof plantslrom the StirlingRange in the carlyvears of thiscenlur\. r

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,r:r//)/ uJn-p'r.rl3 Lulor||l\ Nu!lsiaVol 5. No. 3 (1986) 4. Rinziaoxycoccoides Tulcz., Bull. Cl. Phys.-Math.Acad. lmp. Sci.Saint-Petersbourg ser.2, l0: 3ll (1852).Type: Drummond,5th coll. no. 120(hoio: KW (n.v.);iso: K (n.v.). MEL, PERTH. NSW). Baerkear.,-t-r,r'r.rcr oldts Benth.,Fl. Austral. 3: 75 (1867).nom. illeg.

Pcrennial subshrub to 60 cm across.diffuse, sprawling oI procul]1bent.l,e'al,es close togetherto distant,spreading to divaricateor slightlyrecurvcd; lamina sub-orbicular, o\,ate or broadto narrowoblong, obtusc.2-8 mm Iong,1.3-3 mm wide,strongly rccurved, thick, markedlydiscolorous; petiole c. 0.5 nrm long. Flowel.!crect, l-5 on a shoot,solitary in axilsoI upper leavcs;anthopodium 8-27 mm long, pedunclcc. 0.1 mm long;bracteoles linearto ellipticto oblanceolatc,concavc, 1.7-3.5 mm long. H.vpanthiumobconical to hemispherical,pitted over oil glands,2 4.2 mm diametcri calyx lobeserect, triangular to scmi-circularto obtuse,1.0-1.5 mm long, rcddishwith an entireor laccratedpetaline margin, not kcclcd. Corollq pale to deep rose pink,9.0-19.0 mm diameter; petals sub- orbicularto obovate,clawed, sprcading. Stqmens 10, one oppositeeach petal and calyx lobe, excecding calyx lobes; filaments flat, broad, very shortly united at the base, antepetalousalmost oblong (tapering slightly),2.2-2.5 mm long, antesepaloustapering lrom base,LE-2.0 mm longi anthersc. 0.5 mm long;connective gland srnall, globular. Oyr,'r,3locular, cqual in lengtirto hypanthium,lower Lr/r-2/rfused to tite hypanthium.Style stout,tapering to stigma,equalling antesepalous anthers; stigma small, capitatc. Placentas oval, ovules8- l2 per loculus,approaching radially arrangcd. Undehi.sted /i.l'/l1r hypanthiu m hcmispherical to shortly cylindrical; r'ap.rr.r1eglobular, exceeding calyx lobes. Secr/s reniform.arillatc; testa brown to dark blown. dull, minutelycolliculate.

WESTERN A(ISTRALIA: MiddleMt Barren,C.A. (iardner29l(PERTH): Thumb Peak Range,,1.S. George 7l4l (AD, PERTH); Mt. Bland,Nov. 1932,H. Steetlnan(PERTHl South cast ridge oi Annie Peak. Eyre Range,.1.,S. Ilestc.'n 12797 (PER'l H); Annie Peak, E1'reRange. A.S. Georga7252 (PERTH); Thumb Peak,Fitzgerald River National Park, R.D. Royce 9254(PERTH).

Distribution. Known only from thc Fitzgerald Rjver National Park on the south coast ()1Wcstcrn Australia. Map l.

Hahitat. Fotnd in protcaceous-myrtaceousheath on hiilslopcswith stonv soils.

N.)leJ.Turczaninov and Bentham apparentlydcscribed this taxon indepcndently,from duplicatcsof the samecollection, both applyingthe samespccific cpithet.

Rinzia oxytotroide.r is the largestflowered species in the genusand, whereasthe flowers ol most speciesare not conspicuous.the llowersof this speciesare quitestriking. It has a largc ovulc number and as a cotlsequencethe ovulesare not arrangedin rows but around the placenta.However, they 2rlcnot ncatly radially an.angedas in sectionsRahingtonia and Harmogiu ol B.re(kea L.

Consaryationslalus. lhe conscrvationstatus of R. oxy,tottoidesmust be consideredas excellentas it occursin a large National Park and, from collectionrecords, is apparently not uncommonthere.

5. Rinzia carnosa(S. Moore) Trudgen. comb. nov.-Baeckaautrnosa S. Moore, J. Linn. Soc.Bot. 45: I75(1920). lrpe; BruceRock, H/..4.,Sroward3l5 (holo: BM n.v.:iso:MEL). Baetkea minurifolia E. Cheel,.I. and Proc. Roy. Soc. W. Austral. l0:5 (1923). T.t,pe: Westonia,W. A., April 1918,F. M.C. Sc,4c,ck(ho1o: NSW).

Woody subshrubto 1.3 m tall, much branchcdwith slender(often long) branchlets. leave.sappressed, just overlappingor rarely crowded or distant; lamina sub-orbicularto a

'ruErp 'Eqls 'ululp '!unrJP.lptlllAJ ulur II-S l- ElloroJ ultuoJqo lsBq ultu 9 Z-0 2 JJll\JJq 'Lun8uol unrqluedr(g ruut E g-g Z unlpodoqtuu :rlsod lunJolnluPl scclde snslen t-J 'ruEIlos 'DlEI 'P8uol serol{ etuulo snqtlueuluold snqtpu"lS urnpjclul ruLu t l 9 0 tutu 'Exa^uor 'exe^u t t-l 0 Eurtuel :esnlqo l3r,\el lunpletut sllElxrlpE o r-our'1tl :punlord ;a,l '?llue]ed'eleJIJquII Ertrrel-ruros'pt€^oqorlsniue Ie^ p8uolqo Jtsniuu'"ss.idd€ orPr la'1. 'euelrrpcnbcrlog sll€rloJesuaP:^rsr.'rJLunusL\lt2J\llnLulrJ\n\nllJLuntuepsnlcalJxJlnlJ '^ou ds'ua8Pnrl urqnr Elzulu 9 'J.rnces {le^Ilelej f,q AEu eleq} suollelndod Pue eiuel s1tjo 1:ed urels"e eqt ur 3uu?elJ sseluaeq sEq! raql {lateun1rt4 slred l"Lloll€N ruolj sPlolal ou eiP ejeql se ulPlieaun eJojarJrll st snlels uoll€,\jJsuol s1J olnunall8€ iol JJEId ue{e] Siuls sEq turre.lJ LIJnLuoleq,t\ se.rp ruol.l Jue) puu o8r: slPJ{ 0Z uEtll Jtou .peu Jl3'\\ aseql lo 'SuOrlJelloa l:^otu uerunol 1S€al1P tllolJ u,\\ouI SrDsoutD) y ellq^[J??/r/s uotlDAJaSuo) 'snuJB aql ulqll^\ Peciolf,^JP DtzutY u sroqluuJo tueurqru]]u lllrxepe eql 1"q].]"Jrpur.{eru slq.t, s:trrds zrztlry raqlo 'tlollc3s ur sr?stuetuElu eqt Jo .f,lrllns lelYepe eql ol pcqcslle iuleq ueql JJqlEl ]eq] uI s" pexUrsJoporE srotlluEeqt puc (slueuElll elel3l .^llq qclqA\Jo selJ.d\ rql) I,J))arg +o '-n/J:\ruolJnguorlces ur se pu. :ql o1 rrdnl (rq]'srrrrds raqlo Jql ur se pauell"u el" slustllelrl Jrlt 3Iq/[ snueS rql Jo IBJId,{1lou erP.L)sotl)D) DtzLtlv .to suoulP}s Jqf ttlorv 'tDltqDH 'spuus{ur?ol rrtrupr8 ro ueol uo 3uruo.r8.qnl.s, t?)r)Z.to slalclql ul puno,l

'Z dDl,\'€rl"Isnv uJetsey'{'urlurng pue uueqJell.) o} lsJ\\ PJrEJtltoo?ln) Jqt uoJl punol uotlnqttlst(I '(H r.d1.I '/i\SN) .l'Dgl)Zl9 7.)..)./,.,S') ,U .-t'Bl6l reqot.O'pruotsalN:(HIUEd Zg Y h\sdltllqd 'rr '8881'.{elleA 3 /.1r prt?tloo] Jo lN (ul)l t 9) srlttu l'.(aEw], tlDllat! ? u.ploD :(HlU.l d '-ljhl 'gNVJ) 'eIpr"SlooJ 'sIcoU 9t0B aEnat S L. to INS errotJrA ueenO :(H-LdAd ') 'OH 'gNVJ) 'urqqncuag 'CV) g, LZraupn, I,'J Jo N (uII 6 Zt) s.l[u 8 :(H-LUEd 'arp:e31oo;1 'trII 'Zt6t gggl .tauuo(J ,1 g lo ,lAS,^\ Ee Pr:(HId3(t) iallDg J::l roqolco 'IJo)i 'Surptg e.n.!g 1o EN urS.pupI:VI'IVUISnV NUAlS3,4A srt)u11))t1sPat))ldS 'u^{orq )rEp'el"lnu"l8 ^l.ur.}puE,{lesolc elso1 -,i.in.// :3uol ruur q I-t l J'olellu" lou'rurojluol JParS ilepr,\\ {:c,t 3utuado lou elnsdeJ 'sno..euutli 'lEJllpuII{J pDs'lqae s3qoix,{leJ 3ur1lenbc }snf alnsdeJ ,{11:oq.^ulnrqtuudr(q 'JeAo :tnt.[ p,t.tst4,tpu2 smo: 3ur1co1:elutoi!\] ur :^nlnoollod 9 t selnAo:3uol ultu t 0 J 'tuuedcl 'lnols Eluooeld .]etrdEc Ilacrecs eur8rls:srcqluD snolPledalue 3ut11unbc eli(lS 'runtqtued,{q unrqtuedi(tl 01 pesnj prrq]:c,no1 o1 q13ue1ut lunbe lsorxlP'iPlnJol-t ir,1o 'rElnqolF 'lleus puelS e,\rlJeuuof,:3uol tutu ? 0-t 0 p.xt]r:^JopslJqlu" :epl,{\uIuI E 0-t 0 '3uo1 ruu 10 9 0 snoledesrluu'Jprr\\tutu 10-9 0'3uol uttu t Z t l slPl.dJo mPIJol e1€up" 'seqol lou x.{1ecSurpeerxe IIp.)r€tu snoluladelu" r.qlo qcEe Luor.}luetslp leq^\:ltuos 'sple,$uI 'xede 'poualleU ot pepA\orJ'espq ]p pottun r(llJoqs pe^rn:r sp:e,no19ut:rdr1 sluauElt1:eqo1 xi(1uc qrE3 etrsoddoo,\\l ro .uo pu" Ieled rlie. ilr:^oddooLlo'ZI ,\lelll io 'OI \^u)tllDtsiurpEords'Suolqo lsoLUIu ()1 lElllclqlo-qns slut.d:ssolJE ul tu ! 1-q9'lulLI isor ot alrqi\Dlloto)'.J\ue 'Poleellou'o8pa lullElodellq,\,\\olj?u P qll.{\qslppeJ'8uol 'lEJIlpull^. ruul 0 :eln8ucr:1o1.IelncrJ Irua:^'tJrJcsoqol x{1eo :taleruetp uttu 8 Z-ZZ 0 l-g 'tulolIquIJ ,{1i:oqsot lsJrJeqdsruleqtunrLItuEd,{H Suol ruul Z I-l ,tldrop o1 ,{l,1\ollEqs '3]ploeJupl s.lortr"rq :3uol ruru J g-1 6 rlcunpad :fluol ulrtl E z-9 0 (rlnrpodoqluE:sllxP 'tJara 'stelqtruErq 'JtnulLu rrr ,{rettlos .}o sd]} JPeuio }u pJlo}snlc,-l \1a.Aol! 3loll3d Jr;e1 'ept,tr '3uol ',\\olJq :spuelS le^o strd o] .np q8no: ulru 0 l-9 0 tlltll q Z g 0 XJ,\Uoc,{ldoep llo 'esnlqo .{le^ ot (rplnJirc-rLuJs)xe^uoJ ,ildccp puu 3^oq? e^eJuo:ti(1,to11cqs pur: lr1dr11e ,r:,ry ui;pr.rt A rutoil'rn

I NuytsiaVol. 5, No 3 ( 1986)

a Rinzia communis L Rinzia allinis f Rinzia carnosa

o 75 t50

!o Coolgardie

o SouthernCross I I t o Merredin

I

32.S

Norseman a

I

Lake Grace a .a a ' L a Ravensthorpe1O 'aoo;

o_a ungerup a

o

35"S a Albany 118.E tzt"E

Map 2. Distribution of Rinzia communt, R. alfinis and R. (arnosa. uorlerJeAsrq] 1o ccuccryuSr.^eqt sscssEo] pcpaf,ueje suorlJelloJJeqlrnJ 'oJu€readd€ :srr^\ollrql jo orrsrqt puc sturu€lri oql Jo uorsnjJo eoiSepoqt ur ,{lrlpJol ed,,(teql tp asorl]ruoJJ .re11rp eipJeBIooJ puE ss^o:Juiiqtnos uoJJ su.rurJedseql s.loN

'pues,(er8 Je^o puesud\orq lurd pue ,{eJJ,{puesre^o pu"s,rolleqs 'pues rie{elJu.{o:q e8ue:orpnlJur slros :^urpldSurtplnpun fltueF uo qn]JsDLtuDnsD)ollV puP sJallurulueiroue qlr,r qleeq uedo ,$ol 'eelietUqnrqs uado ,{reAstlDluaulluo)suD.t/ ?'pu"lpoo,r rlt1ssnftIlotng 3urpnllrr sed,{luortetaSe,l:^nouel luorJ u\\out tDtlqDH 'e der'\,J8ur) e)i"-I Jo ls"a puE sso.r.)uiiqlnoq ':rp.te31oo3:eerr uro.U u^\ouI lerlprtsnVurolsey'A ol Jlruapuj uo4nqt.lts:te

'(HJd '.rpjpBloo.) jd) 9199Iaq,\aN y Jo tse^\qlnos ur{ 'rrNSN) 'peor 0Z :(H-L)ll.l 6989.irg.r1rNy 8ur) ol"-I ot (srurlCuorulEs ro) uErursroN '\.irrpl iqruullng elrur06 Jo lse,\\qlnos rul I :(DSJ) 9ZSZ-891y tr\ Sutuuo) /l ? prE,\ol ssorJurolllnoq Luorl (ruI Z fa) seltLut tl iVl'lVdlsnV \dglsqr\\ tuaLut)adspat)4)S

'3]EInuErd^lJuu pue ,{lesolJ'u,rorqptset:et"lluetou'3uoltulu/JglJ'ruro]ruer.rprasuaestoullntfpa)siqa(l -lBlrJprrrlIl 'JsDq 'snoJcEueqJ'Lrqt Jsr,r\lJrlto lE pJrr3pEorq tunrqlupd,{q:tuilsrs::d sueurrls puE sJqol XIIEJ Lunrpodoqluepa^inJer uo luapued n4 p sl4apun s,\\or3ur)jcolrs]ur 'sn1nro1 'seJlueJ o.\\t ur splLr.Jeldot peq:)r]]e :ed 6-t sJIn^O ,(1p:urpnlrFuolp?pr^rp '1u,r.o r.rlt ruJu 11r,u,{.ru,ro ot pJqlllu ,(1uo srtuJJuld .l{ts rqt o} drt pepunor e eru8rls :sreqtus snol"tedelue Surlyunbe'euirts spJeA\ot,{ltq8rls iuuadet el,{tS tUnrqtued,{qol 'urnrqtued,(q 'r"lncol-t pasn1JIeq ot piiql r.r\ol -lo retoun?rppue q13rre1eql JIeq firio 'pe^rnr'llr?uls puelSe,\rlJ3uuoJ :stueruElU.lo aJE.lJ11s lerxepE ol poqJEtlE'3uolulur E 0 J 'esnlqo sriqlue :stu.uIPlr-lsno[:1edelue lq11o q]3uolfi. ol rir roJ p.sn] sluatuelr.JJo setpe ot '3uo1 'pf,ur etn.resrrrdu uru t l-l Suolqo cu ilpel.reu 01,{llq8rls slualuEIrt snoledoselue 'o,^EJuoJ :sleqtur ol luiurqJellP tsud p?puetx. tou tnq etpur8rerueseord"'3uol urur 0 Z-g l 'i1iqir1s 'peo'tq ,{1,tro11eqs.to 1qfru.r1ss:prs iur.rcdut pJuJleU s}u.ut"lrJ snoleledelue 'slEled :Jli(1saq1 punore ouoJ p 3urlulo.l Surpeorxo'oqolxr{lrc puu 1ut:dqoue alrsoddo 'p.,rell 'relnJrqro-qns .uo'01 srrrrrlS Burpr:a.rds o1 3uolqo slEted:r3leruerp ruu ll '!protlep -g l !1rq,\\Dllato,) rtLltsripJ'(,{ln-uorts tou tnq) poloo)i'pcrllnp'rsntqo o} itnrl? 'Fur 'qloorus ;reletuerp p€.rds ol lr.re s.qol xIIpl :pJ.rlln p drap e illrnsn oJe],rns lutu g Z-0 2 'Js"q 'ru:ojrqru.{c leiruorqo IllLuse qtr,{\ ilt.roqs utllqtuDd,iH Suol rulu 6' l-6 6 'etplo?JrrEI 1erupur1.r,r o1rudrll:) selo?]Jerq:3uol ruLut 0-Z 0 elJunped :8uol luul g g-g.ZtunrpodoqluE 'tJlqrueiq :tJiJJ slrrp prel ur ,{.Ie111os e 1o dr1cqt ,ro1oq,{F:oqs b-l s.Dttol! spuelSIro pesrErrllr,r iileiPr ro qloours'IIuoeJPJJns lerxEqE uo olqrsr,rspue131ro "no1rq xr.^,uor r(1deep ,(:r,ro1 fldrrp .,\oqDJ,\pluoJ,{1.ro11eqs..ro iEIJ'Jpr,!\ ruru I I-9 0'3lrol utruI t / 0'esn]qo '-rslnJrqro Ill?lusira'\ uaqr\\ro'31e^oqo-,l\orrt)u o1 tuolqo-,\\orrEu purLuEl :(:^tooqs Sur,roJB IJrnh uo) prsso.rddu{1c:u: ru:r1sruojj siJ.!8epgy c ]ur8:a,lrp'Furddeli.^o snor:e]l.rpenb ir,1rr7 stuetsur"ur eql ruorJ slrlLlJUr-'rqlI,.urs pa\F:l ilrsurp:no:runu qtr^\ rJntEu-r uJq.\\uJdo pur 3ur1,ne:ds'3unor(urt1,r ]Jrrr'ssoJ:ll ru.00l pue IIet urit01 oi qniqS .(,^A SN 'tf '9161 ti )'gNV.):osr:Hf)tg(t;otoq)99L I u)iptltJ. E N r3qrua^oNt'prlprlsnv urolsoy'ApEol sLunDuoLulES ot 3ur;1o1u1 rq] uo )uEf cllhl 06 Jo tse. ul)i I l .sn.]Tl '?auunrq'rlrluupr; 'etlulo]luaJ reulrlqnsptset :pFuol !u!u I I g I c u'lullueuou euttu.S 'elu,rurxo 'utnprLunt opunlo:drsuq turseqrdo:d runrqlucd,{q:sn1on:-1 snlnlol .red 6-y e1n,ro 'snqrtEurBJpure :runteuuoJurntqtupdriq pp rspq'eiplnJol-ttunuu,\O ?3uol !u!u 0Z-g'l snqrJrdrl'elEl:^nFue urlepe:3 eur1c1:rdeluepluaruplrJ :EtauuoJ urnJol"dasetu€ rtlnroluetuEIU!unurprltlll" li. pE anbsn ElLreruEpl:elorAuol srlulod '()1 eururel5

rr /,y uri)f I l,ulLirt.n ,1t0 \uvrsraVot 5. \(, I q86l

Cons.ryation!tatu.s. Rinzia ruhra is known lrom only a lcw collections,and nosl of these are ir-oma smallarea near the 90 Milc Tank on the Lake King to SalmonGums road. Thearea hom therenorth andcast to SouthernCross and Coolgardicis not \lell collected and,as arcas o1 suitable hirbitat occLrr. may wcll containaclditional populations oI RinziQ rabra.Horvever.untilsuchpopuL.ltionsarelountl theconscrvzrtion status (}1 R. r./rrdnusl be r:onsidcreilunsltisl actory.

Ettnrologr'.l-hespecilic cpithct alludcs to lhe dull reddishcolour o1 the hypanthiumand cal_vxlobes. Whilc some other speciesoccasionalll havc these parts reddish,it is pronounccdand apparently-collstilnt in Rinziarubra.

7. Rinzia dimorphandra(F. M uell.ex []enth.)Trudgen. comb. no\'.-/tde('/ie.r dirttorphandra F. Muell. sx Bcnth. Fl. Austral.-.1: 721 (1867). frpc: Sandl pLacesnear Cape le Grand, Marxell (holo:MEL: iso:VEI-).

A Iouncledsubshrub to 25 cm lall and 30 cm across.fairll open:branchcs iong, very slender. 1-c4\'r,.\ appressed to \er\ slightlyspreading. not cluitct(t iust overlappingor rarely up to hall of iengthovc|lapping; lamina narrow-clliptic to elliplicor rarel-Yobovatc, acute to obtuse.1.5-3.9 mm long.0.50.8 rnm wide. deeply cotivex below. shallowlli to moderately concavcabovc. smooth. oil glandssmall; petiole 0.2-0.4 mm long. F/on,ersl-8 clustctcd a1or.jusl belou tllc tips oI the branchlets.rare]v spread along branchlcts.in pairson a conrron pcdunclcor lesscommonl! solitary. erect: peduncles 0.2-0.3 mm long.terminated by a pairol bracteoles;anthopodia L0 2.,1mm long,cach subtended by a pairol bractcoles; bfrcteoleson pedunclesand anlhopodiacllipticlanceolale and shallo\\'lyconcave to clmbilornr. 0.E2.2 mm long. HtpanthiuDi shortly cylindricrl to hemisphericalor obconical.taper ing into the anthopod iLlm. 2.0-2.1 mm diameler;caly\ Iobeserecl or slightly sfreailing,triangular or sub-tfiangLrlarto semielliptic I-1.5 mm 1ong,entire. Corr.,//a pittk or rarclypale pink to rvhitc.6.0-9.0 mm diamctcr';pctals sub orbicularto !ery broadly elliptic,or outlincquite angular, sprcading. Stantans 10. one oppositc cach petal and calyx lobc:antcpctaioLrs [ilaments ob]on!1. of sLighllr-dilated at apcx, L2-1.7mn long,0.6-0.7 ( L0) nn widc.shortl) or to lo\\'-erone quartcr fused to [ilamentsoi antesepalousstamens, equalli !a or slightl_"-erceetling scpal lobes.apex emarginttciantesepiilous lilaments Ilatleneci.tapcring.0.7 I.3 nn l,lng 0.3-0.5rnm wide. apex acuteto obtuscand not equallingtop of anthers:anthers c. 0.3 0.,1mm long; conncctivcgland small,narrow- obovoid. (J|arI thrcc locuLar.nol or just equallingstaruinophorc. lower l1:fuscd to h\panthium.Stylc taperingor lusi1brm.reaching between anthcrs ol antepetalousand antesepalousstamens; sligma not or scarcelvcapitlltc. Placentas cilcular, sLightl! raised. c. 0.4 nrm diamctcr..Orrrles -3,1(5) per locuLus.Und(hi.-(kl ./ruil: hlpanthium shortiy cr,lindlical.base tr uncatc, capsule expanded but not equallingcalyx lobes. Dalited Jruit. h),panthiumhcmisphericalorshallon,er."Secl.rreniforml.l-l.3mmlong,arillate.arillarge; testa.smooth. mid brown.shinr. Elrbr|o; cotylcdonssmall on a slendefneck coilcd on the end ol thc massi\eriidicle.

S<,lcttetl.t1tt,tintcn.i. WfrS'l FRN AIJSTRALIA: 3 km southeast o1 Hil149. Cape Le Grand N"ationalPrrk. R..1.Cranlield r375 (CANB, K. MEI-, NSW, I'ER l H): l2 km sottthol Howick Hill. ca. 75 kn ENE o[ Iisperance. N.N. Donncr 2749(AD,l-. OSH. PERTH. Z): ll.5 km WSW of Howick Hill. Hi. Eithler 19861(AD. CANB. K, PEIITH): 1.8miles (2.9km) alo g tlrcroad lo WiLlenoomHills lrom the Esperanceto CapeArid toad. M.I:. 'l'r'utlgen 1175(BItl. M UL. NSW. PERTH).

Distribution.I-ndemic to WeslernAustralia. found front Esperanceeast to Howick Hill. Map 3. 'luelllos turu runrpodoqlue :IteBerSSpurnlolnlu?J seotdesns::,1 tZ-Z Ie,^.Il"ulq 0 t-t l '"ueld serolj ptel tuur 6 0-/ 0'e8uol trltrlC t-0 2 EuM"l :Psntqolr,\ elnc" areJle^ e^Ecuoc 'eJItdIIIe 'El"loeJu"lqo €rdns'uxe^uoJ epunlo:d er]ul ulnprelul l.^ ESuolqo slsn8ue 'et?Juqrur 'es^se:ddsqns r,lsnunu snld la,Lesse:dde erloC snlccle IoA snl"punlol xelnlJ 'lou ds'ue8pn:1 sIulJJEslzulu 8 'iJotJElsrll,rs,{lqeqord srsnte}s uorte,^resuoJ s}I oi^'()iJEd llTuorteN pu"rC e] odeS) 'sntols o^leseruortp^rsslloJ auo lseal ]p ur punol sr uPuDqdloLutPDtzutll uollD\ta'^uo)

]ooqs a,ttteta8e,re s€ ql,llorBsJnurtuoJ,{llEuoIS"sco xedl olcunpadaq1 'pnq 'uotltsod ur llits sr JJqto eql ellq,r srsf,qtue]€ eq ,{erueuo B e lse8etstuara.}Jlp le ert 'alJunp3d e^rtelJl eluesoql uI {ltuJredd" qinoqll" uoLuLuoc€ uo s:a^\olJ3o :red eqa 'dds sadIlDnA {q peleuru-ropuor]eto8o^ uI lo^pl8 clrlelEi qlIA\spues lo s^rup()lur s,roi8 stut/lo otzuty lnq a"ecesloJd,{q psleuItUopuollela8e^ rrt spuesuo smo:3 otpur:ttldlctuttprrzr/ry sEIIoroJalrq,n o1 >iurd.led ,{lelel lluo ro lurd sleqDipuvqdlotulp ,?z?./ry s€lloroJ lurd eled ro itrq,\\ seq slul/:D rrz&ru clrq^(\ slo'$ou JeAJ"l s"q KlensnstutJJootzury'sdeq:c,loeztsre,noggoa8ue:aqta1lq,$',{3,$srueseqtuI:^cleIJc^os3zis 're3:e1 leel Jo i8unr eqt q8noql ,{11e.reuaBeJe slulJb DtzulAJo osoqttnq se,teelpesse:dde a^eq sarJedsqtog ]c?ri .rour sr pue uJ S1 J \JqJeJ.Lrtu(Jr ntzulll allq,\\ ll"1 ulc 9Z ot qnrqs paqruerq ,{leur}u sI otpuoqtltotutp arzrry ecue:e;e:dtetrqeq pu" ,(Solorldrolu 'lrq"qJo ssJueJeJJrpluPts^uol rrP Jleql pue'lcunlstpe:e,LJq1'le^e,\oH lsed aql ul pesnluor u)aq seq tr qJlq,r qtr,\\s:lutf./n DEulll ot petPlcr,{lesolc sI otpuDqdtoLrttpotzutv saloN spuEsd\olle,{ pue ,{eJB 'us'ot)ads 'elrq^\ uo 'DpunqtnLl Dli^:rnNpue ds otltaqtuoT DtgluDgJo sqnrqsJell"l qti'\\ qtssq ro spuElqniqs,\\ol'qnrJs Ds^oDadsDIsIuDg ut stttot?tt.tpuo4dtoutlP Dlzull! 1D11qDH

et(\a! U p\teD.tpuDrytt.!)lutl A Dtqnt otzutHJo uoufqulsrC I dEt\

t..02

oaoto

srrssrt,r:/ru v tipntlrltot'lP n!.u! A O .rqnt.rzura .

/r/:rry u.ipnrl E mto)trt,J 432 \ultsia Vot 5. No. .t ( t986) longum.Hypanthium obconicum,2.l 3.0 mm diam. Corollaalba, interdum rosea, g.5 10.3mm diam.Stamina i0, petalanec superantia, versus biisim brevitcr connata: lllamenta antepetalinaoblonga, 1.5-1.9 mn longa,versus apicem gradatim angustala vel dilatata, apice emarginato.Ovarium 3-loculare,parte inferiorc ad hypanthium connatum; ovula 3-7 per loculum. Fructus: hypanthium perbrevitercylindraccum, laeve, basi obconicum. Seminarcnitormia, arillata, 1.4-1.5 nm longo;testa laevis, rufescens.

Tlpus: 14 knr W of Newdegare.Wcstern Australia. l2 August 196E,p.G. Il/ilson 1032 (holo:PER l'H; iso:CANB, K, NSW).

A roundcdor erectshrub ro 70 cm tall. open(especially when larger),branches long slendcr. Zearc.t appressedto very slightly spreading, r/s-/2 overlapping or rarely just overlapping;lanina narrowlyoblong or rarelyelliptic to oblanceolate,2.0-4.3 mIL long, 0.7-0.9mm widc. concaveor rarely almost flat above,deeply convex below, apex acuG to obtuse,smooth, oil glandsobscure to easilyvisiblc; petiole 0.2-0.5 mm long. Flowers 2-24 per branchlet,in pairs or sometimessolitary in uppcr leaf axils, erect.pcduncles 0.3_ 0.5 (1.0)mm long,each lerminated bv a pair of bracteoles;anthopodia I.3 4.0 mm long eacl'rsubtended by a pair of bracteoles;bractcoles on pedunclesand anthopodialanceolate and concave-convexto cymbiform. 1.5-2.3mm long, thin, occasionallvred tipped, othcrwisepale. Ilypanthiun obconical,2.l-3.0mm diameter;calyx lobes erect to slightly spreading.sub-triangular to semi-circularor semi-ellipticto sltortly oblong and obiuse, 1.0-2.0mm long, thin, occasionallvribbed, maroon (oiien blotchy). C'oro1lawhitc or less commonlypale pink, 8.5-10.3mm diameter;petals suborbicular to very shortlyoblong, spreading.Stamen.s 10, one oppositeeach petal and calyx lobc; filamentsof antepetalous stamensoblong, slightly tapering or dilatedncar apex, L5-l.9 mm long,0.6-0.9 mm wide, shortly fused to antesepalousfilaments at base,exceeding calyx lobes,apex cmarginate; antesepalousf ilaments flattened. tapering from base0.8-1.3 mm long,0.3_0.5 mm wide. apex acuteand not equallingtop of antherslanthers c. 0.3-0.4mm long; conncctivegland small,globular 1o obovoid. Ov4l1,3 locular,not quite equallingstaminophore, lower /r-rh fused to hypanthium. Stylc slender, fusiform, r{rachingto betwccn anthers of antepetalousand antesepalousstamens; stigma barely capitate. Ovulcs 3-7 per loculus. UndehistedJruit: hypanthium very shortly cylindrical with an obconicalbase, rather dry and chartaceous;capsule not quite equallingcalyx lobes;calyx lobesthin, ercct.Dehisce.l ./iult not seen.S'eedr reniform. ari]]ate,1.4-1.5mm long; aril large and extendingto one end of seedltesta smooth shiny,mid brown (a little reddish);e,zDryo with small cotyledons on a long neck wrapped around radicle.

Seletredspetimens. WESTERN AUSTRALIA: Halfway betweenNewdegate and Lake Grace,A. M. Ashbf 189(AD)t 25 km S of Newdegate,B. R. Muslin 3859(NSW. pERTH): I00 miles N of Stirling Range, I E79,Muir (MEL} ca. 25 km W of Lake Gracenear Tarin Rock, D../.j9 llthibk'y 5300(AD, PEITTH).

Disftibution: Endcmic to Western Australia, found from the Lake King area west to Kukerin.Map 2.

H.abitqt.FolJtd in avariety of vegetationtypes including open heathwith Eucalj.ptusspp. high shrubland(ie. two strata) and low dcnsesandplain scrub with Eucalyptus'titagon'a, on a range o1 soils including yellow sand, loam with lateritic pebbles,sand with lateritic pebblesover lateritc and sandy clay ovcr laterite.

Con.\ervdtion.\tatuJ. None of the collectionsof this speciesis from a conserva'on reservej and as it is known from only I I collcctions and ocurs in an area heavily cleared for agnculture,its conservationstatus must be consideredunsatisfactory. .{lPd

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a Rinzid crussilolid I RinziaJumana

o--=J=.-_go

Km T T a Moora I

I I

32"S,

a Brookton

l Narrogin

a Kojonup

1A ^a

a Mount Barker

Map 4. Distribution ol Rinzia rrassilolia and R. Jnnana. 'se)lP-I '7 de4 rrlerlsny urf,tsf,rlA uotsltqof eql ol qN puE Suruucte) ot N'f,dloqlsuJ^l',U 01 q .8uEd Eurllrls lql ruolj pu|.ol uotlnqulsl(I

'(HJ.Ng.t) ':l,tz 88ZVl)H.@aC .S V "81I S,lt o!['ecual 1oo-r.1lrtlqr:X duolc 'H-Ld3.l) 'edioqlsue,\"u irrrqe,{p lir qS tu)t zz }noq€ :(flNVJ zoas )Enat { L' Jo N 'uoqs 'dnSunurDrref -lahl thl :(Hldf d) ost i r-g:1lrN v Jo ]sry'& sallru z :(H]Nad ) 'CV)00f '3rrr;c1e'1.1ofS'uepp"l|rll:(H-LUld)rI00| ';rdtC I ua]pnL lt A -St'l uaJlnlJ ,92.6ll'S,6E "f! Iro)l seou {\oloq u plEreSTlrl:(H-l.xEd'/!\sN) 9tr 3 14 'pf; '1d]/i ou srelsol uo dnieFuolo N tu) L 6 :(HIU3d ) LOLuaSpruJ ? ft' edroqlsu3red 'A\sN ') 'SNVJ 'lNB 'CV) ',t-no lo .t rrll t 0t :(Hlu3.l tgtl uaBl'nL1 t n uourlEs ot puor uo eruJo lsod 8ur) olE-l lo a (Ltt) 6 82) s3|ut 8l :(Hlu:ld) tdltllrDD 'V 'La6l 'eiued r7r1177rr1/v ) requr:lldrS turtrrls :VI'lVdl-SllV N)l.l-LSA,1\ 7r17r1715' '3l 'lJeu 'peas ppr punole petst,\\l lJpuals t uo lleuls suopol,{loJ'J^lssl?ul el:)lPel '3uol 'lulo]IuJl EullltJ o,t1qu1;):Ll,\\orqIrup elPlnJllloc AI.]nullu els?] :JlElllle ruru I l-9 1 p))slqa(l ,2r.r5 u.1uns tunlqlurlcl,{tlJo JStq'rilspl'\\,{le^ Suluedo'osn}qo se^lP,\:ltntl 'iu0t..,,..'"clsueturts'saqol \,(leo SutpJJcxa^llq8lls elnsde.'lllcllpull,tJr(llloqs tunlqlu€dIq 'ulJo.lluc.l 'lcl.llllloJ 'snlnJol : ntf pDs^ryapun Fuol tuul 9 0-9 0 led Z soln^O 3lluf,J 'eele 'lEr\o u,r\op lrls llJql lo tlJlltu ,loJ slxe o1 peq.PllE sPluac"ld sualuEls snolcdeselue 'dl1 eru8lls eql qlJ\\ puc sno[:]rdcluu Jo srJLlluP u:rJ,\\13qol Sutqcle: popunol E st el€lnqns'lnots el\]S tUnrqluD,{qo} p.sn.}rir led\ol unlqtucdlq Sulllu lou lnq eloqdoulruels '.rr:1nqo13'lEInJol-! i-rrpJ3Jxe ,t1tt1ir1s:o SurJlenbo :L/,,1O pro\uqo ..'l rELrlqolF 'sueruPls puEli ?^rllJuuoi :atnJu snol?lodilu" Jo sluoulelUol pJsn] tllFuJl Jluq \\ l\otullr lo 'IJILIl snolDdasolu" ,{11,roqs3u()1 u[u 9 l-6 0 tll]tl] l3prd\ ^l.Jluls lnq pJualleu s]u3tuEIIJ '3uo1 'e^PJuoJ sepls lo :sr.qtuE Jo do1 turll?nbr pull Jlrlul8lPruJ xede !utu 0 Z-t l 'iuolqo 'saqol r(1to::eu x,{lDf,lo ql8u.l eJl^\l o} Surpeelx.,{ilqAIISslu3tuull] snolul3delue o1 :.qol xIleJ puc Flad qcuo eltsoddo euo'01 .!./ri./rrls iutpee:d.^'lDlnclqio flle1n8a:rt Suolqo or sl"led :releluerp urlu 0 6-119 lurd .lud oI ellq,l\,//o.'/').) ?3p' curlElad ll?proqtuoqt 'pl{rllep 'l Mor.rDup rltr,r pul?peleel'(lo\\ol.l Juo uo u.^.) esnlqo o1 elnlt r)lJ \Jqol \(Lrll :qlooruscJr).llns'i. ].tue!p urLr]91 117'1r:cr.Ieqdsruloq,{lpPoiqolosEqlrlllPLl}l\\|t:rllpurlir -3uol '!ulollqul,ti '3lEAoqo i1.'to,.1suuntltuotl.ill urru t l-9 0 o] 3lucuoJ r(l,lrolleqs ,{i,no..r:ruo1 rrlclrlle :^rloctc?rq :6uol luur [ 0-i 0 srl]unped :3uol tutu 11tr-E 1 erpodoqtuu 'sJ^ual (1tJr:p :tooqs D uo q leddn lo sllYnut ,{.tu1t1os'lJ!l?s/r11olt ,'rlolJLl\aALltrr '{rar I '0pI{\ ot xe,\uo. ,{1deep e,loqe pe^ool3 lo e,rcruoc .{1'to11eqsillluotserco .lo }llll ruul 6'0 .eleLoJJuElqo -E Luut ln Jtl,-loeturl{lalu t to Ftrolqo^\ojjPu ol lEeutl 0.iuol f s-g 0.esntqo 'sllolll?Jllpcnll eurruEl :p.ss3rdde IIJ.!El lo (u!.ls ulo.lJsrr:8ep 69 r) Surpee.rd.^'Sulddrlll.^o sa,rr.r7 ,r1i,ie:1spus turplrlids leql".! o1 pePunoi ssoloPulJ 0t pue II€] tuJ 0[ ol qnlqsqnS

(,^[SN .tSr',J .)'8NVJ :osr:H.t_]Jqd :rntlr7 :oloq) 00ll u)nptu.t. ! J4 9L6L ,{lnf 6Z'V /\\'8ur) e)n.l Jo-:lS'urppr11 rrtnoyl 'r.r\nl r:,nln\r.luJ Jln rrtlrrl..ll :p,ltt,'lt! ul l l-9 | 'DlEIIuc'Brt!rolrL!aJeLlttues llueledns lJllAolrunqol-elrill?J ollllAo'ole,\DJxo epunJold Iseq 'urn:rcutputl,tr.talt,lelqtuntqluPdiq :sntcnll Lunlniol :ed 7l1n,lo:Lun1:Llu')'rLUnlqlund(q 'r:u1nco1-g 't3uol 'tBuolqo pD Jror.r.Jur a1:rd LUnlle^c) 811?ul8lITolJarldP tuul 0'z-t l elsnBup uurllr]adJtue eluaLu€llJ:rllluuoJ LUlrlull?dJ\rltll' uLntolLIaLuElilSIulpnltlls p! rllrl:d nulurPls ruttp tutu ?nb:^nl3,\ PlPutluoo l.llAJlq elu.tll}'lLl:PIluPlJJn\J)Ll 0l 0 6 8 E 'pql" rlr:L tun"Plpull\J Jel uJsoi aprllud u11o.to3tu?lp rutu 97 q 7'uncr:er;qdslLuoq lJ\ r31r^.rq urnrqluEd,(p runSuol uILU 0 t 9 l ulnlpodoqluu :tleFa:g8u LunlolnturJ saJlde snsreAE-I'ruulllossl.loljlllPltuLu60-90'DtUOIulult980eultul?l :esnlqo'exe,\uocou€ld 'elPloif,uEIqo nJuulqo rlsnlluE apunlorcl I..\ DItaiellLLIJs le^ Elcloe.url uInpletul la\ 'PlPJl.lqLul'et:tltrpcnb euplurl rssiJddp runpJcturI.^ rrrtualed etlog sllsn.lJJsnlnJlllll-l 'Aou ds'ueFpnll A t l slunturuoJElzutU 0l

!tt ,r-11r.\/uiitpnrL -t rutortD t\ a

N!\r\a V,,1.5 \, ltlt,Xn) Ildhitat. Rinzio (anlmLrnis is known from a wide rangc of habitats. including open ('!Eut ult'pt tt.;) \roodland, low miillee heath, mallec scrub with Melaleut.' uit,inita, Eutalt'ptusretlunta low mzilleeheath and tall shrubland.on soilsincluding orange brown sand,light brownsand, sandy loiim. sandy gravel (latc.iric) and latente. (onservation status. Rinzia (ommuni.\ is the most widesprcadand common spcciesin thc gcnus and r.vouldappcar to have a reasonablysecurc conservation status as it has been recordcdfron Frank Hann,Fitzgerald River and Stirling Range National Irarks. Howevcr, rt lsnot corrmonin eitheroI the lattertwo Nationalparks and extensive clearing of native vegctzrtionis occurring,or is proposed,within its rangc.

Ilt.rntolog.r.fhe spccilic namc refers to the widespreadand common occurrcnceof the spccles.

i l. Rinziasessilis l ruclgen,sp. nov. Fruticulushumilis, eflusus. vcl interdumrotundatus. Folia quadrifaria,imbricata vel interdumdistantia appressaque, oblonga, eliiptica, vcl suborbicularia,raro lanceolata. supraconcava. infra convexa. obtusa \, el interdum acuta,sessilia; lamina 0.6-3.2 mm longa, 0.4-0.8mm lala. glabra, vei margine gossypina.Flores solitarii, l-5 versusapices ramulorumaggregati: anthopodium 0.8 2.2mm longum.Hypanthium hemisphericum, L7- 2.6mm diam.Corolla prllide rosea vcl fere alba, 5 9 mm diam.Slamina 10, petalis longioraj filarrentavcrsus basim connata; filamenta antepetalina oblonga, versus apicem gradatim angustatavel lcviter dilatata,apicibus tl.tlncatis en]arginatis vel obtusis.Ovarium 3- loculare.parte inferiore ad hyprnthium connatum: ovula 2 pcr loculum_Fructus: h!panthiumhemisphaericum, ovario calyce-lobum superanti. Semina reniformia, arillata, c.1.5 mm longaitcsta nlinute colliculata. brunnea.

T.rpus:10 km North of Lakc Cronin,c. 8.1km E of Hydcn.W.4., l3 Septenber1981. K. Newbey8798; (holo: PLRTH; iso:CANB. K. MEl.. NSW).

Low, spreading.or sometimesroundcd, subshrub to 50 cm tali and 60 cm across: branchietsslender to thick, *'oody. ltavcs scssile.quadrLfarious, overiapping. or rarcly distantand appresscd;lamina oblong to ellipticor suborbicular.rarely lanceolate, obtuse or rarelyacute.0.6 3.2 mm long,0.40.8 mm wjde.!herllowl)r concave above and convex 1odecplv convex (semi-circular) below, glabrous or with lew to manywhite, short to long andtangled, irairs on margins.l:!owers l 5 neartipsofbranchlcts,solitaryinaxilsofupper leavcs,erect; ilnthopodium0.8-2.2 rnm long; peduncle0.10.4 mm long; bracteoles cvmbilorm0.8- L7 mm long,glabrous or marginsproduced into longtangled white hairs. Hvpurtthiunthemisphcrical, 1.7-2.6 mm diameter;calvx lobes slightly spreadingto spleading.triangular to scmi-circuiarto obtuse,0.7 1.2mm long,glabrous or the margins producedinto rangledwhite hairs. Corolla Iight pink to almostwhite,5.0-9.0 mm across: pelalssuborbicular to broad-ellipticor almostsquare. cLawcd, spreading. Sramcnr 10, one opposileeach pctal and calyx iobe. Filantents of antepetalousstamens oblong or tapering, rarell'dilated near apex! I.2- L 7 mm longby 0.6-0.9mm wide,apices emarginatc, rruncate or obluse.not cxceedinqanthcrs; antesepalous filantents oblong to narrow oblong or lapering,0.9 L2 mm long by 0.4-0.5mm wide,fuscd to antepetalousfilamcnts ior hall to threc quarters of theirlcngth, apices acutc to obtuse;ant hers c. 0.25-0.3 mm long.opening rn parallelslits; connective gland globular to shortlycylindrical. Ovar,- 3-locular, top level with staminophore,lower quarterto hail fusedto hypanthium.Style stout. tapering, rcachingto hali wav betweenantepetalous and antisepalousanthers, or rarelyexcecding antepetalousanthcrs; stigma barely capitate to capitate.placentas oval. Ovules2 per loculus, collatcral. Undehisced/rrll. hypanthium hemispherical,capsule expanded, cxceedingcalyx lobes. Dehisted./ruir;hypanthium becomingIlat; valvesopcning widely. '1ooqs .3rrol :8uol lutu g 9-7 1 unrpodoqtup :slrx€J€rl ur ,{relrlos e uo (Zl) g'l s.taao! urrx l0 c alortod:qtoous'xe,\uoi-ou€ld,{1dc:po1,{l^\oll"qs'epr,r ruuI 9l E 0,3uol ruru 'Suolqo-,\\oJJeu ,pe,\lncej 9 6-02'csntqo'Jr]drlle-,rolteu lo Suolqo,{lleuors"aco ol leourl 'pesse:dde ,{l}qBIIS.ro lq3rerls punu€l :pep,r\olool tuuls^lp ,{1a.te:'a1err:e,lrp 01 iurpurlds 'qnlqsqns .{1tq8r1sranzaT ssolJeutc 0t puu Illlt urJ 0Z 01 tceJeol SurpueJdslo c}eltsold '(HluEd "l3l^l :olcelosr:( ^ u) ) :(pa]relasereq) olcJl)'Zal ou ullor qt9 puouunt] :adlJ '(1981) '9ru:g '.Ig nL:t lerrsnV I.{ Duoulats^[lDldo l)arg- (Hfuad l,.\jiosr ( ^ u) ,lA) 'ZZ ,Z :oloq)EIIeISnV tsr,\\ qlnoS l raqulnu uortiellocrltg puotu un)e :)d,t (Z9gl) ltf :01 'rrs 'lJ ! .ZI FrnoqsretodlulES rrS dtul pery qlel^l- sr{qd IInB zolnI ErloJrssBDErzuru

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I OrnBrC.etclnJrllol '3uo1 f1e1nultu',{urqs'u,{\orqp[u e]sJt:f,tpllue uur S J J'urJoJruol .?ra5"tuepued trnl.{ Ltt ,r-?r/ry u.3pn r-L g rutorlEt^J \Lrvtsr.tVol. 5. \o I ( l9lt6) peduncle0.2-0.4 mm long;bracteoles narrow-elliptic to oblonglalmost 11at to clrnbilorm 1.6-3.I1mm long.green or reddish.Hvpanthlum hcmisphcricrl to shortlycylindrical, 2.2 3.5 mm diarneter:callx lobcscrcct. triangular to obtLLse.1.3 2.1 mm long (longerthan hypanthium).gfeen or reddish-brownwlth pctaiinc margin. Corolla whitc.lesscommonly pink,7.5-11.0mm diamctcr;pctals suborbicular to bfoadlv ovate wilh a stout claw. Stamens10, or farely 12,in a conearound thc style,onc oppositecach pctal and calyx lobe,crowded, not to slightlyexscrtcd, shortly fused at the base:anlepetalous fi]amenls oblong,cmarginate, 1.2-2.2 mm long,0.8-1.2mm wide:antescpalous lilaments tapcring, 0.9 1.3 mn long,0.5-0.6mm wide: anthcrs0.3-0.5 mm long, cqualiingor cxceeding lilaments, opening in parallcl slits; connectivegland small, globular. arvrl_t.3 . or rarely 4-locular,globular to shortlycylindrical, equalling staminophore, lowcl quartcf(or less) lusedto hvpanthium;st)'lc stout, cqualling anthers; sligma capitate. I'lacentas oval: ovules 2 per loculus.collat"eral. Lindehist<,tl /ruit: hypanthiumsauccr-shaped, ovarv cxpandedand exceedingcalyx lobcs.DehisteclJiuil: hypanthium almostflat: valvesobtuse. opening very widely.Set rlr reniform,1.5-l.9 mn long,arillate; testa mid to dark brown,dul]. vcrrucosc. Embrlo filiing seecl;radical massive,cotylcdons on a slcndcrncck and appresscdto radicle.

Sele

Distibuion. Endcmicto WesternALLstralia and tound only lrom Watheroosouth to the Darling Rangenear Perth and eastwardsto ncarMeckering. Map 4 llabitat. Kno\,\n from heaths.Allotasuarina thickets and Watdoo (Eutaltptus ' ,(tndo()) woodland.UstLally lrom areaswith lateriticsoils (sand or clay)but alsolrom onelocation with sandysoil ovcr chert.

Norer. The habit oi Rirzia (russildlia is quitc variablc. ranging in lofln from totally prostratelo a smallshrub.

Thc vcrrucoscsceds are qtLite distinctive in the genus.Although somewhat similar testa surfacesale found in somc spcciesof relatedgenera. these species dilcr in a rangeol' charactersincluding other seed characteristics and thc numberand typeof stamen.

Occasionalplants arc 6-mcrous,thcir tlowers having 6 petals,6 calyxlobcs. l2 stamcns and 4 ovrly loculi.otherwise these seem pertecllv normal.

Conservolionslolus.Rinziacrussi/oli4hasnotbeenrccotdcdlromanyconscr\ati'InIcscr\c and is not il cornmonplant, so its conservationstatus must be considereduncertain at thisstage.-l-here are arcas of StatcForcst within its rangewhere it may occur.

Acknowledgcments Access to thc collections oI the Western Austra]ian Herbarium is gralefully acknowledged,loans oi specimenswere kindly madeby the curiltorsof AD. MEL, BRl, NSW. ClnNI| andCBG. l-atindcscliptions we re very kindl_"'preparcd by Mr PaulWilson. who also rcrd thc manuscriptmaking many constructivecomments. The nrapswere preparedby MissA. Napier.Financial suppoft lrom thc AustralianBiological Resources Study program is gratefully ackno\!ledged. (sap€qJpu€ pr^eC :uo^oc loqqv uol,4\oN)uolllpo puo.es ..uqel l€rru€lo9,,(€/61) I lA'ur"ats a8rnqulEH) :t a€tu?Id.,'Ju?ruqo1 ut ,7zr?y(tt8t) 't 80t .leEuerssrard J t'ran?q.S,roneqrS Z-gtZ'LI peeuur-IetruSoceu onurc IaCs^oN u]nreec€utr'1l uouoC (tt8l) J.f ( uueu-Ito8uE 'Sot-ZS:(Z)e :3rzdra.I) ..uallurpJuozueuduaqJrrrnr€N atc.')'tluerd pue V.retSuEul aeer€u,{t!(868t) C.nzueparN (g/6t'ureetsurpotono)98-l:9ttsrrErnl?Npuptprhluecueuv:EreJrrnr:teqtJospoes(tS6l).d.t\..{otrnht '99l-00/:tz prec 108 Irnossr^ uuv srs{tpu€rrleue8otlqd € oearEu{t{pue setEu,(t! (936t) .C g ,s33ugpu? g y.1luosuqo1 ""',",,, .arnl3n,rsa'uarsa,ouur uro{eruopr^. """"",,0^ "u, u,33ffif?ffi5\TTf#3,fl"T!":13 -D .uleqlueg ( uopuol :e^ood) € Io^ (srsuorl€IsnVeroll!, (/98 t ) soluaraJau

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