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Pteridophyta of Peru

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Pteridophyta of Peru CATÁLOGO ANOTADO E ILUSTRADO DE LA FLORA VASCULAR DE LOS LLANOS DE VENEZUELA Blechnum PTERIDOPHYTA (Pteridofitas, Helechos) Recopiladas por Gerardo Aymard (PORT-UNELLEZ), revisadas por Alan R. Smith (UC), Robbin Moran (NY) y Robert Wingfield (CORO-IUTAG) ASPLENIACEAE Ampliamente distribuida en todo el país (Amazo- nas, Bolívar, Delta Amacuro, Falcón, Mérida, Asplenium formosum Willd. Nueva Esparta, Sucre, Táchira, Yaracuy y Zulia). Helecho terrestre, a veces epífita. Bosques de galería. Apure, Barinas, Monagas y Portuguesa (A.R. Smith et al. 1164, PORT, UC). AZOLLACEAE Ampliamente distribuida en todo el país (Amazo- nas, Aragua, Bolívar, Distrito Federal, Falcón, Azolla caroliniana Willd. [Lám. 1-b] Mérida, Miranda, Sucre, Táchira, Trujillo Yara- Helecho flotante. Lagunas y cuerpos de agua esta- cuy y Zulia). cionales. Apure, Cojedes y Portuguesa (B. Stergios et al. 6835, Asplenium fournieri Kuhn [Lám. 1-a] PORT). Helecho terrestre. Bosques de galería. Ampliamente distribuida en todo el país (Amazo- Portuguesa (F. Ortega y B. Stergios 1286, PORT). nas, Delta Amacuro, Distrito Federal, Falcón, En Venezuela sólo en los Llanos. Mérida, Miranda, Sucre, Yaracuy y Zulia). Asplenium hoffmannii Hieron. Azolla microphylla Kaulf. Helecho terrestre. Bosques de galería. Helecho flotante. Lagunas y cuerpos de agua esta- Portuguesa (G. Aymard 2089, PORT, UC). cionales. Sucre. Apure, Cojedes y Portuguesa (B. Stergios 12698, PORT). Asplenium laetum Sw. Ampliamente distribuida en todo el país (Amazo- Hierba terrestre, a veces epífita. Bosques de galería. nas, Delta Amacuro, Distrito Federal, Falcón, Anzoátegui, Monagas y Portuguesa (F. Ortega y B. Mérida, Miranda, Sucre, Yaracuy y Zulia). Stergios 1467, PORT). Ampliamente distribuida en todo el país (Aragua, Bolívar, Distrito Federal, Lara, Mérida, Sucre y BLECHNACEAE Zulia). Blechnum asplenioides Sw. Asplenium melanopus Sodiro Helecho terrestre. Bosques de galería. Helecho terrestre. Bosques de galería. Barinas y Portuguesa (J.A. Steyermark et al. 127087, Portuguesa (A.R. Smith et al. 1062, PORT, UC). PORT, VEN). Zulia. Andes (Lara, Táchira y Zulia) y Bolívar. Asplenium pumilum Sw. Blechnum gracile Kaulf. Helecho terrestre. Bosques de galería. Helecho terrestre o epífita. Bosques de galería. Portuguesa (B. Stergios y G. Aymard 4449, PORT). Anzoátegui y Portuguesa (B. Stergios y A. Utrera Ampliamente distribuida en todo el país (Aragua, 2469, PORT, UC). Bolívar, Distrito Federal, Falcón, Lara, Mérida, Norte de Venezuela (Carabobo y Yaracuy). Miranda, Sucre y Yaracuy). Blechnum occidentale L. [Lám. 1-c] Asplenium purdieanum Hook. Helecho terrestre. Bosques de galería. Helecho terreste. Bosques caducifolios. Barinas, Cojedes y Portuguesa (A.R. Smith et al. 970, Guárico (Plaza et al. s/n, UC). PORT, UC). Zulia. Ampliamente distribuida en todo el país (Amazo- nas, Aragua, Bolívar, Distrito Federal, Falcón, Asplenium serratum L. Lara, Mérida, Miranda, Nueva Esparta, Sucre, Helecho epífita. Bosques de galería y bosques Trujillo, Yaracuy y Zulia). caducifolios. Apure y Barinas (C. Hernández 1066, MER). 197 CATÁLOGO ANOTADO E ILUSTRADO DE LA FLORA VASCULAR DE LOS LLANOS DE VENEZUELA Lámina 1 ASPLENIACEAE: a = Asplenium fournieri, con soros. AZOLLACEAE: b = Azolla caroliniana. BLECHNACEAE: c = Blechnum occidentale. CYATHEACEAE: d= Cyathea microdonta, con soros. DENNSTAEDTIACEAE: e = Pteridium caudatum, con soros; f = Saccoloma elegans, con soros. 198 CATÁLOGO ANOTADO E ILUSTRADO DE LA FLORA VASCULAR DE LOS LLANOS DE VENEZUELA Lomagramma Blechnum serrulatum Rich. DRYOPTERIDACEAE Helecho terrestre. Sotobosque de bosques húmedos. Anzoátegui (A. Fernández et al. 12343, PORT) y Bolbitis portoricensis (Spreng.) Hennipman Apure (G. Davidse y A. González 14148, MO, UC). [Lám. 2-a] Ampliamente distribuida en todo el país (Amazo- Helecho terrestre. Bosques de galería. nas, Bolívar, Delta Amacuro, Falcón, Mérida, Anzoátegui, Barinas y Portuguesa (G. Davidse et al. Miranda y Zulia). 21507, MO, PORT). Ampliamente distribuida en todo el país (Aragua, Bolívar, Carabobo, Mérida, Miranda, Táchira, CYATHEACEAE Yaracuy y Zulia). Cyathea lockwoodiana (P.G. Windisch) Lellinger Bolbitis serratifolia (Mert.) Schott Helecho terrestre. Bosques húmedos. Helecho terrestre. Bosques de galería. Apure y Barinas (R. Liesner y A. González 9328, MO, Anzoátegui, Barinas y Portuguesa (B. Stergios et al. UC). 6613, PORT). Andes (Mérida, Táchira y Zulia). Norte de Venezuela (Aragua, Carabobo y Miran- da), Bolívar y Mérida. Cyathea microdonta (Desv.) Domin [Lám. 1-d] Trichipteris microdonta (Desv.) R.M. Tryon Ctenitis refulgens (Klotzsch) Vareschi [Lám. 2-b] Helecho terrestre. Bosques húmedos y de galería. Helecho terrestre. Bosques húmedos. Anzoátegui, Apure, Barinas y Portuguesa (B. Ster- Apure (J.A. Steyermark et al. 101690, VEN) y Barinas. gios y G. Aymard 5628, PORT). Guayana (Amazonas y Bolívar) y Zulia. Ampliamente distribuida en todo el país (Amazo- nas, Bolívar, Carabobo, Delta Amacuro, Lara Cyclodium guianense (Klotzsch) van der Werff Yaracuy y Zulia). [Lám. 2-c] Helecho terrestre. Bosques húmedos. Apure (G. Davidse y A. González 21735, MO). DENNSTAEDTIACEAE Guayana (Amazonas, Bolívar y Delta Amacuro) y Pteridium caudatum (L.) Maxon [Lám. 1-e] Táchira. Pteridium aquilinum (L.) Kuhn var. caudatum (L.) Sadeb. Cyclopeltis semicordata (Sw.) J. Sm. [Lám. 2-d] Helecho terrestre, creciendo en colonias. Ecotono Polypodium semicordatum Sw. sabana-bosque, lugares quemados y orillas de Polypodium caducum Humb. & Bonpl. caminos. Helecho terrestre. Bosques húmedos. Portuguesa (G. Davidse y A. González 21515, MO, Barinas y Monagas (R.A. Pursell et al. 9134, VEN) PORT, VEN) Ampliamente distribuida en todo el país (Aragua, Ampliamente distribuida en todo el país (Amazo- Bolívar, Falcón, Mérida, Miranda, Portuguesa y nas, Bolívar, Carabobo, Distrito Federal, Falcón, Sucre). Lara, Mérida, Miranda, Sucre, Táchira, Yaracuy y Zulia). Elaphoglossum luridum (Fée) H. Christ Helecho epífita. Bosques húmedos. Saccoloma elegans Kaulf. [Lám. 1-f] Anzoátegui y Apure (H. van der Werff y A. González Helecho terrestre. Bosques húmedos. 4741, MO, VEN). Apure (H. van der Werff y A. González 4754, MO, Guayana (Amazonas, Bolívar y Delta Amacuro). PORT). Norte de Venezuela (Aragua y Miranda) y Guaya- Elaphoglossum ortegae Mickel [Lám. 2-e] na (Amazonas y Bolívar). Helecho terrestre, a veces epífita. Bosques de galería. Saccoloma inaequale (Kunze) Mett. Barinas y Portuguesa (F. Ortega et al. 1423, NY, PORT). Helecho terrestre. Bosques húmedos. Andes (Mérida y Táchira) y Aragua. Barinas (R. Liesner y A. González 9338, MO, UC, VEN). Norte de Venezuela (Aragua, Carabobo, Falcón, Lomagramma guianensis (Aubl.) Ching Miranda y Monagas) y Guayana (Amazonas y Polypodium guianense Aubl. Bolívar). Helecho escandente, hemiepífito. Bosques húmedos. Apure (H. van der Werff y A. González 4674, MO, NY). 199 CATÁLOGO ANOTADO E ILUSTRADO DE LA FLORA VASCULAR DE LOS LLANOS DE VENEZUELA Lámina 2 DRYOPTERIDACEAE: a = Bolbitis portoricensis, fronde vegetativa (izq.), fronde fértil (der.); b = Ctenitis refulgens, con soros y rizoma; c = Cyclodium guianense, con soros; d = Cyclopeltis semicordata, con soros; e = Elaphoglossum ortegae, con fronde fértil; f = Nephrolepis biserrata, con soros. 200 CATÁLOGO ANOTADO E ILUSTRADO DE LA FLORA VASCULAR DE LOS LLANOS DE VENEZUELA Lindsaea Andes (Táchira y Zulia) y Guayana (Amazonas y EQUISETACEAE Bolívar). Equisetum giganteum L. [Lám. 3-c] Nephrolepis biserrata (Sw.) Schott [Lám. 2-f] Helecho terrestre, a veces acúatica. Cultivos de arroz. Helecho terreste. Bosques de galería. Portuguesa (F. Ortega et al. 2725, PORT). Portuguesa (B. Stergios y A. Utrera 2471, PORT, UC). Norte de Venezuela (Aragua, Distrito Federal, Lara Norte de Venezuela (Aragua, Falcón, Nueva Es- y Miranda). parta y Sucre) y Guayana (Amazonas, Bolívar y Delta Amacuro). HYMENOPHYLLACEAE Polybotrya caudata Kunze [Lám. 3-a] Helecho terrestre. Bosques húmedos. Trichomanes elegans Rich. Portuguesa (F. Ortega 636, PORT, UC). Helecho terrestre o en rocas. Bosques húmedos. Ampliamente distribuida en todo el país (Amazo- Apure (J.A. Steyermark et al. 101550, UC, VEN) y nas, Aragua, Bolívar, Delta Amacuro, Miranda, Barinas. Sucre y Táchira). Ampliamente distribuida en todo el país (Amazo- nas, Aragua, Bolívar, Distrito Federal, Miranda, Polybotrya osmundacea Humb. & Bonpl. Sucre, Táchira, Yaracuy y Zulia). Polybotrya aristeguietae Brade Polybotrya vareschii Brade Trichomanes hostmannianum (Klotzsch) Kunze Helecho terrestre. Bosques húmedos. [Lám. 3-d] Anzoátegui, Apure (J.A. Steyermark et al. 101552, Helecho terrestre o en rocas. Bosques húmedos. UC, VEN), Barinas y Monagas. Apure (G. Davidse y A. González 13845, MO, UC, Ampliamente distribuida en todo el país (Ama- VEN). zonas, Aragua, Distrito Federal, Falcón, Lara, Guayana (Amazonas, Bolívar y Delta Amacuro). Miranda, Nueva Esparta, Sucre, Táchira, Yara- cuy y Zulia). Trichomanes pinnatum Hedw. Helecho terrestre. Bosques húmedos. Tectaria heracleifolia (Willd.) Underw. Apure (G. Davidse y A. González 13996, MO, UC, Helecho terrestre. Bosques de galería. VEN) y Portuguesa (B. Stergios y A. Utrera 2468, Anzoátegui y Portuguesa (F. Ortega y B. Stergios PORT). 1316, PORT). Ampliamente distribuida en todo el país (Amazo- Andes (Mérida y Zulia) y Norte de Venezuela (Ara- nas, Bolívar, Delta Amacuro, Mérida, Miranda, gua, Distrito Federal, Falcón, Lara, Miranda, Táchira, Yaracuy y Zulia). Nueva Esparta, Sucre y Yaracuy). ISOËTACEAE Tectaria incisa Cav. [Lám. 3-b] Tectaria martinicensis (Spreng.) Copel. Isoëtes triangula U. Weber [Lám. 3-e] Helecho
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