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Scorpions of Puerto Rico and Its Satellite Islands (Scorpiones: Buthidae, Diplocentridae): an Аnnotated List, Key for Genera, and Bibliography

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Scorpions of Puerto Rico and Its Satellite Islands (Scorpiones: Buthidae, Diplocentridae): an Аnnotated List, Key for Genera, and Bibliography Scorpions of Puerto Rico and its satellite islands (Scorpiones: Buthidae, Diplocentridae): an аnnotated list, key for genera, and bibliography Luis F. de Armas May 2020 — No. 311 Euscorpius Occasional Publications in Scorpiology EDITOR: Victor Fet, Marshall University, ‘[email protected]’ ASSOCIATE EDITOR: Michael E. Soleglad, ‘[email protected]’ TECHNICAL EDITOR: František Kovařík, ‘[email protected]’ Euscorpius is the first research publication completely devoted to scorpions (Arachnida: Scorpiones). Euscorpius takes advantage of the rapidly evolving medium of quick online publication, at the same time maintaining high research standards for the burgeoning field of scorpion science (scorpiology).Euscorpius is an expedient and viable medium for the publication of serious papers in scorpiology, including (but not limited to): systematics, evolution, ecology, biogeography, and general biology of scorpions. Review papers, descriptions of new taxa, faunistic surveys, lists of museum collections, and book reviews are welcome. Derivatio Nominis The name Euscorpius Thorell, 1876 refers to the most common genus of scorpions in the Mediterranean region and southern Europe (family Euscorpiidae). Euscorpius is located at: https://mds.marshall.edu/euscorpius/ Archive of issues 1-270 see also at: http://www.science.marshall.edu/fet/Euscorpius (Marshall University, Huntington, West Virginia 25755-2510, USA) ICZN COMPLIANCE OF ELECTRONIC PUBLICATIONS: Electronic (“e-only”) publications are fully compliant with ICZN (International Code of Zoological Nomenclature) (i.e. for the purposes of new names and new nomenclatural acts) when properly archived and registered. All Euscorpius issues starting from No. 156 (2013) are archived in two electronic archives: • Biotaxa, http://biotaxa.org/Euscorpius (ICZN-approved and ZooBank-enabled) • Marshall Digital Scholar, http://mds.marshall.edu/euscorpius/. (This website also archives all Euscorpius issues previously published on CD-ROMs.) Between 2000 and 2013, ICZN did not accept online texts as “published work” (Article 9.8). At this time, Euscorpius was produced in two identical versions: online (ISSN 1536-9307) and CD-ROM (ISSN 1536-9293) (laser disk) in archive-quality, read-only format. Both versions had the identical date of publication, as well as identical page and figure numbers. Only copies distributed on a CD-ROM from Euscorpius in 2001-2012 represent published work in compliance with the ICZN, i.e. for the purposes of new names and new nomenclatural acts. In September 2012, ICZN Article 8. What constitutes published work, has been amended and allowed for electronic publications, disallowing publication on optical discs. From January 2013, Euscorpius discontinued CD-ROM production; only online electronic version (ISSN 1536-9307) is published. For further details on the new ICZN amendment, see http://www.pensoft.net/journals/zookeys/article/3944/. Publication date: 21 May 2020 http://zoobank.org/urn:lsid:zoobank.org:pub:0343EFAE-9A2A-4FC8-A84B-579B22D741B3 Euscorpius - Occasional Publications in Scorpiology. 2020, No. 311 Scorpions of Puerto Rico and its satellite islands (Scorpiones: Buthidae, Diplocentridae): an аnnotated list, key for genera, and bibliography Luis F. de Armas P. O. Box 4327, San Antonio de los Baños, Artemisa 38100, Cuba; [email protected] http://zoobank.org/urn:lsid:zoobank.org:pub:0343EFAE-9A2A-4FC8-A84B-579B22D741B3 Summary An annotated list of the scorpions of Puerto Rico and its satellite islands (mainly Vieques, Culebra, Caja de Muertos, Magueyes, Desecheo, and Mona), a key for the seven genera present in those insular territories, a key for the three species of Microtityus, and the basic bibliography on taxonomy and natural history are provided. Of the 16 confirmed species described from these Greater Antillean islands, four are known only from a single sex and only one or two adult specimens. Seven species (six of them from the genus Tityus), are endemic to the Puerto Rico Island; Vieques has two species, whereas Mona Island and Culebrita Island have one species each. At least two species are shared with Hispaniola, whereas one is also found in the Virgin Islands. The only introduced species is Isometrus maculatus (De Geer, 1778). The buthid Tityus dasyurus Pocock, 1897, described from “Porto Rico”, likely is not a Puerto Rican taxon. Introduction The genera Cazierius Francke, 1978 and Rhopalurus Thorell, 1876 [the Greater Antillean species of the latter were The island of Puerto Rico (Fig. 1) is situated in the Caribbean transferred by Esposito et al. (2017) to Heteroctenus Pocock, Sea and is part of the Greater Antilles, being the smaller among 1893] were recorded from Mona Island by Armas (2005) and those insular territories (the others are Cuba, Hispaniola, and Santiago-Blay (2009), respectively. The presence in Puerto Jamaica). Until the present century, the scorpion fauna of Rico of the genus Microtityus Kjellesvig-Waering, 1966, Puerto Rico and its satellite islands remained poorly known; predicted by Armas (1984: 5), was recently confirmed by for example, Armas (2001a) listed six species belonging to Teruel et al. (2014, 2015), increasing to seven the number of four genera of the families Buthidae and Diplocentridae. scorpion genera present in Puerto Rico and its satellite islands. Fortunately, during the last two decades this situation has In the present contribution, we provide an annotated list of favourably changed and now the list has increased to about 20 the scorpions described or recorded from Puerto Rico and species (Teruel et al., 2015). its satellite islands, a key for the identification of the seven In 1984, Jorge A. Santiago-Blay concluded his six-year genera present in those insular territories, another key for studies on the scorpions of the Greater Puerto Rico Region, members of the genus Microtityus in these islands, and the which includes the islands of Puerto Rico, Mona, Desecheo, basic bibliography on taxonomy and natural history of the Caja de Muertos, Culebra, Vieques, and also the Virgin Islands involved taxa. Solid keys for species of Centruroides Marx, (Santiago-Blay, 1984). A list of the species was published 1890 and Tityus C. L. Koch, 1836 are not possible due to the (Santiago-Blay, 1983), but the new species discovered during insufficient taxonomical information for some of their species, that project waited for 25 years for a formal description mainly those described by Santiago-Blay (2009). (Santiago-Blay, 2009). Unfortunately, these descriptions were very brief, and the promised details are yet unpublished. Of Methods & Material the eight new species described by Santiago-Blay (2009), three have been relegated as junior synonyms (Armas, 2009; The taxonomical information and bibliography are actual until Esposito et al., 2017; Teruel et al., 2017), and perhaps other April 30, 2020. The bibliography is mainly devoted to those two species might be suffering the same fate. Recently, Teruel papers containing relevant data on taxonomy and natural history, et al. (2018) redescribed Tityus estherae Santiago-Blay, although other matters are not excluded. Taxa are alphabetically 2009 and provided additional information on its taxonomy, arranged. Terminology follows Stahnke (1971). The map (Fig. distribution, and natural history. 1) has been adapted from Microsoft Encarta 2009. 2 Euscorpius - 2020, No. 311 Figure 1. Geographical position of Puerto Rico and its most relevant satellite islands. Systematics from the synonymy with C. griseus, and raised it to the species Annotated List level, a nomenclatural action overlooked by Crew & Esposito (2020: fig. 11), who again mentioned it as a subspecies of C. Family Buthidae griseus. If C. griseus borinquensis is a good species, then it Centruroides bani Armas & Marcano Fondeur, 1987. would be most strongly demonstrated. Consequently, at this Distribution. Hispaniola and Mona Island (Puerto Rico). moment caution seems preferable.This is the most common This species was recorded from Mona Island by Esposito et scorpion in Puerto Rico and Virgin Islands, also being al. (2017: 27), but without mentioning the material examined. synanthropic (Santiago-Blay, 2009). Teruel et al. (2017) synonymizes Centruroides mariaorum Santiago-Blay, 2009, described from Mona Island, to C. bani, Centruroides jorgeorum Santiago-Blay, 2009. and added new localities from this small insular territory, as Distribution. Only known from the type locality: Puerto well as notes on its habitat. Rico: Patillas: Barrio Guardarraya, close to Barrio Pollo. This species was barely described on the basis of a single Centruroides griseus (C. L. Koch, 1844). adult female, which is the only known specimen. On July 28, Distribution. Virgin Islands and Puerto Rico, including 2010, personal effort for rediscovering it in the type locality some satellite islands of the last (Vieques, Culebra, Caja de was fruitless. It remains as a species pendant of confirmation Muertos, Magueyes). The presence of C. griseus in Desecheo (Teruel et al., 2015: 12). Island (Torres-González, 1965, cited by Santiago-Blay, 2009: 112) was considered by Teruel et al. (2015: 13) as needed Centruroides sasae Santiago-Blay, 2009. of verification and probably erroneous. The Puerto Rican Distribution. Only known from the type locality: Caja de population was described by Armas (1982) as Centruroides Muertos Island. This species was described on the basis of griseus borinquensis, but later downgraded by him to a single adult male, which is the
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