Radiocarbon Variations from Tasmanian Conifers: Results from Three Early Holocene Logs
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Evolution of the Female Conifer Cone Fossils, Morphology and Phylogenetics
DEPARTMENT OF BIOLOGICAL AND ENVIRONMENTAL SCIENCES EVOLUTION OF THE FEMALE CONIFER CONE FOSSILS, MORPHOLOGY AND PHYLOGENETICS Daniel Bäck Degree project for Bachelor of Science with a major in Biology BIO602, Biologi: Examensarbete – kandidatexamen, 15 hp First cycle Semester/year: Spring 2020 Supervisor: Åslög Dahl, Department of Biological and Environmental Sciences Examiner: Claes Persson, Department of Biological and Environmental Sciences Front page: Abies koreana (immature seed cones), Gothenburg Botanical Garden, Sweden Table of contents 1 Abstract ............................................................................................................................... 2 2 Introduction ......................................................................................................................... 3 2.1 Brief history of Florin’s research ............................................................................... 3 2.2 Progress in conifer phylogenetics .............................................................................. 4 3 Aims .................................................................................................................................... 4 4 Materials and Methods ........................................................................................................ 4 4.1 Literature: ................................................................................................................... 4 4.2 RStudio: ..................................................................................................................... -
Pollination Drop in Relation to Cone Morphology in Podocarpaceae: a Novel Reproductive Mechanism Author(S): P
Pollination Drop in Relation to Cone Morphology in Podocarpaceae: A Novel Reproductive Mechanism Author(s): P. B. Tomlinson, J. E. Braggins, J. A. Rattenbury Source: American Journal of Botany, Vol. 78, No. 9 (Sep., 1991), pp. 1289-1303 Published by: Botanical Society of America Stable URL: http://www.jstor.org/stable/2444932 . Accessed: 23/08/2011 15:47 Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at . http://www.jstor.org/page/info/about/policies/terms.jsp JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. Botanical Society of America is collaborating with JSTOR to digitize, preserve and extend access to American Journal of Botany. http://www.jstor.org AmericanJournal of Botany 78(9): 1289-1303. 1991. POLLINATION DROP IN RELATION TO CONE MORPHOLOGY IN PODOCARPACEAE: A NOVEL REPRODUCTIVE MECHANISM' P. B. TOMLINSON,2'4 J. E. BRAGGINS,3 AND J. A. RATTENBURY3 2HarvardForest, Petersham, Massachusetts 01366; and 3Departmentof Botany, University of Auckland, Auckland, New Zealand Observationof ovulatecones at thetime of pollinationin the southernconiferous family Podocarpaceaedemonstrates a distinctivemethod of pollencapture, involving an extended pollinationdrop. Ovules in all generaof the family are orthotropousand singlewithin the axil of each fertilebract. In Microstrobusand Phyllocladusovules are-erect (i.e., the micropyle directedaway from the cone axis) and are notassociated with an ovule-supportingstructure (epimatium).Pollen in thesetwo genera must land directly on thepollination drop in theway usualfor gymnosperms, as observed in Phyllocladus.In all othergenera, the ovule is inverted (i.e., the micropyleis directedtoward the cone axis) and supportedby a specializedovule- supportingstructure (epimatium). -
Methods for Measuring Frost Tolerance of Conifers: a Systematic Map
Review Methods for Measuring Frost Tolerance of Conifers: A Systematic Map Anastasia-Ainhoa Atucha Zamkova *, Katherine A. Steele and Andrew R. Smith School of Natural Sciences, Bangor University, Bangor LL57 2UW, Gwynedd, UK; [email protected] (K.A.S.); [email protected] (A.R.S.) * Correspondence: [email protected] Abstract: Frost tolerance is the ability of plants to withstand freezing temperatures without unrecov- erable damage. Measuring frost tolerance involves various steps, each of which will vary depending on the objectives of the study. This systematic map takes an overall view of the literature that uses frost tolerance measuring techniques in gymnosperms, focusing mainly on conifers. Many different techniques have been used for testing, and there has been little change in methodology since 2000. The gold standard remains the field observation study, which, due to its cost, is frequently substituted by other techniques. Closed enclosure freezing tests (all non-field freezing tests) are done using various types of equipment for inducing artificial freezing. An examination of the literature indicates that several factors have to be controlled in order to measure frost tolerance in a manner similar to observation in a field study. Equipment that allows controlling the freezing rate, frost exposure time and thawing rate would obtain results closer to field studies. Other important factors in study design are the number of test temperatures used, the range of temperatures selected and the decrements between the temperatures, which should be selected based on expected frost tolerance of the tissue and species. Citation: Atucha Zamkova, A.-A.; Steele, K.A.; Smith, A.R. -
Podocarpus Has a fleshy Enlarged Stem with the Seed Capsule Attached
Some members of the Podocarpaceae It now seems as if the general lock down is being eased across the country and I am looking to do other things than write these notes which seemed helpful eleven weeks ago but hopefully soon will not be so. Not quite a bakers dozen but one more than a S.I. preferred unit. Also I must get on with the June FACTT Newsletter which I like to think is as eagerly awaited as these notes. This note is about a family, members of which are found in fossils of Gondwana times and now in those countries which formed the original landmass and some other countries. Podocarpus has a fleshy enlarged stem with the seed capsule attached. It is easy to see the possibility of birds spreading the seeds, unlike Nothofagus where the seed is less likely to be spread widely one would think. Regarding the movement of continents it needs to be mentioned that a million is a very large number. If, as Australia is presently doing, a landmass moves 70mm a year in just 1 million years; a short period of time in geological terms; the distance traveled is 70km. Australia is moving so fast in fact that GPS can’t keep up. Some people think that taxonomists fall into two broad categories - Lumpers and Splinters. Lumpers tend to say -Well the species are so similar they are in fact one. Splitters on the other hand tend to the view that the fine details represent sufficient differences to recognise they are different species. -
Abstract Clark, Catherine Margaret
ABSTRACT CLARK, CATHERINE MARGARET. Genetic diversity and phylogeography in a Tasmanian rainforest conifer (Lagarostrobos franklinii (Hook f.) Quinn Podocarpaceae. (Under the direction of Dr. Ronald Sederoff and Dr. Thomas Wentworth). Genetic variation in Huon pine (Lagarostrobos franklinii), a Tasmanian rainforest conifer, was estimated using several marker systems and two spatial scales. Genealogy based methods were used to infer population history of eight Huon pine stands based on chloroplast DNA variation. Chloroplast nucleotide diversity (π) was low (0.00093± 0.00006) in a multilocus haplotype generated by three universal chloroplast primers (trnS-trnT, trnD-trnT, psbC-trnS). Five haplotypes were identified; two were€ widely distributed but the most frequently occurring haplotype was found only in trees in the western portion of the range. Genetic differentiation among populations was significant and showed a high degree of structure (GST = 0.26077 p ≤ 0.00001). Pairwise comparisons between populations revealed significant structure between the southeastern and northwestern watersheds and significant isolation€ by distance€ ( p ≤ 0.0220). Nucleotide variation was also assessed in segments of three nuclear genes, 4Cl (4- € coumarate: coenzyme A ligase), ITS2 (intergenic spacer region of ribosomal DNA) and G3pdh (glyceraldehyde 3-phosphate dehydrogenase). A total of 1,154 base pairs were sequenced from 79 individuals (158 alleles) representing seven geographic locations. Estimates of nucleotide diversity (π = 0.00089 ± 0.00007) and theta (θ) (0.00061 ± 0.00031) were low for the combined loci and similar to chloroplast estimates. There was a higher level of variation at the 4Cl locus (π = 0.00167 ± 0.00014) associated with recombination. Nucleotide diversity for nuclear loci was highest in the subalpine Mt. -
Tassie's Trees Black Peppermint - Eucalyptus Gum-Top Stringy Bark - Eucalyptus Amygdalina Delegatensis
Tassie's Trees Black peppermint - Eucalyptus Gum-top stringy bark - Eucalyptus amygdalina delegatensis Black peppermint is a tree of the more open This impressive gum tree is one of the giants of forests that can grow up to 30 metres tall. It can Tasmanian forests and can reach heights of up to be found mostly in the drier areas of Tasmania, 90 metres. This tree is the dominant forest species often in association with white gum and silver that grow in the cooler, higher altitude areas. It is wattle. This species is only found in Tasmania. one of the ash group of eucalypts which include swamp gum and stringy bark. White gum - Eucalyptus viminalis Silver wattle - Acacia dealbata White gum is a tree with many forms. It can grow Silver wattle is a distinctive tree or large shrub of to over 90 metres tall in the Evercreech forests the wattle family. It is widely distributed near Fingal in the north east, but it is usually throughout eucalypt forests and woodlands in found growing among black peppermint in drier eastern Australia. The tree can easily be identified eucalypt forests. White gum is a major species in by its blue-green feathery foliage and yellow the open woodlands of Tasmania. flowers during spring. It is more valued for horticultural uses than for its timber. Tassie's Trees Native Laurel - Anopterus King Billy pine - Athrotaxis glandulosus selaginoides More an understorey shrub rather than a canopy Although related to the famous redwoods of tree, this endemic species is wide-spread in wet California, the King Billy pine is only a medium eucalypt forests and rainforest up to 1200m above sized tree, usually between 25 and 30 metres sea level. -
The Ecology and Conservation of Bryophytes in Tasmanian Wet Eucalypt Forest
The ecology and conservation of bryophytes in Tasmanian wet eucalypt forest Perpetua A. M. Turner B.Sc. (Hons) Submitted in fulfilment of the requirements of the degree of Doctor of Philosophy School of Geography and Environmental Studies University of Tasmania Hobart July 2003 Declaration This thesis contains no material that has been accepted for the award of any other degree or diploma in any tertiary institution and, to the best of my knowledge and belief, the thesis contains no material previously published or written by another person, except where due reference is made in the text. Perpetua A.M. Turner (nee Blanks) 11th July 2003 Authority of Access This thesis may be made available forloan. Copying of any part of this thesis is prohibited fortwo years fromthe date this statement was signed; after that time limited copying is permitted in accordance with the Copyright Act 1968. Perpetua A.M. Turner (nee Blanks) 11 th July 2003 11 Abstract The aim of this study was to determine the factors that affect bryophyte richness and species composition in wet eucalypt forest, including old growth and forest disturbed by wildfire or silvicultural practice. Approximately one third of the total bryophyte flora for Tasmania was recorded in old growth mixed forest, with more liverwort than moss species found. Bryophyte species composition was significantly different between groups of sites of forest from the northwest, central and southern areas of the state. Mean annual temperature, altitude, rainfall of the driest month and aspect were most significant in predicting variation in bryophyte species composition. The use of vascular plants as surrogates for the conservation ofbryophyte species was examined. -
Document Title 1
Document title 1 Contents CHAPTER 1. Context.................................................................................................................................. 2 1.1 Purpose ...................................................................................................................................................................................... 3 1.2 Scope .......................................................................................................................................................................................... 3 1.3 Policy and legislative framework .......................................................................................................................................... 3 1.4 Development and custodianship of this document ......................................................................................................... 4 CHAPTER 2. Rainforest Ecology ................................................................................................................ 5 2.1 Definition and classification ................................................................................................................................................... 6 2.1.1 Rainforest types ............................................................................................................................................................... 6 2.1.2 Further classification ...................................................................................................................................................... -
Forest to Fjaeldmark: Rainforest and Related Scrub
Edition 2 From Forest to Fjaeldmark The Vegetation Communities Rainforest and related scrub Eucryphia lucida Edition 2 From Forest to Fjaeldmark (revised – May 2018) 1 Rainforest and related scrub Community (Code) Page Athrotaxis cupressoides-Nothofagus gunnii short rainforest (RPF) 10 Athrotaxis cupressoides open woodland (RPW) 12 Athrotaxis cupressoides rainforest (RPP) 15 Athrotaxis selaginoides-Nothofagus gunnii short rainforest (RKF) 17 Athrotaxis selaginoides rainforest (RKP) 19 Athrotaxis selaginoides subalpine scrub (RKS) 21 Coastal rainforest (RCO) 23 Highland low rainforest and scrub (RSH) 25 Highland rainforest scrub with dead Athrotaxis selaginoides (RKX) 27 Lagarostrobos franklinii rainforest and scrub (RHP) 29 Nothofagus-Atherosperma rainforest (RMT) 31 Nothofagus-Leptospermum short rainforest (RML) 34 Nothofagus-Phyllocladus short rainforest (RMS) 36 Nothofagus gunnii rainforest scrub (RFS) 39 Nothofagus rainforest (undifferentiated) (RMU) 41 Rainforest fernland (RFE) 42 General description Diselma archeri and/or Pherosphaera hookeriana. These are mapped within the unit Highland This group comprises most Tasmanian vegetation coniferous shrubland (HCS), which is included within dominated by Tasmanian rainforest species (sensu the Highland treeless vegetation section. Jarman and Brown 1983) regardless of whether the dominant species are trees, shrubs or ferns. Rainforests and related scrub vegetation generally Tasmanian cool temperate rainforest has been occur in high rainfall areas of Tasmania that exceed defined floristically -
Ethnobotany, Phytochemistry and Pharmacology of Podocarpus Sensu Latissimo (S.L.) ⁎ H.S
Available online at www.sciencedirect.com South African Journal of Botany 76 (2010) 1–24 www.elsevier.com/locate/sajb Review Ethnobotany, phytochemistry and pharmacology of Podocarpus sensu latissimo (s.l.) ⁎ H.S. Abdillahi, G.I. Stafford, J.F. Finnie, J. Van Staden Research Centre for Plant Growth and Development, School of Biological and Conservation Sciences, University of KwaZulu-Natal Pietermaritzburg, Private Bag X01, Scottsville 3209, South Africa Received 26 August 2009; accepted 2 September 2009 Abstract The genus Podocarpus sensu latissimo (s.l.) was initially subdivided into eight sections. However, based on new information from different morphological and anatomical studies, these sections were recognised as new genera. This change in nomenclature sometimes is problematic when consulting ethnobotanical data especially when selecting plants for pharmacological screening, thus there is a need to clear any ambiguity with the nomenclature. Species of Podocarpus s.l. are important timber trees in their native areas. They have been used by many communities in traditional medicine and as a source of income. Podocarpus s.l. is used in the treatment of fevers, asthma, coughs, cholera, distemper, chest complaints and venereal diseases. Other uses include timber, food, wax, tannin and as ornamental trees. Although extensive research has been carried out on species of Podocarpus s.l over the last decade, relatively little is known about the African species compared to those of New Zealand, Australia, China and Japan. Phytochemical studies have led to the isolation and elucidation of various terpenoids and nor- and bis- norditerpenoid dilactones. Biflavonoids of the amentoflavone and hinokiflavone types have also been isolated. -
Podocarp Evolution: a Molecular Phylogenetic Perspective Edward Biffin, John G
1 Podocarp Evolution: A Molecular Phylogenetic Perspective Edward Biffin, John G. Conran, and Andrew J. Lowe ABSTRACT. Phylogenetic reconstructions of the relationships among extant taxa can be used to infer the nature of the processes that have generated contemporary patterns of biotic diversity. In this study, we present a molecular phylogenetic hypothesis for the conifer family Podocarpaceae based upon three DNA fragments that have been sampled for approximately 90 taxa. We use Bayesian relaxed- clock methods and four fossil con- straints to estimate divergence times among the lineages of Podocarpaceae. Our dating analyses suggest that although the family is old (Triassic–Jurassic), the extant species groups are of recent evolutionary origin (mid- to late Cenozoic), a pattern that could reflect a temporal increase in the rate lineage accumulation or, alternatively, a high and constant rate of extinction. Our data do not support the hypothesis that Podocarpaceae have diversified at a homogeneous rate, instead providing strong evidence for a three- to eightfold increase in diversification associated with the Podocarpoid– Dacrydioid clade, which radiated in the mid- to late Cretaceous to the earliest Cenozoic, around 60–94 mya. This group includes a predominance of taxa that develop broad leaves and/or leaf- like shoots and are distributed predominantly throughout the tropics. Tropical podocarps with broad leaves may have experienced reduced extinction and/or increased speciation coincident with the radiation of the angiosperms, the expansion of megathermal forests, and relatively stable tropical climates that were widespread through the Tertiary. Edward Biffin, John G. Conran, and Andrew J. INTRODUCTION Lowe, Australian Centre for Evolutionary Biol- ogy and Biodiversity, School of Earth and Envi- Patterns of species diversity reflect the balance of speciation and extinction ronmental Science, The University of Adelaide, Adelaide, South Australia 5005, Australia. -
3193VEA Otway Report PDF
ANGAHOOK-OTWAY INVESTIGATION DISCUSSION PAPER VICTORIAN ENVIRONMENTAL ASSESSMENT COUNCIL SEPTEMBER 2003 WHAT IS VEAC? The Victorian Environmental Assessment Council was established in 2002 – under the Victorian Environmental Assessment Council Act 2001 – to replace the Environment Conservation Council (ECC) as the body providing the State Government with independent advice on strategic public land-use planning. The five members of VEAC are: (Chair) Dr Brian Robinson,AM, FTSE, B.Sc., Ph.D. – The former chair of the Environment Protection Authority; a wealth of experience in sustainable development and natural resources management. Dr Sarah Ewing, B.Sc (Hons), M.Sc., Ph.D., Grad. Dip. Ed. – Currently a member of the Victorian Catchment Management Council and formerly a deputy member of the Australian Landcare Council; many years experience in catchment and natural resource management issues. Mr Duncan Malcolm – Currently the Chairman of the Gippsland Coastal Board and Watermark Inc.; immediate past Chairman of Lakes and Wilderness Tourism Association, immediate past chair and current board member of Irrigation Association of Australia and member of Victorian Coastal Council. Dr David Mercer, BA (Hons), Ph.D., Dip.Ed. – Fellow of The Environment Institute of Australia and New Zealand; broad expertise in natural resource management, recreation and tourism and also highly regarded academic experience. Mrs Eda Ritchie, Grad. Dip. Bus. – Formerly a member of the Environment Conservation Council,Trust for Nature Board and the Chairperson of the Western Region Coastal Board; has a strong background in farming in Western Victoria. SUBMISSIONS INVITED Submissions are now invited from interested groups and individuals for consideration for the Draft Proposals Paper to be published early in 2004.