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Phylogenetic Relationships and Historical Biogeography of Tribes and Genera in the Subfamily Nymphalinae (Lepidoptera: Nymphalidae)
Blackwell Science, LtdOxford, UKBIJBiological Journal of the Linnean Society 0024-4066The Linnean Society of London, 2005? 2005 862 227251 Original Article PHYLOGENY OF NYMPHALINAE N. WAHLBERG ET AL Biological Journal of the Linnean Society, 2005, 86, 227–251. With 5 figures . Phylogenetic relationships and historical biogeography of tribes and genera in the subfamily Nymphalinae (Lepidoptera: Nymphalidae) NIKLAS WAHLBERG1*, ANDREW V. Z. BROWER2 and SÖREN NYLIN1 1Department of Zoology, Stockholm University, S-106 91 Stockholm, Sweden 2Department of Zoology, Oregon State University, Corvallis, Oregon 97331–2907, USA Received 10 January 2004; accepted for publication 12 November 2004 We infer for the first time the phylogenetic relationships of genera and tribes in the ecologically and evolutionarily well-studied subfamily Nymphalinae using DNA sequence data from three genes: 1450 bp of cytochrome oxidase subunit I (COI) (in the mitochondrial genome), 1077 bp of elongation factor 1-alpha (EF1-a) and 400–403 bp of wing- less (both in the nuclear genome). We explore the influence of each gene region on the support given to each node of the most parsimonious tree derived from a combined analysis of all three genes using Partitioned Bremer Support. We also explore the influence of assuming equal weights for all characters in the combined analysis by investigating the stability of clades to different transition/transversion weighting schemes. We find many strongly supported and stable clades in the Nymphalinae. We are also able to identify ‘rogue’ -
List of Animal Species with Ranks October 2017
Washington Natural Heritage Program List of Animal Species with Ranks October 2017 The following list of animals known from Washington is complete for resident and transient vertebrates and several groups of invertebrates, including odonates, branchipods, tiger beetles, butterflies, gastropods, freshwater bivalves and bumble bees. Some species from other groups are included, especially where there are conservation concerns. Among these are the Palouse giant earthworm, a few moths and some of our mayflies and grasshoppers. Currently 857 vertebrate and 1,100 invertebrate taxa are included. Conservation status, in the form of range-wide, national and state ranks are assigned to each taxon. Information on species range and distribution, number of individuals, population trends and threats is collected into a ranking form, analyzed, and used to assign ranks. Ranks are updated periodically, as new information is collected. We welcome new information for any species on our list. Common Name Scientific Name Class Global Rank State Rank State Status Federal Status Northwestern Salamander Ambystoma gracile Amphibia G5 S5 Long-toed Salamander Ambystoma macrodactylum Amphibia G5 S5 Tiger Salamander Ambystoma tigrinum Amphibia G5 S3 Ensatina Ensatina eschscholtzii Amphibia G5 S5 Dunn's Salamander Plethodon dunni Amphibia G4 S3 C Larch Mountain Salamander Plethodon larselli Amphibia G3 S3 S Van Dyke's Salamander Plethodon vandykei Amphibia G3 S3 C Western Red-backed Salamander Plethodon vehiculum Amphibia G5 S5 Rough-skinned Newt Taricha granulosa -
Superior National Forest
Admirals & Relatives Subfamily Limenitidinae Skippers Family Hesperiidae £ Viceroy Limenitis archippus Spread-wing Skippers Subfamily Pyrginae £ Silver-spotted Skipper Epargyreus clarus £ Dreamy Duskywing Erynnis icelus £ Juvenal’s Duskywing Erynnis juvenalis £ Northern Cloudywing Thorybes pylades Butterflies of the £ White Admiral Limenitis arthemis arthemis Superior Satyrs Subfamily Satyrinae National Forest £ Common Wood-nymph Cercyonis pegala £ Common Ringlet Coenonympha tullia £ Northern Pearly-eye Enodia anthedon Skipperlings Subfamily Heteropterinae £ Arctic Skipper Carterocephalus palaemon £ Mancinus Alpine Erebia disa mancinus R9SS £ Red-disked Alpine Erebia discoidalis R9SS £ Little Wood-satyr Megisto cymela Grass-Skippers Subfamily Hesperiinae £ Pepper & Salt Skipper Amblyscirtes hegon £ Macoun’s Arctic Oeneis macounii £ Common Roadside-Skipper Amblyscirtes vialis £ Jutta Arctic Oeneis jutta (R9SS) £ Least Skipper Ancyloxypha numitor Northern Crescent £ Eyed Brown Satyrodes eurydice £ Dun Skipper Euphyes vestris Phyciodes selenis £ Common Branded Skipper Hesperia comma £ Indian Skipper Hesperia sassacus Monarchs Subfamily Danainae £ Hobomok Skipper Poanes hobomok £ Monarch Danaus plexippus £ Long Dash Polites mystic £ Peck’s Skipper Polites peckius £ Tawny-edged Skipper Polites themistocles £ European Skipper Thymelicus lineola LINKS: http://www.naba.org/ The U.S. Department of Agriculture (USDA) prohibits discrimination http://www.butterfliesandmoths.org/ in all its programs and activities on the basis of race, color, national -
Assessing Landscape Effects on Genetics and Dispersal of the Rocky Mountain Apollo Butterfly Arnassiusp Smintheus Using a Resistance Mapping Approach
Western University Scholarship@Western Electronic Thesis and Dissertation Repository 10-27-2017 10:30 AM Assessing Landscape Effects on Genetics and Dispersal of the Rocky Mountain Apollo Butterfly arnassiusP smintheus using a Resistance Mapping Approach Ning Chen The University of Western Ontario Supervisor Dr. Nusha Keyghobadi The University of Western Ontario Graduate Program in Biology A thesis submitted in partial fulfillment of the equirr ements for the degree in Master of Science © Ning Chen 2017 Follow this and additional works at: https://ir.lib.uwo.ca/etd Part of the Biology Commons Recommended Citation Chen, Ning, "Assessing Landscape Effects on Genetics and Dispersal of the Rocky Mountain Apollo Butterfly Parnassius smintheus using a Resistance Mapping Approach" (2017). Electronic Thesis and Dissertation Repository. 5058. https://ir.lib.uwo.ca/etd/5058 This Dissertation/Thesis is brought to you for free and open access by Scholarship@Western. It has been accepted for inclusion in Electronic Thesis and Dissertation Repository by an authorized administrator of Scholarship@Western. For more information, please contact [email protected]. Abstract Landscape variables that best explain genetic differentiation may not also best explain dispersal patterns, but many studies use genetic differentiation as a proxy for dispersal. I tested the effects of landscape on both genetic differentiation and dispersal in parallel, to explore whether landscape effects on genetic differentiation between populations and landscape effects on dispersal would be comparable in such contexts. I used circuit theory (Circuitscape) and least cost transect analysis to evaluate the effects of landscape on both movement and genetic differentiation of the butterfly, Parnassius smintheus, in the Jumpingpound Ridge study system. -
Guidance Document on the Strict Protection of Animal Species of Community Interest Under the Habitats Directive 92/43/EEC
Guidance document on the strict protection of animal species of Community interest under the Habitats Directive 92/43/EEC Final version, February 2007 1 TABLE OF CONTENTS FOREWORD 4 I. CONTEXT 6 I.1 Species conservation within a wider legal and political context 6 I.1.1 Political context 6 I.1.2 Legal context 7 I.2 Species conservation within the overall scheme of Directive 92/43/EEC 8 I.2.1 Primary aim of the Directive: the role of Article 2 8 I.2.2 Favourable conservation status 9 I.2.3 Species conservation instruments 11 I.2.3.a) The Annexes 13 I.2.3.b) The protection of animal species listed under both Annexes II and IV in Natura 2000 sites 15 I.2.4 Basic principles of species conservation 17 I.2.4.a) Good knowledge and surveillance of conservation status 17 I.2.4.b) Appropriate and effective character of measures taken 19 II. ARTICLE 12 23 II.1 General legal considerations 23 II.2 Requisite measures for a system of strict protection 26 II.2.1 Measures to establish and effectively implement a system of strict protection 26 II.2.2 Measures to ensure favourable conservation status 27 II.2.3 Measures regarding the situations described in Article 12 28 II.2.4 Provisions of Article 12(1)(a)-(d) in relation to ongoing activities 30 II.3 The specific protection provisions under Article 12 35 II.3.1 Deliberate capture or killing of specimens of Annex IV(a) species 35 II.3.2 Deliberate disturbance of Annex IV(a) species, particularly during periods of breeding, rearing, hibernation and migration 37 II.3.2.a) Disturbance 37 II.3.2.b) Periods -
Spotting Butterflies Says Ulsh
New & Features “A butterfly’s lifespan generally cor- responds with the size of the butterfly,” Spotting Butterflies says Ulsh. The tiny blues often seen in the mountains generally only live about How, when and where to find Lepidoptera in 10 days. Some species, however, will the Cascades and Olympics overwinter in the egg, pupa or chrysalid form (in the cocoon prior to becoming winged adults). A few Northwest spe- cies overwinter as adults, and one—the mourning cloak—lives for ten months, and is the longest lived butterfly in North America. The first thing that butterflies do upon emerging from the chrysalis and unfold- ing their wings is to breed. In their search for mates, some butterflies “hilltop,” or stake out spots on high trees or ridgelines to make themselves more prominent. A butterfly’s wing colors serve two distinct purposes. The dorsal, or upperside of the wings, are colorful, and serve to attract mates. The ventral, or underside of the wings generally serves to camouflage the insects. So a butterfly such as the satyr comma has brilliant orange and yellow Western tiger and pale tiger swallowtail butterflies “puddling.” When looking for for spots when seen with wings open, and butterflies along the trail keep an eye on moist areas or meadows with flowers. a bark-like texture to confuse predators when its wings are closed. By Andrew Engelson Butterfly Association, about where, how As adults, butterflies also seek out Photos by Idie Ulsh and when to look for butterflies in our nectar and water. Butterflies generally mountains. Ulsh is an accomplished Butterflies are the teasers of wildlife. -
Large-Scale Movement Behavior in a Reintroduced Predator Population
doi: 10.1111/ecog.03030 41 126–139 ECOGRAPHY Research Large-scale movement behavior in a reintroduced predator population Frances E. Buderman, Mevin B. Hooten, Jacob S. Ivan and Tanya M. Shenk F. E. Buderman (http://orcid.org/0000-0001-9778-9906) ([email protected]) Dept of Fish, Wildlife, and Conservation Biology, Colorado State Univ., Fort Collins, CO, USA. – M. B. Hooten, U.S. Geological Survey, Colorado Cooperative Fish and Wildlife Research Unit, Dept of Fish, Wildlife, and Conservation Biology and Statistics, Colorado State Univ., Fort Collins, CO, USA, and Graduate Degree Program in Ecology, Colorado State Univ., Fort Collins, CO, USA. – J. S. Ivan, Colorado Parks and Wildlife, Fort Collins, CO, USA. – T. M. Shenk, National Park Service, Great Plains Cooperative Ecosystem Studies Unit, Univ. of Nebraska, Lincoln, NE, USA. Ecography Understanding movement behavior and identifying areas of landscape connectivity is 41: 126–139, 2018 critical for the conservation of many species. However, collecting fine-scale movement doi: 10.1111/ecog.03030 data can be prohibitively time consuming and costly, especially for rare or endangered species, whereas existing data sets may provide the best available information on animal Subject Editor: Timothy Keitt. movement. Contemporary movement models may not be an option for modeling Editor-in-Chief: Miguel Araújo. existing data due to low temporal resolution and large or unusual error structures, but Accepted 17 July 2017 inference can still be obtained using a functional movement modeling approach. We use a functional movement model to perform a population-level analysis of telemetry data collected during the reintroduction of Canada lynx to Colorado. -
MOTHS and BUTTERFLIES LEPIDOPTERA DISTRIBUTION DATA SOURCES (LEPIDOPTERA) * Detailed Distributional Information Has Been J.D
MOTHS AND BUTTERFLIES LEPIDOPTERA DISTRIBUTION DATA SOURCES (LEPIDOPTERA) * Detailed distributional information has been J.D. Lafontaine published for only a few groups of Lepidoptera in western Biological Resources Program, Agriculture and Agri-food Canada. Scott (1986) gives good distribution maps for Canada butterflies in North America but these are generalized shade Central Experimental Farm Ottawa, Ontario K1A 0C6 maps that give no detail within the Montane Cordillera Ecozone. A series of memoirs on the Inchworms (family and Geometridae) of Canada by McGuffin (1967, 1972, 1977, 1981, 1987) and Bolte (1990) cover about 3/4 of the Canadian J.T. Troubridge fauna and include dot maps for most species. A long term project on the “Forest Lepidoptera of Canada” resulted in a Pacific Agri-Food Research Centre (Agassiz) four volume series on Lepidoptera that feed on trees in Agriculture and Agri-Food Canada Canada and these also give dot maps for most species Box 1000, Agassiz, B.C. V0M 1A0 (McGugan, 1958; Prentice, 1962, 1963, 1965). Dot maps for three groups of Cutworm Moths (Family Noctuidae): the subfamily Plusiinae (Lafontaine and Poole, 1991), the subfamilies Cuculliinae and Psaphidinae (Poole, 1995), and ABSTRACT the tribe Noctuini (subfamily Noctuinae) (Lafontaine, 1998) have also been published. Most fascicles in The Moths of The Montane Cordillera Ecozone of British Columbia America North of Mexico series (e.g. Ferguson, 1971-72, and southwestern Alberta supports a diverse fauna with over 1978; Franclemont, 1973; Hodges, 1971, 1986; Lafontaine, 2,000 species of butterflies and moths (Order Lepidoptera) 1987; Munroe, 1972-74, 1976; Neunzig, 1986, 1990, 1997) recorded to date. -
CA Checklist of Butterflies of Tulare County
Checklist of Buerflies of Tulare County hp://www.natureali.org/Tularebuerflychecklist.htm Tulare County Buerfly Checklist Compiled by Ken Davenport & designed by Alison Sheehey Swallowtails (Family Papilionidae) Parnassians (Subfamily Parnassiinae) A series of simple checklists Clodius Parnassian Parnassius clodius for use in the field Sierra Nevada Parnassian Parnassius behrii Kern Amphibian Checklist Kern Bird Checklist Swallowtails (Subfamily Papilioninae) Kern Butterfly Checklist Pipevine Swallowtail Battus philenor Tulare Butterfly Checklist Black Swallowtail Papilio polyxenes Kern Dragonfly Checklist Checklist of Exotic Animals Anise Swallowtail Papilio zelicaon (incl. nitra) introduced to Kern County Indra Swallowtail Papilio indra Kern Fish Checklist Giant Swallowtail Papilio cresphontes Kern Mammal Checklist Kern Reptile Checklist Western Tiger Swallowtail Papilio rutulus Checklist of Sensitive Species Two-tailed Swallowtail Papilio multicaudata found in Kern County Pale Swallowtail Papilio eurymedon Whites and Sulphurs (Family Pieridae) Wildflowers Whites (Subfamily Pierinae) Hodgepodge of Insect Pine White Neophasia menapia Photos Nature Ali Wild Wanderings Becker's White Pontia beckerii Spring White Pontia sisymbrii Checkered White Pontia protodice Western White Pontia occidentalis The Butterfly Digest by Cabbage White Pieris rapae Bruce Webb - A digest of butterfly discussion around Large Marble Euchloe ausonides the nation. Frontispiece: 1 of 6 12/26/10 9:26 PM Checklist of Buerflies of Tulare County hp://www.natureali.org/Tularebuerflychecklist.htm -
Manduca Sexta and Hyles Lineata (Sphingidae), and Helicoverpa Zea (Noctuidae)
VOLUME 60, NUMBER 2 101 weedy Pieridae including Pieris rapae L. and Pontia Argentine Andean and Patagonian Pierid fauna. J.Res.Lepid. 28:137-238. protodice Bdv. & LeC., but it is almost never seen above —— 1997. Impactos antropogenicos sobre la fauna de mariposas 1500m and is completely absent in climates comparable (Lepidoptera: Rhopalocera) de Patagonia austral y Tierra del to that at Las Lenas. The erect, even bushy growth form Fuego. Anales Instituto de la Patagonia (Punta Arenas, Chile), Ser.Cs.Nat. 25: 117-l26. of this plant has no analogue in the native brassicaceous —— 2002. The Californian urban butterfly fauna is dependent on flora of the high Andes. It would seem P. nymphula has alien plants. Diversity & Distributions 8: 31-40. successfully colonized this plant by focusing strictly on small rosettes, whose growth form, with tightly ARTHUR M. SHAPIRO, Center for Population Biology, imbricated leaves, is familiar to it as the mature plant is University of California, Davis, CA 95616 not. Received for publication 9 February 2005; revised and accepted 13 I thank Joanne Smith-Flueck and Santiago Cara for July 2005 companionship afield. LITERATURE CITED GRAVES, S.D. & A. M. SHAPIRO. 2003. Exotics as host plants of the California butterfly fauna. Biol. Cons. 110: 413-433. SHAPIRO,A. M. 1991. The zoogeography and systematics of the Journal of the Lepidopterists’ Society 60(2), 2006, 101–103 SURVIVAL OF FREEZING AND SUBSEQUENT SUMMER ECLOSION BY THREE MIGRATORY MOTHS: MANDUCA SEXTA AND HYLES LINEATA (SPHINGIDAE), AND HELICOVERPA ZEA (NOCTUIDAE). Additional key words: overwintering, Heliothis virescens Hyles lineata (Fabricius) and Helicoverpa zea al., 1995), Nova Scotia (Ferguson, 1955), and Quebec (Boddie) are well known migrants whose overwintering (Handfield, 1999) often in September and October, the limits are apparently poorly known. -
BMC Evolutionary Biology Biomed Central
BMC Evolutionary Biology BioMed Central Research article Open Access Timing major conflict between mitochondrial and nuclear genes in species relationships of Polygonia butterflies (Nymphalidae: Nymphalini) Niklas Wahlberg*1, Elisabet Weingartner2, Andrew D Warren3,4 and Sören Nylin2 Address: 1Laboratory of Genetics, Department of Biology, University of Turku, 20014 Turku, Finland, 2Department of Zoology, Stockholm University, 106 91 Stockholm, Sweden, 3McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, SW 34th Street and Hull Road, PO Box 112710, Gainesville, FL 32611-2710, USA and 4Museo de Zoología, Departamento de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional Autónoma de México, Apdo. Postal 70-399, México, DF 04510 México Email: Niklas Wahlberg* - [email protected]; Elisabet Weingartner - [email protected]; Andrew D Warren - [email protected]; Sören Nylin - [email protected] * Corresponding author Published: 7 May 2009 Received: 20 August 2008 Accepted: 7 May 2009 BMC Evolutionary Biology 2009, 9:92 doi:10.1186/1471-2148-9-92 This article is available from: http://www.biomedcentral.com/1471-2148/9/92 © 2009 Wahlberg et al; licensee BioMed Central Ltd. This is an Open Access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/2.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited. Abstract Background: Major conflict between mitochondrial and nuclear genes in estimating species relationships is an increasingly common finding in animals. Usually this is attributed to incomplete lineage sorting, but recently the possibility has been raised that hybridization is important in generating such phylogenetic patterns. -
Biodiversity Profile of Afghanistan
NEPA Biodiversity Profile of Afghanistan An Output of the National Capacity Needs Self-Assessment for Global Environment Management (NCSA) for Afghanistan June 2008 United Nations Environment Programme Post-Conflict and Disaster Management Branch First published in Kabul in 2008 by the United Nations Environment Programme. Copyright © 2008, United Nations Environment Programme. This publication may be reproduced in whole or in part and in any form for educational or non-profit purposes without special permission from the copyright holder, provided acknowledgement of the source is made. UNEP would appreciate receiving a copy of any publication that uses this publication as a source. No use of this publication may be made for resale or for any other commercial purpose whatsoever without prior permission in writing from the United Nations Environment Programme. United Nations Environment Programme Darulaman Kabul, Afghanistan Tel: +93 (0)799 382 571 E-mail: [email protected] Web: http://www.unep.org DISCLAIMER The contents of this volume do not necessarily reflect the views of UNEP, or contributory organizations. The designations employed and the presentations do not imply the expressions of any opinion whatsoever on the part of UNEP or contributory organizations concerning the legal status of any country, territory, city or area or its authority, or concerning the delimitation of its frontiers or boundaries. Unless otherwise credited, all the photos in this publication have been taken by the UNEP staff. Design and Layout: Rachel Dolores