Diversity and Evolution of Asterids
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Floral Symmetry Affects Speciation Rates in Angiosperms Risa D
Received 25 July 2003 Accepted 13 November 2003 Published online 16 February 2004 Floral symmetry affects speciation rates in angiosperms Risa D. Sargent Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia V6T 1Z4, Canada ([email protected]) Despite much recent activity in the field of pollination biology, the extent to which animal pollinators drive the formation of new angiosperm species remains unresolved. One problem has been identifying floral adaptations that promote reproductive isolation. The evolution of a bilaterally symmetrical corolla restricts the direction of approach and movement of pollinators on and between flowers. Restricting pollin- ators to approaching a flower from a single direction facilitates specific placement of pollen on the pollin- ator. When coupled with pollinator constancy, precise pollen placement can increase the probability that pollen grains reach a compatible stigma. This has the potential to generate reproductive isolation between species, because mutations that cause changes in the placement of pollen on the pollinator may decrease gene flow between incipient species. I predict that animal-pollinated lineages that possess bilaterally sym- metrical flowers should have higher speciation rates than lineages possessing radially symmetrical flowers. Using sister-group comparisons I demonstrate that bilaterally symmetric lineages tend to be more species rich than their radially symmetrical sister lineages. This study supports an important role for pollinator- mediated speciation and demonstrates that floral morphology plays a key role in angiosperm speciation. Keywords: reproductive isolation; pollination; sister group comparison; zygomorphy 1. INTRODUCTION The importance of pollinator-mediated selection in angiosperms is well supported by theory (Kiester et al. -
Early Evolution of the Angiosperm Clade Asteraceae in the Cretaceous of Antarctica
Early evolution of the angiosperm clade Asteraceae in the Cretaceous of Antarctica Viviana D. Barredaa,1,2, Luis Palazzesia,b,1, Maria C. Telleríac, Eduardo B. Oliverod, J. Ian Rainee, and Félix Forestb aDivisión Paleobotánica, Museo Argentino de Ciencias Naturales “Bernardino Rivadavia,” Consejo Nacional de Investigaciones Cientificas y Técnicas, Buenos Aires C1405DJR, Argentina; bJodrell Laboratory, Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3DS, United Kingdom; cLaboratorio de Sistemática y Biología Evolutiva, Museo de La Plata, La Plata B1900FWA, Argentina; dCentro Austral de Investigaciones Científicas, Consejo Nacional de Investigaciones Cientificas y Técnicas, 9410 Ushuaia, Tierra del Fuego, Argentina; and eDepartment of Palaeontology, GNS Science, Lower Hutt 5040, New Zealand Edited by Michael J. Donoghue, Yale University, New Haven, CT, and approved July 15, 2015 (received for review December 10, 2014) The Asteraceae (sunflowers and daisies) are the most diverse Here we report fossil pollen evidence from exposed Campanian/ family of flowering plants. Despite their prominent role in extant Maastrichtian sediments from the Antarctic Peninsula (Fig. 1, Fig. S1, terrestrial ecosystems, the early evolutionary history of this family and SI Materials and Methods, Fossiliferous Localities)(7)thatradi- remains poorly understood. Here we report the discovery of a cally changes our understanding of the early evolution of Asteraceae. number of fossil pollen grains preserved in dinosaur-bearing deposits from the Late Cretaceous of Antarctica that drastically pushes back Results and Discussion the timing of assumed origin of the family. Reliably dated to ∼76–66 The pollen grains reported here and discovered in the Late Cre- Mya, these specimens are about 20 million years older than previ- taceous of Antarctica are tricolporate, microechinate, with long ously known records for the Asteraceae. -
BIGHEAD KNAPWEED (Centaurea Macrocephala Puschk.)
PNW386 BIGHEAD KNAPWEED (Centaurea macrocephala Puschk.) Bighead knapweed is native to woody crown, each topped by a the mountain grasslands of 3-inch flower head. When Caucasia (specifically Armenia competing with other plants, and Romania) and also to bighead knapweed is generally subalpine meadows in Turkey. only 2 to 3 feet tall with one or European growers cultivate it as two stalks bearing flower heads an ornamental flower and sell it about 1.5 inches in diameter. in the fresh flower markets. Broadly lance-shaped leaves have Although not as popular as sharp-pointed tips, shallowly cornflower (bachelor's button) or toothed edges and rough mountain bluet, it is also surfaces. Blades of the stalked available in the United States in basal and rosette leaves may be flower seed catalogs and 10 inches long and 3 inches wide; nurseries under various common blade and stalk together may names, including Lemon fluff exceed 15 inches. Stem leaves and Globe Centaury. gradually change from having Occasionally, it appears in dried simple stalks to winged stalks to flower arrangements. stalkless as they become smaller Weed specialists in Washington going up the stem. The smallest found bighead knapweed in the Under frroorable conditions, bighead leaves are clustered on the early 1980s escaping from knapweed grows 4 to 5 feet tall. (Each swollen stem at the base of the abandoned gardens in Pend stripe on the stake is 4 inches.) flower head. Oreille and Whitman counties, and added it to the Class A noxious weed list. Since then, Okanogan County, Washington, and Quesnel, British Columbia have reported it. -
State of Colorado 2016 Wetland Plant List
5/12/16 State of Colorado 2016 Wetland Plant List Lichvar, R.W., D.L. Banks, W.N. Kirchner, and N.C. Melvin. 2016. The National Wetland Plant List: 2016 wetland ratings. Phytoneuron 2016-30: 1-17. Published 28 April 2016. ISSN 2153 733X http://wetland-plants.usace.army.mil/ Aquilegia caerulea James (Colorado Blue Columbine) Photo: William Gray List Counts: Wetland AW GP WMVC Total UPL 83 120 101 304 FACU 440 393 430 1263 FAC 333 292 355 980 FACW 342 329 333 1004 OBL 279 285 285 849 Rating 1477 1419 1504 1511 User Notes: 1) Plant species not listed are considered UPL for wetland delineation purposes. 2) A few UPL species are listed because they are rated FACU or wetter in at least one Corps Region. 3) Some state boundaries lie within two or more Corps Regions. If a species occurs in one region but not the other, its rating will be shown in one column and the other column will be BLANK. Approved for public release; distribution is unlimited. 1/22 5/12/16 Scientific Name Authorship AW GP WMVC Common Name Abies bifolia A. Murr. FACU FACU Rocky Mountain Alpine Fir Abutilon theophrasti Medik. UPL UPL FACU Velvetleaf Acalypha rhomboidea Raf. FACU FACU Common Three-Seed-Mercury Acer glabrum Torr. FAC FAC FACU Rocky Mountain Maple Acer grandidentatum Nutt. FACU FAC FACU Canyon Maple Acer negundo L. FACW FAC FAC Ash-Leaf Maple Acer platanoides L. UPL UPL FACU Norw ay Maple Acer saccharinum L. FAC FAC FAC Silver Maple Achillea millefolium L. FACU FACU FACU Common Yarrow Achillea ptarmica L. -
Invasive Asteraceae Copy.Indd
Family Asteraceae Family: Asteraceae Spotted Knapweed Centaurea biebersteinii DC. Synonyms Acosta maculosa auct. non Holub, Centaurea maculosa auct. non Lam. Related Species Russian Knapweed Acroptilon repens (L.) DC. Description Spotted knapweed is a biennial to short-lived perennial plant. Seedling cotyledons are ovate, with the first leaves lance-shaped, undivided, and hairless. (Young seedlings can appear grass-like.) Stems grow 1 to 4 feet tall, and are many-branched, with a single flower at the end of each branch. Rosette leaves are indented or divided Old XID Services photo by Richard about half-way to the midrib. Stem leaves are alternate, pinnately divided, Spotted knapweed flower. and get increasingly smaller toward the tip of each branch. Flower heads are urn-shaped, up to 1 inch wide, and composed of pink, purple, or sometimes white disk flowers. A key characteristic of spotted knap- weed is the dark comb-like fringe on the tips of the bracts, found just below the flower petals. These dark-tipped bracts give this plant its “spotted” appearance. Russian knapweed is a creeping perennial plant that is extensively branched, with solitary urn-shaped pink or purple flower heads at the end of each branch. Similar in appearance to spotted knapweed, Russian knapweed can be distinguished by its slightly smaller flower heads, flower head bracts covered in light hairs, with papery tips, and scaly dark brown or black rhizomes, which have a burnt appearance. Family: Asteraceae Spotted Knapweed Leaves and stems of both spotted and Russian knapweeds are covered in fine hairs, giving the plants a grayish cast. -
A Biographical Index of British and Irish Botanists
L Biographical Index of British and Irish Botanists. TTTEN & BOULGER, A BIOaEAPHICAL INDEX OF BKITISH AND IRISH BOTANISTS. BIOGRAPHICAL INDEX OF BRITISH AND IRISH BOTANISTS COMPILED BY JAMES BEITTEN, F.L.S. SENIOR ASSISTANT, DEPARTMENT OF BOTANY, BBITISH MUSEUM AKD G. S. BOULGEE, E.L. S., F. G. S. PROFESSOR OF BOTANY, CITY OF LONDON COLLEGE LONDON WEST, NEWMAN & CO 54 HATTON GARDEN 1893 LONDON PRINTED BY WEST, NEWMAN AND HATTON GAEDEN PEEFACE. A FEW words of explanation as to the object and scope of this Index may fitly appear as an introduction to the work. It is intended mainly as a guide to further information, and not as a bibliography or biography. We have been liberal in including all who have in any way contributed to the literature of Botany, who have made scientific collections of plants, or have otherwise assisted directly in the progress of Botany, exclusive of pure Horticulture. We have not, as a rule, included those who were merely patrons of workers, or those known only as contributing small details to a local Flora. Where known, the name is followed by the years of birth and death, which, when uncertain, are marked with a ? or c. [circa) ; or merely approximate dates of "flourishing" are given. Then follows the place and day of bu'th and death, and the place of burial ; a brief indication of social position or occupation, espe- cially in the cases of artisan botanists and of professional collectors; chief university degrees, or other titles or offices held, and dates of election to the Linnean and Eoyal Societies. -
Alphabetical Lists of the Vascular Plant Families with Their Phylogenetic
Colligo 2 (1) : 3-10 BOTANIQUE Alphabetical lists of the vascular plant families with their phylogenetic classification numbers Listes alphabétiques des familles de plantes vasculaires avec leurs numéros de classement phylogénétique FRÉDÉRIC DANET* *Mairie de Lyon, Espaces verts, Jardin botanique, Herbier, 69205 Lyon cedex 01, France - [email protected] Citation : Danet F., 2019. Alphabetical lists of the vascular plant families with their phylogenetic classification numbers. Colligo, 2(1) : 3- 10. https://perma.cc/2WFD-A2A7 KEY-WORDS Angiosperms family arrangement Summary: This paper provides, for herbarium cura- Gymnosperms Classification tors, the alphabetical lists of the recognized families Pteridophytes APG system in pteridophytes, gymnosperms and angiosperms Ferns PPG system with their phylogenetic classification numbers. Lycophytes phylogeny Herbarium MOTS-CLÉS Angiospermes rangement des familles Résumé : Cet article produit, pour les conservateurs Gymnospermes Classification d’herbier, les listes alphabétiques des familles recon- Ptéridophytes système APG nues pour les ptéridophytes, les gymnospermes et Fougères système PPG les angiospermes avec leurs numéros de classement Lycophytes phylogénie phylogénétique. Herbier Introduction These alphabetical lists have been established for the systems of A.-L de Jussieu, A.-P. de Can- The organization of herbarium collections con- dolle, Bentham & Hooker, etc. that are still used sists in arranging the specimens logically to in the management of historical herbaria find and reclassify them easily in the appro- whose original classification is voluntarily pre- priate storage units. In the vascular plant col- served. lections, commonly used methods are systema- Recent classification systems based on molecu- tic classification, alphabetical classification, or lar phylogenies have developed, and herbaria combinations of both. -
Anaphalis Margaritacea (L) Benth
Growing and Using Native Plants in the Northern Interior of B.C. Anaphalis margaritacea (L) Benth. and Hook. F. ex C.B. Clarke pearly everlasting Family: Asteraceae Figure 79. Documented range of Anaphalis margaritacea in northern British Columbia. Figure 80. Growth habit of Anaphalis margaritacea in cultivation. Symbios Research & Restoration 2003 111 Growing and Using Native Plants in the Northern Interior of B.C. Anaphalis margaritacea pearly everlasting (continued) Background Information Anaphalis margaritacea can be found north to Alaska, the Yukon and Northwest Territories, east to Newfoundland and Nova Scotia, and south to North Carolina, Kentucky, Arizona, New Mexico and California. It is reported to be common throughout B.C. except in the northeast (Douglas et al. 1998). Growth Form: Rhizomatous perennial herb, with few basal leaves, alternate stem leaves light green above, woolly white underneath; flower heads in dense flat-topped clusters, yellowish disk flowers; involucral bracts dry pearly white; mature plant size is 20-90 cm tall (MacKinnon et al. 1992, Douglas 1998). Site Preferences: Moist to dry meadows, rocky slopes, open forest, landings, roadsides and other disturbed sites from low to subalpine elevations, throughout most of B.C. In coastal B.C., it is reported to be shade-intolerant and occupies exposed mineral soil on disturbed sites and water- shedding sites up to the alpine (Klinka et al. 1989). Seed Information Seed Size: Length: 0.97 mm (0.85 - 1.07 mm). Width : 0.32 mm (0.24 - 0.37 mm). Seeds per gram: 24,254 (range: 13,375 - 37,167). Volume to Weight Conversion: 374.0 g/L at 66.7.5% purity. -
1 Sex Ratio and Spatial Distribution of Male and Female Antennaria Dioica
Sex ratio and spatial distribution of male and female Antennaria dioica (Asteraceae) plants Sandra Varga* and Minna-Maarit Kytöviita Department of Biological and Environmental Science, University of Jyväskylä, P.O. Box 35, FIN- 40014 Jyväskylä, Finland E-mail addresses: [email protected] and [email protected] (S. Varga), minna- [email protected] (M.-M. Kytöviita) *Author for correspondence: Sandra Varga Department of Biological and Environmental Science University of Jyväskylä FIN-40014 Jyväskylä Finland Email: [email protected] and [email protected] Phone: +358 14 260 2304 Fax: +358 14 260 2321 1 Abstract Sex ratio, sex spatial distribution and sexual dimorphism in reproduction and arbuscular mycorrhizal colonisation were investigated in the dioecious clonal plant Antennaria dioica (Asteraceae). Plants were monitored for five consecutive years in six study plots in Oulanka, northern Finland. Sex ratio, spatial distribution of sexes, flowering frequency, number of floral shoots and the number and weight of inflorescences were recorded. In addition, intensity of mycorrhizal fungi in the roots was assessed. Both sexes flowered each year with a similar frequency, but the overall genet sex ratio was strongly female-biased. The bivariate Ripley’s analysis of the sex distribution showed that within most plots sexes were randomly distributed except for one plot. Sexual dimorphism was expressed as larger floral and inflorescence production and heavier inflorescences in males. In addition, the roots of both sexes were colonised to a similar extent by arbuscular mycorrhizal fungi. The female sex-biased flowering ratios reported are not consistent among years and cannot be explained in terms of spatial segregation of the sexes or sex lability. -
Fdn22 Northern Dry-Bedrock Pine (Oak) Woodland *Spinulose Shield Fern Or Glandular Wood Fern ( Dryopteris Carthusiana Or D
FIRE-DEPENDENT FOREST/WOODLAND SYSTEM FDn22 Northern Floristic Region Northern Dry-Bedrock Pine (Oak) Woodland Dry pine or oak woodlands on shallow, excessively drained, loamy soils on bedrock ridges and hillsides or on rock ledges and terraces adjacent to rivers. Crown and surface fires were common historically. Vegetation Structure & Composition Description is based on summary of vegetation data from 47 plots (relevés). Ground-layer cover of forbs and grami- noids typically ranges from 25-75%. The most common vascular plants are Canada mayflower (Maianthemum canadense), wild sarsaparilla (Aralia nudicaulis), large-leaved aster (Aster macrophyllus), poverty grass (Danthonia spicata), wintergreen (Gaultheria procumbens), and bracken (Pteridium aqui- linum). Lichen- and moss-covered bedrock and boulders typically make up at least 25% of the ground layer. Shrub layer is typically dominated by deciduous species, usually with patchy to interrupted cover (25-75%). Lowbush blue- berry (Vaccinium angustifolium), juneberries (Amelanchier spp.), red maple saplings, and bush honeysuckle (Diervilla lonicera) are the most common species in the shrub layer. Subcanopy is usually absent, but when present, red maple and paper birch are frequent components. Canopy is composed of conifers, hardwoods, or conifers mixed with hardwoods, and is usually patchy (25-50% cover), with openings in areas of exposed bedrock or boulders. Red pine and white pine are dominant on many sites. On other sites, jack pine or northern pin oak are dominant. In mixed forests, conifers often form a supercanopy above hardwood species. Paper birch is often present in the hardwood canopy. Landscape Setting & Soils Glacially scoured bedrock—Common. Landscape is hummocky to rugged. Parent material is non-calcareous drift, usually less than 20in (50cm) deep over bedrock. -
Wildflowers of Twin Pillars North Trail, Bingham Springs, Trail Station
Wildflowers of Twin Pillars North Trail, Bingham Springs, Trail Station Meadows & upper Desolation Canyon Mill Creek Wilderness & Ochoco National Forest Data from personal observations & Data Provided by the Oregon Flora Project & Consortium of Pacific Northwest Herbaria Taxa with a question mark have been found within 10 miles of this site but not yet documented onsite. Last updated May 28, 2018 Common Name Scientific Name Family ____ Black Elderberry Sambucus racemosa v. melanocarpa Adoxaceae ____ Bulbil Onion Allium geyeri v. tenerum Amaryllidaceae ____ Tolmie's Onion Allium tolmiei v. tolmiei Amaryllidaceae ____ Gray’s Lovage Ligusticum grayi Apiaceae ____ Slender-fruited Lomatium Lomatium leptocarpum Apiaceae ____ Gray's Desert Parsley Lomatium papilioniferum ? Apiaceae ____ Common Sweet Cicely Osmorhiza berteroi Apiaceae ____ Western Sweet Cicely Osmorhiza occidentalis Apiaceae ____ Gairdner’s Yampah Perideridia gairdneri Apiaceae ____ Sierra Snakeroot Sanicula graveolens Apiaceae ____ Mountain Dogbane Apocynum androsaemifolium Apocynaceae ____ False Solomon Seal Maianthemum racemosum ssp. amplexicauleAsparagaceae ____ Star-flowered False Solomon SealMaianthemum stellatum Asparagaceae ____ Hyacinth Cluster-lily Triteleia hyacinthina ? Asparagaceae ____ Yarrow Achillea millefolium Asteraceae ____ Pale Agoseris Agoseris glauca v. glauca Asteraceae ____ Sagebrush Agoseris Agoseris parviflora Asteraceae ____ Pearly Everlasting Anaphalis margaritacea Asteraceae ____ Pussytoes Antenaria sp. Asteraceae ____ Woodrush Pussytoes Antennaria -
Full of Beans: a Study on the Alignment of Two Flowering Plants Classification Systems
Full of beans: a study on the alignment of two flowering plants classification systems Yi-Yun Cheng and Bertram Ludäscher School of Information Sciences, University of Illinois at Urbana-Champaign, USA {yiyunyc2,ludaesch}@illinois.edu Abstract. Advancements in technologies such as DNA analysis have given rise to new ways in organizing organisms in biodiversity classification systems. In this paper, we examine the feasibility of aligning two classification systems for flowering plants using a logic-based, Region Connection Calculus (RCC-5) ap- proach. The older “Cronquist system” (1981) classifies plants using their mor- phological features, while the more recent Angiosperm Phylogeny Group IV (APG IV) (2016) system classifies based on many new methods including ge- nome-level analysis. In our approach, we align pairwise concepts X and Y from two taxonomies using five basic set relations: congruence (X=Y), inclusion (X>Y), inverse inclusion (X<Y), overlap (X><Y), and disjointness (X!Y). With some of the RCC-5 relationships among the Fabaceae family (beans family) and the Sapindaceae family (maple family) uncertain, we anticipate that the merging of the two classification systems will lead to numerous merged solutions, so- called possible worlds. Our research demonstrates how logic-based alignment with ambiguities can lead to multiple merged solutions, which would not have been feasible when aligning taxonomies, classifications, or other knowledge or- ganization systems (KOS) manually. We believe that this work can introduce a novel approach for aligning KOS, where merged possible worlds can serve as a minimum viable product for engaging domain experts in the loop. Keywords: taxonomy alignment, KOS alignment, interoperability 1 Introduction With the advent of large-scale technologies and datasets, it has become increasingly difficult to organize information using a stable unitary classification scheme over time.