Volume 29: 14–22 METAMORPHOSIS ISSN 1018–6490 (PRINT) LEPIDOPTERISTS’ SOCIETY OF AFRICA ISSN 2307–5031 (ONLINE)

The (: Papilionoidea) of Semuliki National Park, western Uganda Published online: 21 May 2018

Scott Forbes 33 Hope Terrace, Edinburgh, EH9 2AP, Scotland, U.K. E-mail: [email protected]

Copyright © Lepidopterists’ Society of Africa

Abstract: A checklist of the butterflies (Lepidoptera: Papilionoidea) of Semuliki National Park, Uganda is provided. The list was compiled from records made by the author as well as from historical records and the records of a number of field workers. A total of 448 species are listed for the Park.

Key words: Lepidoptera, Papilionoidea, butterflies, Semuliki National Park, Uganda.

Citation: Forbes, S. (2018). The butterflies (Lepidoptera: Papilionidae) of Semuliki National Park, western Uganda. Metamorphosis 29: 14–22.

INTRODUCTION

Semuliki National Park (SNP), with an area of 219 km2, and lying at Lat. -1.455, Long. 31.152 (DD), is Uganda’s only remaining lowland forest. The conservation status of the Park includes being located within the Eastern Afromontane hotspot region within the Albertine Rift, the western arm of the Rift Valley. (Fig. 1). It has also been ranked in a global irreplaceability list (LeSaout et al., 2013) and as a Key Biodiversity Area (IUCN, 2015).

Much of the park is flat, lying at an altitude of 670 m in the east, rising to undulating sections of 760 m. Only 10 km2 lies above 750 m, at its western boundary. The Park is situated within the Semliki Valley and is bounded to the north-northwest by the steep escarpment of the Rwenzori Mountain Range, which isolates the Park from the rest of East Africa. To the east is a demarcated boundary where the rainforest is supplanted by the open grasslands of the Semliki Valley, which stretches down to the shores of Lake Albert. The northern boundary is the Semliki River, the international boundary between Uganda and the Democratic Republic of Congo (DRC) and to the west the Lamia River borders the Park. The Ituri Forest of the DRC lies along the eastern margin of the Congo Basin, and SNP is the easternmost extension of this typical -Congolean forest habitat. It makes up one of a few of the eastern forest fragments of the Figure 1 - The Albertine Rift (pale shading), with Semuliki Congo Basin forest system, which totals 54% of National Park (red; arrowed). Map from Protected Area Watch Africa’s tropical forests (Mayaux et al., 2013). for the Albertine Rift Project (PAWAR), 2016.

This lowland forest is classified as moist, semi- identified as the Cynometra-Celtis zone, one of four deciduous (Langdale-Brown et al., 1964) and, being different types of lowland forest types described by Hamilton (1981) (Table 1). An area of approximately 30% is dominated by a single leguminous tree, Received: 28 February 2018 Published: 21 May 2018 Cynometra alexandri. Much of the Park overlies grey alluvial clay soils that tend to be saline, have reduced Copyright: This work is licensed under the Creative Commons fertility, and are poorly drained (Howard, 1991). Attribution-NonCommercial-NoDerivs 3.0 Unported License. To view a copy of this license, send a letter to Creative These give rise to swamp communities, which Commons, Second Street, Suite 300, San Francisco, California, 94105, USA, or visit: http://creative commons.org/licenses/by- contribute to around 7% of the total area. It is probably nc-nd/3.0/

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Table 1 – Summary of salient characteristics of Semuliki represented by large differences in tree species-area National Park. relationships at sites within close proximity (Howard, 1991). Characteristic Description Co-ordinates Lat. -1.455, Long. 31.152 (DD) MATERIALS AND METHODS Designation Gazetted as National Park in 1993; full status in

1994 Biogeography Afrotropical region, Uganda zone Sampling Forest Single-dominant, moist-evergreen and semi- Classification evergreen Guineo-Congolian lowland rainforest Due to permit restrictions implemented for Ugandan Location Semliki Valley, north-northwest of the Rwenzori National Parks, it is difficult to remove material for mountains in the Albertine Rift. Sharing a 1 km external identification, which is sometimes necessary for boundary with the North Rwenzori Forest Reserve and contiguous with a 10 km boundary these . The fruit-feeding assemblage, of the Parc National des Virunga in the DRC. which includes the Nymphalidae genera Bebearia, Bordered by the Lamia and Semliki Rivers to the Euphaedra, Cymothoe, Bicyclus and Charaxes have been west and north; by the Fort Portal to Bundibugyo sampled consistently over a period of years (2012 to road to the south and community agricultural land to the east 2017), but predominantly in only two transects 500 m in Biodiversity Eastern Afromontane Division length around 7 kms apart (Kirumya and Sempaya trails). Hotspot Genitalic dissections were retained in glycerol vials Elevation Medium altitude, 670 to 760 m; <10 km2 >750 pinned under the corresponding specimens. m Soils Rift valley sediments, grey halomorphic alluvial Source of other records and data clay soils Recent History Western sections of the Park were used for Records from Davenport & Howard (1996) have been cultivation in the 1970’s and 1980’s. Encroachers removed in the late 1980’s. merged with records by Forbes for the period from 2013 Roughly 25% is now considered undisturbed to 2017. The latter records were gathered during research Climate Two seasons. Dry from December to February for a completed PhD entitled ‘The ecology and and a long diffuse wet season with two peaks, conservation of a fruit-feeding butterfly assemblage in a from April to May and October to November Ugandan lowland rainforest’, together with other records Rivers Drained by Ntotoro, Rugo, Mampongi and Kirumya rivers provided by D. Bernaud, R. Ducarme, J-P. Lequeux, and Precipitation Approximately 1350 mm – 1500 mm annually H. Selb. A maximum of two citations per species record Temperature 15°C to 25°C are given even though, in some instances, more records Current Forest Monthly firewood collection by women from the exist. As further records are identified or recorded in the Use surrounding communities – can be as many as field, updated versions of this checklist will be uploaded 700 individuals. Unrestricted access for the Batwa, the local indigenous community, for to the SNP butterflies Facebook page, with images being firewood collection, medicinal plant harvesting uploaded to the SNP butterflies Instagram page. and beehive access. Regular security patrols. Small outpost constructed by fishermen by the Museum collections consulted were: Museum of Central Semliki River Africa, Tervuren, Belgium; National Museum of Management Periodic debarking of invasive and exotic trees. Scotland, Edinburgh (NMS) and Glasgow University Continual maintenance of KR trail specifically used for birders, quarterly maintenance of RM Museum, Glasgow, Scotland. National Museums of trail. Biannual grassland burning in December (NMK) citations were provided by Mr Steve and January at the start of the dry season Collins of the African Butterfly Research Institute Threats Community encroachment on the western (ABRI), Nairobi, Kenya. border. Poaching for bush meat and illegal fishing. River pollution from production runoff entering the water course Taxonomy running into the Park Afrotropical butterfly taxonomy, to generic level, follows Williams (2015), Dhungel & Wahlberg (2018) and part of the major Pleistocene (2 500 000 – 12 000 yrs BP) Espeland et al. (2018). forest refugium from the eastern DRC (Hamilton, 1981), which has resulted in large species diversity of many RESULTS AND CONCLUSIONS communities characterised by typical Guineo-Congolian fauna, with many species at the eastern limit of their A total of 448 species have been recorded from SNP to ranges. In addition to differences in forest type associated date (Table 2). This is an increase of 142 species since with altitudinal and climatic factors, historical uses of the the last official census by Davenport & Howard (1996), Park have created a mosaic of climax vegetation types. who recorded 309 species. The current total is These were primarily created from the early 1970’s when significantly short of the over 600 species that an African viable forest sections of the Park, to the east of the equatorial lowland forest would be predicted to contain Kirumya River, were destroyed by agricultural (S. Collins, pers. comm.). Using the Davenport (2001) encroachment to provide for community agricultural land figure as a midpoint for total butterfly species in Uganda for subsistence farming. The last of these cultivators were (1 251), the SNP contains just over 36% of Ugandan finally evicted in the mid 1980’s. Thus, considerable butterfly species. local variation related to site factors, such as past management, soil type and drainage conditions are The butterfly families that require further sampling are

Forbes/ Metamorphosis 29: 14–22 16 the and Hesperiidae, which should contribute and dated phylogenomic analysis of butterflies. at least 40% of the total number of species of an Current Biology 28: 1–9. Afrotropical lowland forest (S. Collins, pers. comm.). HAMILTON, A. C. 1981. The quarternary history of Currently, the combined species total of these two African forests: its relevance to conservation. African families contributes just below 30% of the total but the Journal of Ecology 19: 1–6. Lycaenidae and Hesperiidae of the lowland forests in the HOWARD, P. 1991. Nature conservation in Uganda’s DRC (Ducarme, 2018) should also be found in the SNP tropical forest reserves. IUCN, Gland, Switzerland. since they share a similar habitat and altitude. IUCN. 2015. Global Standard for the Identification of Key Biodiversity Areas KBA’s. Gland, Switzerland. ACKNOWLEDGEMENTS LANGDALE-BROWN, I., OSMASTON, H.A. & WILSON, J.G. 1964. The Vegetation of Uganda and I thank Justice Olibokiriho at SNP for continuing to its bearing on land-use. Uganda Government Printer, sample and identify new records, and for his assistance in Entebbe, Uganda. the field. Joachim Mutaibisa and Martha Komuhangi LEQUEUX, J-P. 2016. Semuliki National Park – were also both instrumental in providing excellent field Papilionidae and Charaxinae records (unpublished). assistance and ongoing sampling. LESAOUT, S., HOFFMANN, M., SHI, Y., HUGHES, A., BERNARD, C., BROOKS, T.M., BERTZKY, B., Many thanks to the following collaborators for help and BUTCHART, S.H.M., STUART, S.N., BADMAN, guidance in compiling this species list: T. & RODRIGUES A.S.L. 2013. Protected areas and

Mr Dominique Bernaud – Subfamily Heliconiinae effective biodiversity conservation. Science 342: Mr Röbert Ducarme – NE DRC, Semliki Valley, 803–805. Albertine Rift, Congo Basin, Blue and Mitumba MAYAUX, P., PEKEL, J.F., DESCLÉE, B., DONNAY, Mountains and southern North Kivu distributional F., LUPI, A., ACHARD F., CLERICI, M., BODART, records C., BRINK, A., NASI, R. & BELWARD, A. 2013. Mr Jean-Pierre Lequeux – Family Papilionidae and State and evolution of the African rainforests between Subfamily Charaxinae 1990 and 2010. Philosophical Transactions of the Mr Haydon Warren-Gash – Family Pieridae Royal Society of London 368: 20120300. Mr Harald Selb – Family Lycaenidae SELB, H.E. 2014. Semuliki National Park – Uganda, Field Survey (unpublished). Mr Steve Collins (ABRI) and Dr Keith Bland of the WILLIAMS, M.C. 2015. Classification of the National Museum of Scotland NMS reviewed the final Afrotropical butterflies to generic level. draft of this document. Their input is valued and much Metamorphosis 26: 102–108. appreciated.

LITERATURE CITED

BERNAUD, D. 2016. Semuliki National Park, Uganda – provisional checklist of Acraeinae (unpublished). DAVENPORT, T.R.B. 2001. The butterflies of Uganda: an annotated catalogue. Wildlife Conservation Society, . DAVENPORT, T.R.B. & HOWARD, P. 1996. Semliki National Park, Biodiversity Report. Uganda Forest Department. DHUNGEL, B. & WAHLBERG, N. 2018. Molecular systematics of the subfamily Limenitidinae (Lepidoptera: Nymphalidae). PeerJ 6:e4311; DOI 10.7717/peerj.4311. DUCARME, R. 2017. Relevé faunistique des lépidoptères Rhopalocères du Nord Est de la Rèpublique Dèmocratique du Congo par milieu naturel (unpublished). DUCARME, R. 2018. Checklist of the butterflies (Lepidoptera: Papilionoidia) of the Democratic Republic of the Congo. Metamorphosis 29 [in press] ESPELAND, M., BREINHOLT, J., WILLMOTT, K.R., WARREN, A.D., VILA, R., TOUSSAINT, E.F.A., MAUNSELL, S.C., ADUSE-POKU, K., TALAVERA, G., EASTWOOD, R., JARZYNA, M.A., GURALNICK, R., LOHMAN, D.J., PIERCE, N.E. & KAWAHARA, A.Y. 2018. A comprehensive

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Table 2 – Checklist of the butterflies of Semuliki National Park, Osmodes Holland, 1892 Western Uganda. O. laronia (Hewitson, 1868) Forbes/Davenport O. thora (Plötz, 1884) Davenport Genus Acleros Mabille, 1885 TAXON COLLECTOR(S)/ A. neavei Evans, 1937 Davenport FAMILY PAPILIONIDAE Latreille, A. ploetzi Mabille, 1889 Davenport 1802 Genus Hypoleucis Mabille, 1891 Subfamily Papilioninae Latreille, 1802 H. tripunctata draga Evans, 1937 Davenport Tribe Leptocircini Kirby, 1896 Genus Meza Hemming, 1939 Genus Graphium Scopoli, 1777 M. cybeutes pallida (Evans, 1937) Davenport G. angolanus baronis (Ungemach, 1932) Davenport Genus Andronymus Holland, 1896 G. leonidas leonidas (Fabricius, 1793) Forbes/Davenport A. neander neander (Plötz, 1884) Davenport G. policenes policenes (Cramer, 1775) Davenport Genus Zophopetes Mabille, 1904 G. antheus (Cramer, 1779) Forbes/Davenport Z. dysmephila (Trimen, 1868) Davenport Tribe Papilionini Latreille, 1802 Z. nobilior (Holland, 1896) Davenport Genus Papilio Linnaeus, 1758 Genus Gamia Holland, 1896 P. mackinnoni mackinnoni Sharpe, 1891 Forbes/Davenport G. shelleyi (Sharpe, 1890) Davenport P. charopus montuosus Joicey & Talbot, Davenport Genus Gretna Evans, 1937 1927 G. cylinda (Hewitson, 1876) Davenport P. sosia debilis Storace, 1951 Forbes/Davenport Genus Caenides Holland, 1896 P. chrapkowskoides chrapkowskoides Forbes/Davenport C. dacela (Hewitson, 1876) Davenport Storace, 1952 Genus Monza Evans, 1937 P. nireus nireus Linnaeus, 1758 Forbes/Davenport M. punctata crola Evans, 1937 Davenport P. cynorta Fabricius, 1793 Forbes/Davenport Tribe Baorini Doherty, 1886 P. dardanus dardanus Brown, 1776 Forbes/Davenport Genus Borbo Evans, 1949 P. phorcas congoanus Rothschild, 1896 Forbes/Davenport B. fallax (Gaede, 1916) Forbes/Davenport P. nobilis crippsianus Stoneham, 1936 Forbes/Davenport Total for the subfamily Hesperiinae 24 P. hesperus hesperus Westwood, 1843 Forbes Total for the family Hesperiidae 34 P. lormieri semlikana Le Cerf, 1924 Davenport FAMILY NYMPHALIDAE P. demodocus demodocus Esper, 1798 Forbes/Davenport Rafinesque, 1815 P. mechowi mechowi Dewitz, 1881 Forbes/Davenport Subfamily Danainae Boisduval, 1833 P. zenobia Fabricius, 1775 Forbes/Davenport Tribe Danaini Boisduval, 1833 Total for the family Papilionidae 18 Subtribe Danaina Boisduval, 1833 FAMILY HESPERIIDAE Latreille, Genus Danaus Kluk, 1780 1809 D. chrysippus chrysippus Linnaeus, 1758) Forbes/Davenport Subfamily Coeliadinae Evans, 1937 Genus Tirumala Moore, 1880 Genus Coeliades Hübner, 1818 T. petiverana (Doubleday, 1847) Forbes/Davenport C. chalybe chalybe (Westwood, 1852) Davenport T. formosa mercedonia (Karsch, 1894) Forbes/Davenport C. forestan forestan (Stoll, 1782) Forbes/Davenport Subtribe Amaurina Le Cerf, 1922 C. libeon (Druce, 1875) Forbes/Davenport Genus Amauris Hübner, 1816 Total for the subfamily Coeliadinae 3 A. niavius niavius (Linnaeus, 1758) Forbes/Davenport Subfamily Pyrginae Burmeister, 1878 A. tartarea tartarea Mabille, 1876 Forbes/Davenport Tribe Tagiadini Mabille, 1878 A. hecate hecate (Butler, 1866) Forbes/Davenport Genus Eagris Guenée, 1862 A. albimaculata magnimacula Rebel, Davenport E. lucetia (Hewitson, 1875) Forbes/Davenport 1914 E. decastigma purpura Evans, 1937 Davenport A. crawshayi oscarus Thurau, 1904 Davenport Genus Calleagris Aurivillius, 1925 Total for the subfamily Danainae 8 C. hollandi (Butler, 1897) Davenport Subfamily Libytheinae Boisduval, 1833 C. lacteus lacteus (Mabille, 1877) Davenport Genus Libythea Fabricius, 1807 Genus Tagiades Hübner, 1819 L. labdaca Westwood, 1851 Forbes/Davenport T. flesus (Fabricius, 1781) Davenport Total for the subfamily Libytheinae 1 Tribe Celaenorrhinini Swinhoe, 1912 Subfamily Satyrinae Boisduval, 1833 Genus Sarangesa Moore, 1881 Tribe Elymniini Herrich-Schaeffer, 1864 S. lucidella lucidella (Mabille, 1891) Davenport Genus Elymnias Hübner, 1816 S. brigida sanaga Miller, 1964 Davenport E. bammakoo rattrayi Sharpe, 1902 Forbes/Davenport Total for the subfamily Pyrginae 7 Tribe Melanitini Reuter, 1896 Subfamily Hesperiinae Latreille, 1809 Genus Gnophodes Doubleday, 1849 Tribe incertae sedis G. betsimena parmeno Doubleday, 1849 Forbes/Davenport Genus Kedestes Watson, 1893 G. chelys (Fabricius, 1793) Forbes/Davenport K. rogersi Druce, 1907 Forbes Genus Melanitis Fabricius, 1807 Genus Teniorhinus Holland, 1892 M. leda (Linnaeus, 1758) Forbes/Davenport T. ignita (Mabille, 1877) Davenport M. libya Distant, 1882 Forbes T. watsoni Holland, 1892 Davenport M. ansorgei Rothschild, 1904 Forbes/Davenport Genus Ceratrichia Butler, 1870 Tribe Satyrini Boisduval, 1833 C. semlikensis Joicey & Talbot, 1921 Forbes/Davenport Subtribe Mycalesina Reuter, 1896 C. clara medea Evans, 1937 LepSoc Africa Genus Bicyclus Kirby, 1871 C. wollastoni Heron, 1909 Davenport B. alboplaga (Rebel, 1914) Forbes/Davenport Genus Ceratricula Larsen, 2013 B. angulosa angulosa (Butler, 1868) Forbes/Davenport C. semilutea congdoni Larsen, 2013 LepSoc Africa B. buea (Strand, 1912) Forbes/Davenport Genus Pardaleodes Butler, 1870 B. campina campina (Aurivillius, 1901) Forbes P. tibullus torensis Bethune-Baker, 1906 Forbes B. hyperanthus (Bethune-Baker, 1908) Forbes P. sator pusiella Mabille, 1877 Forbes/Davenport B. graueri graueri (Rebel, 1914) Forbes/Davenport P. incerta incerta (Snellen, 1872) Forbes/Davenport B. sebetus (Hewitson, 1877) Forbes/Davenport

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B. mesogena ugandae (Riley, 1926) Forbes C. pleione bebra Rothschild, 1900 Forbes/Davenport B. sambulos cyaneus Condamin, 1961 Forbes/Davenport C. paphianus subpallida Joicey & Talbot, Forbes/Davenport B. collinsi Aduse-Poku, Lees & Wahlberg, Forbes/Davenport 1925 2016 C. kahldeni Homeyer & Dewitz, 1882 Davenport B. auricruda fulgida Fox, 1963 Forbes/Davenport C. nichetes nichetes Grose-Smith, 1883 NMS B. vulgaris (Butler, 1868) Forbes/Davenport C. lycurgus bernardiana Plantrou, 1978 Forbes/Davenport B. jefferyi Fox, 1963 Forbes/Davenport C. zelica depuncta Joicey & Talbot, 1921 NMS B. sandace (Hewitson, 1877) Forbes/Davenport C. trajanus vansomereni (Poulton, 1929) Forbes B. sanaos (Hewitson, 1866) Forbes/Davenport C. eurinome lequeuxi (Turlin, 2009) Forbes/Davenport B. sophrosyne sophrosyne (Plötz, 1880) Forbes/Davenport C. crossleyi ansorgei (Rothschild, 1903) Forbes/Lequeux B. mollitia (Karsch, 1895) Forbes/Davenport C. eupale latimargo Joicey & Talbot, 1921 Forbes/Davenport B. smithi smithi (Aurivillius, 1899) Forbes/Davenport C. minor karinae Bouyer, 1999 Davenport/Selb B. golo (Aurivillius, 1893) Forbes/Selb Total for the subfamily Charaxinae 41 B. ignobilis (Butler, 1870) Forbes Subfamily Nymphalinae Rafinesque, B. xeneoides Condamin, 1961 Forbes 1815 B. safitza safitza (Hewitson, 1850) Forbes/Davenport Tribe Kallimini Doherty, 1886 B. campus (Karsch, 1893) Forbes/Davenport Genus Catacroptera Karsch, 1894 B. funebris (Guérin-Méneville, 1844) Forbes/Davenport C. cloanthe cloanthe (Stoll, 1781) Forbes B. procora (Karsch, 1893) Forbes/Davenport Tribe Junoniini Reuter, 1896 B. uniformis (Bethune-Baker, 1908) Forbes/Davenport Genus Junonia Hübner, 1819 Genus Brakefieldia Aduse-Poku, Lees & J. chorimene (Guérin-Méneville, 1844) Forbes/Davenport Wahlberg, 2016 J. gregorii Butler, 1896 Forbes B. phaea phaea (Karsch, 1894) Forbes J. terea tereoides (Butler, 1901) Forbes/Davenport B. peitho reducta (Libert, 2006) Forbes/Davenport J. sophia sophia (Fabricius, 1793) Forbes/Davenport Subtribe Ypthimina Reuter, 1896 J. westermanni suffusa (Rothschild & Davenport Genus Ypthima Hübner, 1818 Jordan, 1903) Y. asterope asterope (Klug, 1832) Forbes J. oenone oenone (Linnaeus, 1758) Forbes/Davenport Y. doleta Kirby, 1880 Forbes/Selb J. stygia (Aurivillius, 1894) Forbes/Davenport Y. albida albida Butler, 1888 Forbes/Davenport J. ansorgei (Rothschild, 1899) Forbes/Davenport Genus Ypthimomorpha van Son, 1955 Genus Salamis Boisduval, 1833 Y. itonia (Hewitson, 1865) Forbes/Selb S. cacta cacta (Fabricius, 1793) Forbes/Davenport Total for the subfamily Satyrinae 38 Genus Protogoniomorpha Wallengren, Subfamily Charaxinae Guenée, 1865 1857 Tribe Pallini Rydon, 1971 P. temora temora (Felder & Felder, 1867) Forbes/Davenport Genus Palla Hübner, 1819 P. parhassus (Drury, 1782) Forbes/Davenport P. violinitens bwamba van Someren, 1975 Forbes/Davenport P. anacardii nebulosa (Trimen, 1881) Forbes Tribe Charaxini Guenée, 1865 Genus Precis Hübner, 1819 Genus Charaxes Ochsenheimer, 1816 P. pelarga (Fabricius, 1775) Forbes/Davenport C. varanes vologeses (Mabille, 1876) Forbes/Davenport P. actia Distant, 1880 Selb C. fulvescens monitor Rothschild, 1900 Forbes/Davenport P. antilope (Feisthamel, 1850) Davenport C. acuminatus kigezia van Someren, 1963 Forbes P. tugela pyriformis (Butler, 1896) Forbes/Davenport C. candiope (Godart, 1824) Forbes/Davenport Genus Hypolimnas Hübner, 1819 C. protoclea nothodes Jordan, 1911 Forbes/Davenport H. misippus (Linnaeus, 1764) Forbes C. cynthia kinduana Le Cerf, 1923 Forbes/Davenport H. salmacis salmacis (Drury, 1773) Forbes/Davenport C. lucretius maximus van Someren, 1971 Forbes/Davenport H. monteironis major Rothschild, 1918 Forbes/Davenport C. epijasius Reiche, 1850 Forbes H. anthedon anthedon (Doubleday, 1845) Forbes/Davenport C. castor castor (Cramer, 1775) Davenport Tribe incertae sedis C. brutus angustus Rothschild, 1900 Forbes/Davenport Genus Kallimoides Shirôzu & C. pollux pollux (Cramer, 1775) Forbes/Davenport Nakanishi, 1984 C. eudoxus mechowi Rothschild, 1900 NMS K. rumia rattrayi (Shrpe, 1904) Forbes/Davenport C. numenes aequatorialis van Someren, Forbes/Davenport Genus Vanessula Dewitz, 1887 1972 V. milca latifascia Joicey & Talbot, 1928 Forbes C. tiridates tiridatinus Röber, 1936 Forbes/Davenport Total for the subfamily Nymphalinae 23 C. bipunctatus ugandensis van Someren, Forbes/Davenport Subfamily Cyrestinae Guenée, 1865 1972 Genus Cyrestis Boisduval, 1832 C. smaragdalis caerulea Jackson, 1951 Forbes C. camillus camillus (Fabricius, 1781) Forbes/Davenport C. imperialis ugandicus van Someren, Forbes/Davenport Total for the subfamily Cyrestinae 1 1972 Subfamily Biblidinae Boisduval, 1833 C. ameliae victoriae van Someren, 1972 Forbes/Davenport Tribe Biblidini Boisduval, 1833 C. pythodoris pythodoris Hewitson, 1873 Forbes Genus Byblia Hübner, 1819 C. hadrianus Ward, 1871 Forbes B. anvatara crameri Aurivillius, 1894 Forbes/Davenport C. zingha (Stoll, 1780) Forbes/Davenport B. ilithyia (Drury, 1773) Forbes/Davenport C. etesipe etesipe (Godart, 1824) Forbes/Davenport Genus Mesoxantha Aurivillius, 1899 C. anticlea adusta Rothschild, 1900 Forbes M. ethosea reducta Rothschild, 1918 Forbes/Davenport C. hildebrandti katangensis Talbot, 1928 NMS Genus Ariadne Horsfield, 1829 C. virilis lenis Henning, 1989 Forbes/Davenport A. enotrea suffusa (Joicey & Talbot, 1921) Forbes/Davenport C. catachrous van Someren & Jackson, Forbes/Davenport A. pagenstecheri (Suffert, 1904) Forbes/Davenport 1952 Genus Neptidopsis Aurivillius, 1899 C. etheocles carpenteri van Someren & Forbes/Davenport N. ophione nucleata Grünberg, 1911 Forbes/Davenport Jackson, 1957 Genus Eurytela Boisduval, 1833 C. cedreatis Hewitson, 1874 Forbes/Davenport E. hiarbas hiarbas (Drury, 1782) Forbes/Davenport C. viola picta van Someren & Jackson, Forbes/Davenport E. dryope angulata Aurivillius, 1899 Forbes/Davenport 1952 Tribe Epicaliini Guenée, 1865

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Genus Sevenia Koçak, 1996 P. eurytis eurytis (Doubleday, 1847) Forbes/Davenport S. occidentalium occidentalium (Mabille, Forbes/Davenport Tribe Argynnini Swainson, 1833 1876) Subtribe Argynnina Swainson, 1833 S. garega (Karsch, 1892) Forbes Genus Issoria Hübner, 1819 S. boisduvali omissa (Rothschild, 1918) Forbes I. baumanni excelsior (Butler, 1896) Forbes/Davenport Total for the subfamily Biblidinae 11 Total for the subfamily Heliconiinae 49 Subfamily Apaturinae Boisduval, 1840 Subfamily Limenitidinae Behr, 1864 Genus Apaturopsis Aurivillius, 1899 Tribe Pseudoneptini Dhungel & A. cleochares cleochares (Hewitson, Forbes Wahlberg, 2018 1873) Genus Pseudoneptis Snellen, 1882 Total for the subfamily Apaturinae 1 P. bugandensis ianthe Hemming, 1964 Forbes/Davenport Subfamily Heliconiinae Swainson, 1822 Tribe Cymothoini Dhungel & Wahlberg, Tribe Acraeini Boisduval, 1833 2018 Subtribe Acraeina Boisduval, 1833 Genus Cymothoe Hübner, 1819 Genus Acraea Fabricius, 1807 C. beckeri theodosia Staudinger, 1890 Forbes A. kraka Aurivillius, 1893 Davenport C. confusa Aurivillius, 1887 Forbes/Davenport A. rogersi lankesteri Carpenter, 1941 Lequeux/Bernaud C. cyclades (Ward, 1871) Forbes/Davenport A. egina egina (Cramer, 1775) Lequeux/Bernaud C. ochreata Grose Smith, 1890 Forbes/Davenport A. abdera abdera Hewitson, 1852 Forbes C. lurida tristis Overlaet, 1952 Forbes/Davenport A. cepheus (Linnaeus, 1758) Lequeux/Bernaud C. colmanti Aurivillius, 1898 Forbes A. asboloplintha asboloplintha Karsch, Davenport C. haynae diphyia Karsch, 1894 Forbes 1894 C. herminia johnstoni (Butler, 1902) Forbes A. zetes zetes (Linnaeus, 1758) Bernaud C. caenis (Drury, 1773) Forbes/Davenport A. leucographa leucographa Ribbe, 1889 Davenport C. coccinata vrydaghi Overlaet, 1944 Forbes/Davenport A. quirina quirina (Fabricius, 1781) Lequeux/Bernaud C. reginaeelisabethae reginaeelisabethae Forbes/Davenport A. epaea epaea (Cramer, 1779) Lequeux/Bernaud Holland, 1920 A. parageum parageum (Grose-Smith, Lequeux/Bernaud C. sangaris sangaris (Godart, 1824) Forbes/Davenport 1900) C. hobarti hobarti Butler, 1900 Forbes/Davenport A. schubotzi (Grünberg, 1911) Lequeux/Bernaud C. jodutta mostinckxi Overlaet, 1952 Forbes/Davenport A. ducarmei Bernaud & Pierre, 2012 Lequeux/Bernaud C. meridionalis ghesquierei Overlaet, Forbes A. poggei nelsoni Grose-Smith & Kirby, Bernaud 1944 1892 Genus Harma Doubleday, 1848 A. epiprotea (Butler, 1874) Lequeux/Bernaud H. theobene superna (Fox, 1968) Forbes/Davenport A. alcinoe camerunica (Aurivillius, 1893) Forbes/Davenport Tribe Neptini Newman, 1870 A. consanguinea intermedia (Aurivillius, Forbes Genus Neptis Fabricius, 1807 1899) N. saclava marpessa Hopffer, 1855 Forbes/Davenport A. pseuderyta Godman & Salvin, 1890 Lequeux/Bernaud N. conspicua Neave, 1904 Davenport A. macarista macarista (Sharpe, 1906) Lequeux/Bernaud N. constantiae constantiae Carcasson, Forbes A. umbra hemileuca (Jordan, 1914) Davenport 1961 Subtribe Actinotina Henning, 1992 N. metanira Holland, 1892 Selb Genus Telchinia Hübner, 1819 N. metella metella (Doubleday, 1850) Forbes/Davenport T. perenna perenna (Doubleday, 1847) Selb/Bernaud N. nysiades Hewitson, 1868 Forbes T. cinerea (Neave, 1904) NMS N. nemetes margueriteae Fox, 1968 Forbes/Davenport T. orinata (Oberthür, 1893) Davenport N. nicomedes Hewitson, 1874 Davenport T. penelope penelope (Staudinger, 1896) Bernaud N. strigata kakamega Collins & Larsen, Davenport/Selb T. quirinalis (Grose Smith, 1900) Lequeux/Bernaud 1996 T. parrhasia servona (Godart, 1819) Bernaud N. nicoteles Hewitson, 1874 Davenport T. peneleos pelasgia (Grose-Smith, 1900) Bernaud N. melicerta (Drury, 1773) Forbes/Davenport T. grosvenori (Eltringham, 1912) Davenport N. agouale parallela Collins & Larsen, Selb T. oreas oreas (Sharpe, 1891) NMS 1996 T. iturina (Grose-Smith, 1890) Bernaud N. lermanni Aurivillius, 1896 Davenport T. pharsalus pharsalus (Ward, 1871) Bernaud N. trigonophora intermedia Schultze, Forbes/Selb T. encedon encedon (Linnaeus, 1758) Davenport 1920 T. encedana (Pierre, 1976) Forbes N. kiriakoffi Overlaet, 1955 Selb T. jodutta jodutta (Fabricius, 1793) Bernaud N. serena serena Overlaet, 1955 Selb T. aurivillii aurivillii (Staudinger, 1896) Bernaud N. morosa Overlaet, 1955 Forbes T. alciopoides (Joicey & Talbot, 1921) Lequeux/Bernaud N. seeldrayersi Aurivillius, 1895 Davenport T. lycoa (Godart, 1819) Bernaud N. alta Overlaet, 1955 Selb T. serena (Fabricius, 1775) Davenport N. ochracea ochracea Neave, 1904 Davenport T. acerata (Hewitson, 1874) Bernaud N. continuata Holland, 1892 Davenport T. althoffi rubrofasciata (Aurivillius, Lequeux/Bernaud Tribe Adoliadini Doubleday, 1845 1895) Subtribe Bebearina Hemming, 1960 T. uvui uvui (Grose-Smith, 1890) Davenport Genus Evena Westwood, 1850 T. bonasia bonasia (Fabricius, 1775) Bernaud E. crithea (Drury, 1773) Forbes/Davenport T. sotikensis sotikensis (Sharpe, 1892) Davenport E. angustatum (Felder & Felder, 1867) Forbes/Davenport T. viviana (Staudinger, 1896) Davenport E. oberthueri (Karsch, 1894) LepSoc Africa T. orestia orestia (Hewitson, 1874) Lequeux/Bernaud Genus Aterica Boisduval, 1833 Tribe Vagrantini Pinratana & Eliot, 1996 A. galene extensa Heron, 1909 Forbes/Davenport Genus Lachnoptera Doubleday, 1847 Genus Pseudargynnis Karsch, 1892 L. anticlia (Hübner, 1819) Davenport P. hegemone (Godart, 1819) Forbes/Davenport Genus Phalanta Horsfield, 1829 Genus Cynandra Schatz, 1887 P. phalantha aethiopica (Rothschild & Forbes C. opis bernardii Lagnel, 1967 Forbes/Davenport Jordan, 1903) Genus Euryphura Staudinger, 1891

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E. chalcis chalcis (Felder & Felder, 1860) Forbes Tribe Leptosiaini Braby, 2014 E. plautilla (Hewitson, 1865) Forbes Genus Leptosia Hübner, 1818 Genus Euphaedra Hübner, 1819 L. wigginsi wigginsi (Dixey, 1915) Forbes/Davenport E. medon inaequabilis Thurau, 1904 Forbes/Davenport L. nupta pseudonupta Bernardi, 1959 Forbes/Davenport E. zaddachi crawshayi Butler, 1895 Forbes L. alcesta inalcesta Bernardi, 1959 Davenport E. mirabilis nubila Hecq, 1986 Forbes Tribe Nepheroniini Braby, 2014 E. caerulescens caerulescens Grose Forbes Genus Nepheronia Butler, 1870 Smith, 1890 N. pharis silvanus (Stoneham, 1957) Forbes/Davenport E. christyi Sharpe, 1904 Forbes N. thalassina verulanus (Ward, 1871) Forbes/Davenport E. eberti hamus Berger, 1940 Forbes N. argia argia (Fabricius, 1775) Forbes/Davenport E. diffusa diffusa Gaede, 1916 Forbes/Davenport N. buquetii buquetii (Boisduval, 1836) Forbes/Davenport E. imitans Holland, 1893 Forbes Tribe Teracolini Reuter, 1896 E. sarita sarita (Sharpe, 1891) Forbes Genus Colotis Hübner, 1819 E. sardetta Gaede, 1916 Forbes C. euippe complexivus (Butler, 1886) Forbes/Davenport E. viridicaerulea nitidula van Someren, Forbes C. hetaera ankolensis Stoneham, 1940 Selb 1935 C. elgonensis basilewskyi Berger, 1956 Davenport E. preussi preussi Staudinger, 1891 Forbes C. antevippe zera (Lucas, 1852) Forbes/Davenport E. albofasciata Berger, 1981 Forbes Genus Eronia Hübner, 1823 E. eleus eleus (Drury, 1782) Forbes/Davenport E. cleodora Hübner, 1823 Forbes/Davenport E. hybrida Hecq, 1978 Forbes Genus Afrodryas Stoneham, 1957 E. alacris Hecq, 1978 Forbes A. leda (Boisduval, 1847) Selb E. rattrayi rattrayi Sharpe, 1904 Forbes Tribe Pierini Swainson, 1820 E. ruspina (Hewitson, 1865) Forbes Subtribe Appiadina Kuzsenov, 1921 E. edwardsii langoueensis Oremans, 2010 Forbes/Davenport Genus Appias Hübner, 1819 E. harpalyce spatiosa (Mabille, 1876) Forbes/Davenport A. sabina sabina (Felder & Felder, 1865) Davenport Genus Euriphene Boisduval, 1847 A. phaola intermedia Dufrane, 1848 Forbes/Davenport E. ribensis (Ward, 1871) Forbes/Davenport A. epaphia epaphia (Cramer, 1779) Forbes/Davenport E. atossa australis d’Abrera, 1980 Forbes/Davenport Subtribe Aporiina Chapman, 1895 E. amaranta (Karsch, 1894) Forbes/Glasgow Genus Belenois Hübner, 1819 E. saphirina saphirina (Karsch, 1894) Forbes/Davenport B. creona severina (Stoll, 1781) Forbes/Davenport Genus Bebearia Hemming, 1960 B. aurota (Fabricius, 1793) Forbes B. carshena (Hewitson, 1871) Forbes/Davenport B. crawshayi Butler, 1894 Davenport B. absolon entebbiae (Lathy, 1906) Forbes/Davenport B. calypso minor Talbot, 1943 Davenport B. abesa pandera Hecq, 1988 Forbes B. thysa meldolae Butler, 1872 Forbes/Davenport B. zonara (Butler, 1871) Forbes/Davenport B. theora laeta (Weymer, 1903) Selb B. mandinga beni Hecq, 1990 Forbes B. solilucis loveni (Aurivillius, 1921) Forbes/Davenport B. oxione squalida (Talbot, 1928) Forbes Genus Dixeia Talbot, 1932 B. partita (Aurivillius, 1895) Forbes/Davenport D. pigea (Boisduval, 1836) Davenport B. barce maculata (Aurivillius, 1912) Forbes/Davenport Genus Mylothris Hübner, 1819 B. plistonax (Hewitson, 1874) Forbes M. chloris chloris (Fabricius, 1775) Forbes/Davenport B. cocalia badiana (Rebel, 1914) Forbes/Davenport M. rhodope (Fabricius, 1775) Forbes/Selb B. sophus audeoudi (Riley, 1936) Forbes M. continua continua Talbot, 1944 Forbes B. seeldrayersi (Aurivillius, 1899) Forbes/Davenport Total for the subfamily Pierinae 27 B. brunhilda iturina (Karsch, 1894) Forbes/Davenport Total for the family Pieridae 34 B. laetitioides (Joicey & Talbot, 1921) Forbes/Davenport FAMILY RIODINIDAE Grote, 1895 B. flaminia flaminia (Staudinger, 1891) Forbes Subfamily Nemeobiinae Bates, 1868 Genus Hamanumida Hübner, 1819 Tribe Abisarini Stichel, 1928 H. daedalus (Fabricius, 1775) Forbes/Davenport Subtribe Abisarina Stichel, 1928 Tribe Pseudacraeini Dhungel & Genus Afriodinia d’Abrera, 2009 Wahlberg, 2018 A. neavei neavei (Riley, 1932) Unknown Genus Pseudacraea Westwood, 1850 Total for the family Riodinidae 1 P. eurytus eurytus (Linnaeus, 1758) Forbes/Davenport FAMILY LYCAENIDAE Leach, 1815 P. lucretia protracta (Butler, 1875) Forbes/Davenport Subfamily Poritiinae Doherty, 1886 P. semire (Cramer, 1779) Forbes/Davenport Tribe Pentilini Aurivillius, 1914 P. warburgi Aurivillius, 1892 Selb Genus Ptelina Clench, 1965 Total for the subfamily Limenitidinae 90 P. carnuta (Hewitson, 1873) Selb Total for the family Nymphalidae 263 Genus Pentila Westwood, 1851 FAMILY PIERIDAE Swainson, 1820 P. pauli clarensis Neave, 1903 Davenport/Selb Subfamily Pseudopontiinae Reuter, 1896 P. umangiana connectens Hulstaert, 1924 Davenport/Selb Genus Pseudopontia Plötz, 1870 P. inconspicua Druce, 1910 Davenport/Selb P. mabira Mitter & Collins, 2011 Selb P. tachyroides tachyroides Dewitz, 1879 Selb Total for subfamily Pseudopontiinae 1 P. cloetensi catauga Rebel, 1914 Selb Subfamily Coliadinae Swainson, 1821 Genus Liptenara Bethune-Baker, 1915 Genus Eurema Hübner, 1819 L. hiendlmayri (Dewitz, 1887) Davenport E. hecabe solifera (Butler, 1875) Forbes/Davenport Genus Telipna Aurivillius, 1895 E. floricola leonis (Butler, 1886) Forbes/Davenport T. aurivillii jefferyi Jackson, 1969 Forbes/Davenport E. senegalensis (Boisduval, 1836) Forbes/Davenport T. sanguinea depuncta Talbot, 1937 Forbes/Davenport E. desjardinsi regularis (Butler, 1876) Forbes/Davenport T. consanguinea consanguinea Rebel, Selb E. hapale (Mabille, 1882) Forbes/Davenport 1914 Genus Catopsilia Hübner, 1819 T. nyanza katangae Stempffer, 1961 Selb C. florella (Fabricius, 1775) Forbes/Davenport Genus Ornipholidotos Bethune-Baker, Total for the subfamily Coliadinae 6 1914 Subfamily Pierinae Swainson, 1820 O. overlaeti intermedia Libert, 2005 Davenport

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O. jacksoni jacksoni Stempffer, 1961 Selb Subfamily Polyommatinae Swainson, Tribe Liptenini Röber, 1892 1827 Genus Liptena Westwood, 1851 Tribe Lycaenesthini Toxopeus, 1929 L. opaca ugandana Stempffer, Bennett & Forbes Genus Anthene Doubleday, 1847 May, 1974 A. larydas (Cramer, 1780) Davenport/Selb L. xanthostola xantha (Grose-Smith, Davenport/Selb A. crawshayi crawshayi (Butler, 1899) Selb 1901) A. ligures ligures (Hewitson, 1874) Davenport/Selb L. praestans Schultze, 1923 Davenport/Selb A. sylvanus (Drury, 1773) Selb Genus Tetrarhanis Karsch, 1893 A. rubricinctus rubricinctus (Holland, Davenport T. ilala ilala (Riley, 1929) Forbes 1891) T. ilma ugandae (Stempffer, 1964) Davenport/Selb A. ituria (Bethune-Baker, 1910) Davenport/Selb Genus Falcuna Stempffer & Bennett, A. princeps (Butler, 1876) Selb 1963 A. liodes liodes (Hewitson, 1874) Selb F. margarita (Suffert, 1904) Davenport Genus Neurellipes Bethune-Baker, 1910 F. orientalis bwamba Stempffer & Selb N. fulvimacula (Mabille, 1890) Davenport Bennett, 1963 N. kampala triangularis Libert, 2010 Selb Genus Larinopoda Butler, 1871 N. scintillula scintillula (Holland, 1891) Davenport L. tera (Hewitson, 1873) Davenport/Selb Genus Triclema Karsch, 1893 L. lagyra lagyra (Hewitson 1866) Davenport/Selb T. staudingeri (Grose-Smith & Kirby, Davenport Genus Citrinophila Kirby, 1887 1894) C. erastus pallida Hawker-Smith, 1933 Forbes T. phoenicis Karsch, 1893 Selb Genus Toxochitona Stempffer, 1956 T. inconspicua latefascia Libert, 2010 Forbes/Selb T. sankuru Stempffer, 1961 Davenport Tribe Polyommatini Swainson, 1827 Genus Mimacraea Butler, 1872 Subtribe incertae sedis M. landbecki Druce, 1910 Forbes Genus Cupidopsis Karsch, 1895 M. krausei krausei Dewitz, 1889 Forbes/Davenport C. cissus cissus (Godart, 1824) Selb Genus Mimeresia Stempffer, 1961 Genus Pseudonacaduba Stempffer, M. drucei ugandae (Stempffer, 1954) Forbes 1943 M. moreelsi purpurea (Hawker-Smith, Forbes P. aethiops (Mabille, 1877) Davenport/Selb 1933) P. sichela sichela Wallengren, 1857) Davenport Tribe Epitolini Jackson, 1962 Genus Uranothauma Butler, 1895 Genus Epitola Westwood, 1851 U. falkensteinii (Dewitz, 1879) Davenport/Selb E. uranoides uranoides Libert, 1999 Selb U. cyara tenuimarginata (Grünberg, Davenport/Selb E. urania Kirby, 1887 Forbes 1908) Genus Cerautola Libert, 1999 Genus Cacyreus Butler, 1897 C. mittoni (Jackson, 1964) NMK C. lingeus (Stoll, 1782) Davenport/Selb C. subargentea subargentea (Jackson, Selb C. audeoudi Stempffer, 1936 Davenport/Selb 1965) Genus Leptotes Scudder, 1876 Genus Geritola Libert, 1999 L. pirithous pirithous (Linnaeus, 1767) Selb G. cynaea (Jackson, 1964) Davenport L. marginalis (Stempffer, 1942) Selb Genus Stempfferia Jackson, 1962 Genus Tuxentius Larsen, 1982 S. cercenoides (Holland, 1890) Davenport T. margaritaceus (Sharpe, 1892) Davenport/Selb S. tumentia (Druce, 1910) Davenport Genus Zizeeria Chapman, 1910 Genus Cephetola Libert, 1999 Z. knysna knysna (Trimen, 1862) Selb C. bwamba (Jackson, 1964) NMK Genus Actizera Chapman, 1910 C. pinodes budduana (Talbot, 1937) Selb A. lucida (Trimen, 1883) Selb Genus Epitolina Aurivillius, 1895 Genus Zizula Chapman, 1910 E. dispar (Kirby, 1887) Selb Z. hylax (Fabricius, 1775) Davenport Genus Phytala Westwood, 1851 Genus Azanus Moore, 1881 P. elais ugandae Jackson, 1964 Forbes/Selb A. isis Drury, 1773 Davenport/Selb Genus Hewitsonia Kirby, 1871 A. mirza (Plötz, 1880) Forbes/Selb H. ugandae ugandae Jackson, 1962 Selb A. moriqua (Wallengren, 1857) Selb H. inexpectata Bouyer, 1997 Selb A. natalensis (Trimen, 1887) Davenport H. intermedia Jackson, 1962 Forbes/Selb Genus Eicochrysops Bethune-Baker, Total for the subfamily Poritiinae 42 1924 Subfamily Miletinae Reuter, 1896 E. hippocrates (Fabricius, 1793) Forbes/Davenport Tribe Lachnocnemini Clench, 1955 Genus Euchrysops Butler, 1900 Genus Lachnocnema Trimen, 1887 E. osiris (Hopffer, 1855) Davenport L. exiguus Holland, 1890 Selb E. barkeri (Trimen, 1893) Davenport/Selb Tribe Spalgini Toxopeus, 1929 Genus Thermoniphas Karsch, 1895 Genus Spalgis Moore, 1879 T. alberici (Dufrane, 1945) Forbes/Davenport S. jacksoni jacksoni Stempffer, 1967 Selb T. distincta (Talbot, 1935) Davenport Tribe Miletini Reuter, 1896 T. fontainei Stempffer, 1956 Forbes Genus Megalopalpus Röber, 1886 Genus Oboronia Karsch, 1893 M. simplex Röber, 1886 Selb O. punctatus (Dewitz, 1879) Forbes/Davenport M. metaleucus Karsch, 1893 Selb O. pseudopunctatus (Strand, 1912) Forbes/Davenport M. zymna (Westwood, 1851) Forbes/Davenport O. guessfeldtii (Dewitz, 1879) Davenport Total for the subfamily Miletinae 5 Total for the subfamily Polyommatinae 40 Subfamily Aphnaeinae Distant, 1884 Subfamily Swainson, 1830 Genus Hübner, 1819 Tribe Amblypodini Doherty, 1886 A. orcas (Drury, 1782) Forbes/Davenport Genus Myrina Fabricius, 1807 Total for the subfamily Aphnaeinae 1 M. sharpei sharpei Bethune-Baker, 1906 Williams/Dobson Tribe Iolaini Riley, 1958

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Genus Iolaus Hübner, 1819 I. bellina exquisita (Riley, 1928) Selb I. pollux albocaerulea (Riley, 1929) Forbes Tribe Hypolycaenini Swinhoe, 1910 Genus Hypolycaena Felder, 1862 H. antifaunus latimacula (Joicey & Davenport/Selb Talbot, 1921) H. philippus philippus (Fabricius, 1793) Davenport/Selb H. nigra Bethune-Baker, 1914 Davenport H. hatita ugandae Sharpe, 1904 Forbes/Davenport Tribe Oxylidini Eliot, 1973 Genus Oxylides Hübner, 1819 O. albata (Aurivillius, 1895) Forbes/Selb O. feminina feminina (Sharpe, 1904) Selb Tribe Deudorigini Doherty, 1886 Genus Deudorix Hewitson, 1863 D. antalus (Hopffer, 1855) Davenport Total for the subfamily Theclinae 10 Total for the family Lycaenidae 98 Species total 448