Pseudoscorpiones), with Remarks on the Composition of the Family

Total Page:16

File Type:pdf, Size:1020Kb

Pseudoscorpiones), with Remarks on the Composition of the Family DOI: 10.18195/issn.0312-3162.24(4).2008.387-394 f\ccord.' ottflC WcstCrIl AlIstrllliall !V1l1SClIlll 24: 387-394 (2008) The first troglomorphic pseudoscorpion of the family Olpiidae (Pseudoscorpiones), with remarks on the composition of the family Mark S. Harvey1,2 and Mei Chen Leng' [Department of Terrestrial Zoology, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia 6986, Australia. Email: [email protected]. 'School of Animal Biology, University of Western Australia, Crawley, Western Australia 6009, Australia. Abstract -A new genus and species of Olpiidae, Lillllllcolpilllll lillllllCi, is de- scribed from a pisolite mesa in the Pilbara region of Western Australia. The sole specimen exhibits troglomorphic facies with highly reduced eyes, pallid colouration and slightly attenuated appendages, differing from all other cave- dwelling olpiids which lack troglomorphic facies. The affinities of the new genus are difficult to determine, but it is most similar to a group of genera characterised by the position of trichobothrium ish: f'.SClIdollOrJlS, AlIS!rollOrJlS, XCllolpilllll, ElIryolpilllll and possibly Hc!crofpilllll. The family Olpiidae is di- vided into two subfamilies, Olpiinae and IIesperolpiinae. INTRODUCTION (1986), who based his system on the observations Pseudoscorpions of the family Olpiidae are of Heurtault (1980a, 1980b, 1982) and Heurtault virtually cosmopolitan in distribution, usually and Rebiere (1983). occurring in xeric environments under rocks, The specimen was examined with an Olympus in leaf litter or beneath the bark of trees. Cave- BH-2 compound microscope and illustrated with dwelling olpiids are, however, quite rare, with the aid of a drawing tube. Measurements were only five species recorded from subterranean taken at the highest possible magnification using habitats: Ca/oclzeirlls tenerifae Mahnert 2002 an ocular graticule. The specimen was examined by from the Canary Islands (Mahnert 2(02), and preparing a temporary slide mount by immersing l Progaryplls gracilis Mahnert 2001, P. /iliac Mahnert the specimen in 20 ;;, lactic acid at room temperature 2001, P. nigrimanlls Mahnert 2001, and P. sdifer for several days, and mounting it on a microscope Mahnert 2001 from Brazil (Mahnert 2(01). All of slide with 10 or 12 mm coverslips supported by these species have large eyes and seem to exhibit small sections of 0.25 mm or 0.50 mm diameter no troglomorphic morphological modifications. nylon fishing line. After study the specimen Among specimens collected from subterranean were returned to 75% ethanol with the dissected habitats within mesas in the Pilbara region portions placed in 12 x 3 mm glass genitalia of Western Australia was a single male of an microvials (BioQuip Products, Inc.). Images of the unusual olpiid. The specimen has reduced eyes, whole animal were taken using a Micropublisher is slightly pale in colouration, and has slightly 5.0 digital camera mounted on a Leica MZ16 attenuate appendages. To further document the microscope. subterranean pseudoscorpion fauna of Western Australia we present a description of this species Family Olpiidae Banks 1895 based upon a single male specimen. We also Subfamily Olpiinae Banks 1895 propose a division of the Olpiidae into two subfamilies, Olpiinae and Hesperolpiinae, based Remarks primarily upon the length of the venom ducts. For most of the past 80 years, the Olpiidae were The specimen examined in the present study defined by a series of characters states outlined by is lodged in the Western Australian Museum, Chamberlin (1930, 1931) and Beier (1932). Although Perth (WAM). Terminology and mensuration numerous new genera have been described since mostly follows Chamberlin (1931), with the the 1930's, the family definition has barely altered. exception of the nomenclature of the chelicera The Olpiidae were recently restricted by Judson (Judson 2(07), pedipalps, legs and with some (1992, 1(93) by the removal of the garypinines to m inor mod ifications to the term inology of the a separate family, Carypinidae, and the removal trichobothria (Harvey 1992). In particular, it of the hesperolpiines as a subfamily of C;,1l'ypidae. should be noted that the terminology for the The status of the garypinines as a family was trichobothria used by Harvey (1992) differs confirmed by Judson (200S), who referred to his slightly from that used by other workers. The unpublished thesis (Judson 1(92) in defining the setal notation of metatarsus IV follows Muchmore group. With the exclusion of the garypinines from 388 M.S. Harvey, M.C. Leng the family Olpiidae, and the transfer of a further that the type species of Xenolpium, X. pacificum five genera (Neominniza Beier 1930, Oreolpium (With 1907) from New Zealand, possessed leg Benedict and Malcolm 1978, Protogarypinus Beier morphology characteristic of the Olpiini requiring 1954, TeratolpiuJ1l Beier 1959 and Thaumatolpium the automatic synonymy of the two tribes. In Beier 1931) from the Olpiidae to the Garypinidae the meantime, Hoff (1964) included a third tribe by Harvey and Stahlavsky (unpublished data), the - Hesperolpiini Hoff 1964 - within the Olpiinae family now contains 33 genera (Table 1). based upon the long venom ducts within the Hoff (1945) divided the Olpiinae into the tribes chelal fingers, and the patella of leg I shorter than Olpiini Banks 1895 and Xenolpiini Hoff 1945 the femur, with a freely mobile joint. Hoff (1964) based upon the morphology of the first pair included Hesperolpium and Aphelolpium within of legs in which the femur are longer than the the Hesperolpiini, which was augmented by the patella in Olpiini, with a freely moveable joint, inclusion of Planctolpium Hoff 1964 by Muchmore but the two segments are subequal in Xenolpiini, (1979), previously described as a member of the thus allowing only restricted movement. Both Garypidae (Hoff 1964). Long venom ducts are tribes were rediagnosed by Hoff (1964), but were also found in a variety of other olpiids: Apolpium subsequently used only intermittently by later Chamberlin 1930 (Chamberlin 1931; Tooren researchers. They were abandoned by Heurtault 2002; Harvey, personal observation), Calocheirus (1980b), who presented new data on the type Chamberlin 1930 (Mahnert 1986, 2002), Cardiolpium species of several Old World genera, including Mahnert 1986 (Mahnert 1986), Ectactolpium Beier the type genera of the two tribes, Olpium L. 1947 (Harvey, personal observation), Nanolpium Koch 1873 and Xenolpium Chamberlin 1930. In Beier 1947 (Harvey, personal observation), particular, Heurtault (1979, 1980a, 1980b) found Progarypus Beier 1931 (Mahnert 2001; Harvey, Table 1 Genera assigned to the Olpiidae. Family Olpiidae Banks 1895 Subfamily Olpiinae Banks 1895 AntiLlolpium Muchmore 1991 Austrohoms Beier 1966 Banksolpium Muchmore 1986 Beierolpium Heurtault 1977 Calocheiridius Beier and Turk 1952 Euryolpium Redikorzev 1938 Halominniza Mahnert 1975 Heterolpium Sivaraman 1980 Hoffhorus Heurtault 1977 Homs Chamberlin 1930 Indolpium Hoff 1945 Leptolpium Tooren 2002 Linnaeolpiurn gen. novo Minniza Simon 1881 Neopachyolpium Hoff 1945 Nipponogarypus Morikawa 1955 Novohorus Hoff 1945 Olpiolum Beier 1931 Olpium L. Koch 1873 Pachyolpium Beier 1931 Parolpium Beier 1931 Pseudohorus Beier 1946 Xenolpium Chamberlin 1930 Subfamily Hesperolpiinae Hoff 1964 Aphelolpium Hoff 1964 Apolpium Chamberlin 1930 Calocheirus Chamberlin 1930 Cardiolpium Mahnert 1986 Ectactolpium Beier 1947 Hesperolpium Chamberlin 1930 Nanolpium Beier 1947 Planctolpium Hoff 1964 Progarypus Beier 1931 Stenolpiodes Beier 1959 Figure 1 sp. nov., holotype male Stenolpiun1 Beier 1955 Linnaeolpium linnaei (WAM T82354). Firsttroglomorphic Olpiidae 389 personal observation), Stcllolpiodcs Beier 1959 venom ducts characteristic of the rlesperolpiinae U~larvey, personal observation) and Stcllolpilllll are also found in Ceogarypidae, Carypidae, Beier 1955 (Mahnert 1984; Harvey, personal Larcidae and some Carypinidae (Allzlilyolpilll/1 observation). All of these genera, along with and Ncoal/1blyolpillllz). The lack of any features ApiTclolpilllll, f-icspcrolpilllll and Pla/lctolpilllll, arc providing firm evidence for the monophyly here regarded as hesperolpiines, based upon the of the Olpiidae suggests that the Olpiinae and criteria explicitly outlined by Hoff (1964). Hesperolpiinae may not be sister taxa, concordant Although we concur with Heurtault (1979, with the analyses of Murienne cl al. (2008), and 1980a, 1980b) that a satisfactory division between that other character systems need to be explored the Olpiini and Xenolpiini is unobtainable, we to establish their affinities and status within the agree with Hoff (1964) that the hesperolpiines Carypoidea. appear to represent a distinct diagnosable entity, which is here recognised as a subfamily of the Genus LilllwcolpiulIl gen. novo Olpiidae. The proposal by Judson (1992, 1993) that the ~Iesperolpiinae represents a member of Type species the Garypidae remains untested (but see below). Li/lllllColpilll11 lillllaci sp. novo The Hesperolpiinae contains 11 genera, whereas the relimited Olpiinae contains 22 genera (Table Diagnosis 1). Nearly all Olpiinae possess short venom ducts Lillllllcolpilll11 differ from all other olpiid genera where they generally do not reach ct in the fixed by the following combination of characters: two chelal finger or t in the moveable chelal finger. blades in the cheliceral rallum (Figure 8), reduced The most obvious exception is Mi/llliza barkizalllac eyes (Figure 2), the lack of enlarged tactile setae Mahnert
Recommended publications
  • Information to Users
    INFORMATION TO USERS The most advanced technology has been used to photograph and reproduce this manuscript from the microfilm master. UMI films the text directly from the original or copy submitted. Thus, some thesis and dissertation copies are in typewriter face, while others may be from any type of computer printer. The quality of this reproduction is dependent upon the quality of the copy submitted. Broken or indistinct print, colored or poor quality illustrations and photographs, print bleedthrough, substandard margins, and improper alignment can adversely affect reproduction. In the unlikely event that the author did not send UMI a complete manuscript and there are missing pages, these will be noted. Also, if unauthorized copyright material had to be removed, a note will indicate the deletion. Oversize materials (e.g., maps, drawings, charts) are reproduced by sectioning the original, beginning at the upper left-hand corner and continuing from left to right in equal sections with small overlaps. Each original is also photographed in one exposure and is included in reduced form at the back of the book. Photographs included in the original manuscript have been reproduced xerographically in this copy. Higher quality 6" x 9" black and white photographic prints are available for any photographs or illustrations appearing in this copy for an additional charge. Contact UMI directly to order. University Microfilms International A Bell & Howell Information Company 300 North Zeeb Road. Ann Arbor, Ml 48106-1346 USA 313/761-4700 800/521-0600 Order Number 9111799 Evolutionary morphology of the locomotor apparatus in Arachnida Shultz, Jeffrey Walden, Ph.D.
    [Show full text]
  • Pseudalbiorix, a New Genus of Ideoroncidae (Pseudoscorpiones, Neobisioidea) from Central America
    2006 (2007). The Journal of Arachnology 34:610–626 PSEUDALBIORIX, A NEW GENUS OF IDEORONCIDAE (PSEUDOSCORPIONES, NEOBISIOIDEA) FROM CENTRAL AMERICA Mark S. Harvey: Department of Terrestrial Invertebrates, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia 6986, Australia. E-mail: [email protected] Rene´ Barba Dı´az: Dept. de Invertebrados, Instituto de Ecologı´a y Sistema´tica, A.P. 8029, C.P. 10800, Habana 8, Cuba. Current address: Grupo BioKarst, Sociedad Espeleolo´gica de Cuba, Apartado Postal 2778, Habana 13, C.P. 11300, Cuba William B. Muchmore: Department of Biology, University of Rochester, Box 270211, Rochester, New York 14627-0211, USA Abel Pe´rez Gonza´lez: Grupo Biokarst, Sociedad Espeleolo´gica de Cuba. Current address: Dept. Invertebrados, Lab. Aracnologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista s/n, Sa˜o Cristo´va˜o 20.940-040, Rio de Janeiro, Brazil ABSTRACT. A new genus of Ideoroncidae, Pseudalbiorix, is described from Central America, and is found to consist of four species: the type species P. reddelli (Muchmore 1982), new combination from southern Mexico, P. veracruzensis (Hoff 1945), new combination from Belize, Guatemala and southern Mexico, and P. muchmorei Barba & Pe´rez, new species and P. armasi Barba & Pe´rez, new species from western Cuba. Pseudalbiorix reddelli and P. veracruzensis are transferred from the genus Albiorix. Mem- bers of this genus differ from all other ideoroncids principally in the morphology of the chelal externo- distal condyle. All post-embryonic stages of P. reddelli are described. Keywords: Pseudoscorpions, Mexico, Cuba, taxonomy, morphology, new species, biospeleology, trog- lobite The pseudoscorpion family Ideoroncidae ican species of Typhloroncus, and several spe- consists of several genera found in disparate cies of Albiorix, are restricted to caves.
    [Show full text]
  • Reprint Covers
    TEXAS MEMORIAL MUSEUM Speleological Monographs, Number 7 Studies on the CAVE AND ENDOGEAN FAUNA of North America Part V Edited by James C. Cokendolpher and James R. Reddell TEXAS MEMORIAL MUSEUM SPELEOLOGICAL MONOGRAPHS, NUMBER 7 STUDIES ON THE CAVE AND ENDOGEAN FAUNA OF NORTH AMERICA, PART V Edited by James C. Cokendolpher Invertebrate Zoology, Natural Science Research Laboratory Museum of Texas Tech University, 3301 4th Street Lubbock, Texas 79409 U.S.A. Email: [email protected] and James R. Reddell Texas Natural Science Center The University of Texas at Austin, PRC 176, 10100 Burnet Austin, Texas 78758 U.S.A. Email: [email protected] March 2009 TEXAS MEMORIAL MUSEUM and the TEXAS NATURAL SCIENCE CENTER THE UNIVERSITY OF TEXAS AT AUSTIN, AUSTIN, TEXAS 78705 Copyright 2009 by the Texas Natural Science Center The University of Texas at Austin All rights rereserved. No portion of this book may be reproduced in any form or by any means, including electronic storage and retrival systems, except by explict, prior written permission of the publisher Printed in the United States of America Cover, The first troglobitic weevil in North America, Lymantes Illustration by Nadine Dupérré Layout and design by James C. Cokendolpher Printed by the Texas Natural Science Center, The University of Texas at Austin, Austin, Texas PREFACE This is the fifth volume in a series devoted to the cavernicole and endogean fauna of the Americas. Previous volumes have been limited to North and Central America. Most of the species described herein are from Texas and Mexico, but one new troglophilic spider is from Colorado (U.S.A.) and a remarkable new eyeless endogean scorpion is described from Colombia, South America.
    [Show full text]
  • Review of the Family Bochicidae, with New Species and Records (Arachnida: Pseudoscorpionida)
    University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Center for Systematic Entomology, Gainesville, Insecta Mundi Florida March 1998 Review of the family Bochicidae, with new species and records (Arachnida: Pseudoscorpionida) William B. Muchmore Rochester, New York Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi Part of the Entomology Commons Muchmore, William B., "Review of the family Bochicidae, with new species and records (Arachnida: Pseudoscorpionida)" (1998). Insecta Mundi. 363. https://digitalcommons.unl.edu/insectamundi/363 This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 12, Nos. 1 & 2, March-June, 1998 117 Review of the family Bochicidae, with new species and records (Arachnida: Pseudoscorpionida) William B. Muchmore Department of Biology, University of Rochester Box 27021 1, Rochester, New York 14627-0211 Abstract: A history of the family is presented. All of the genera and species previously assigned to the family are reviewed, and 4 new species are described: Leucohya parva from Venezuela and Guyana, Mexobisiunz venii from Belize and Guatemala, M. dorninicanu~nfrom the Dominican Republic, and Paravachoniuln sprousei from Mexico. Two sub- families are established: Bochicinae (including the genera Alatillobisium, Bochica, Troglobochica, Troglohya, and Vacholziu~n)and Leucohyinae (including Apohya, Leucohya, Mexobisiurn, and Paravachoniurn). Introduction genus Mirobisiu7n Beier in the Bochicinae along with Bochica. The history of the family Bochicidae is long 1937: C. F. Roewer followed Beier's classifica- and complex.
    [Show full text]
  • Fauna Survey Report Lake Disappointment Potash Project Reward Minerals
    Fauna Survey Report Lake Disappointment Potash Project Reward Minerals Ltd October 2017 Report Number: 01-000018-1 FINAL On behalf of: Reward Minerals Limited PO Box 1104 NEDLANDS WA 6909 T: (08) 9386 4699 E: [email protected] Prepared by: Greg Harewood Zoologist PO Box 755 BUNBURY WA 6231 M: 0402 141 197 E: [email protected] LAKE DISAPPOINTMENT POTASH PROJECT – REWARD MINERALS LTD – FAUNA SURVEY REPORT – OCTOBER 2017 – FINAL TABLE OF CONTENTS SUMMARY ...................................................................................................... III 1. INTRODUCTION ....................................................................................... 1 1.1 BACKGROUND ......................................................................................... 1 1.2 STUDY AREA ............................................................................................ 2 1.3 SURVEY SCOPE ....................................................................................... 2 2. METHODS ................................................................................................. 3 2.1 FAUNA INVENTORY – LITERATURE REVIEW ....................................... 3 2.1.1 Database Searches ................................................................................ 3 2.1.2 Previous Fauna Surveys in the Area ...................................................... 4 2.2 FAUNA INVENTORY - FAUNA SURVEYS ............................................... 6 2.2.1 Survey Timing and Weather ..................................................................
    [Show full text]
  • The Pseudoscorpion of Illinois
    BULLETIN of the ILLINOIS NATURAL HISTORY SURVEl HARLOW B. MILLS, Chief The Pseudoscorpion of Illinois C. CLAYTON HOFF ""''^^^m, ^/% Printed by Authority of the STATE OF ILLINOIS ADLAI E. STEVENSON, Governor DEPARTMENT OF REGISTRATION AND EDUCATION NOBLE J. PUFFER, Director STATE OF ILLINOIS Adlai E. Stevenson", Governor DEPARTMENT OF REGISTRATION AND EDUCATION Noble J. Puffer, Director NATURAL HISTORY SURVEY DIVISION Harlow B. Mills, Chief \'oliime 24 BULLETIN Article 4 The Pseudoscorpions of Illinois C. CLAYTON HOFF Printed by Authority of the State of Illinois URBAN A, ILLINOIS June 1949 STATE OF ILLINOIS Adlai E. Stevenson, Governor DEPARTMENT OF REGISTRATION AND EDUCATION Noble J. Puffer, Director BOARD OF NATURAL RESOURCES AND CONSERVATION Noble J. Piffer, Chairman A. E. Emerson, Ph.D., Biology George D. Stodrard, Ph.D., Litt.D., L.H.D., L. H. Tiffany, Ph.D., Forestry LL.D., President of the University oj Illinois L R. HowsoN, B.S.C.E., C.E., Walter H. Newhouse, Ph.D., Geology Engineering Roger Adams, Ph.D., D.Sc, Chemistry NATURAL HISTORY SURVEY DIVISION Urbana, Illinois Scientific and Technical Staff Harlow B. Mills, Ph.D., Chief Bessie B. Henderson, M.S., Assistant to the Chief Section of Economic Entomology Section of Applied Botany and Plant Pa thology George C. Decker, Ph.D., Entomologist and Head Leo R. Tehon, Ph.D., Botanist and Head Cedric Carter, Ph.D., Plant Pathologist J. H. Bigger, M.S., Entomologist J. L. L. English, Ph.D., Entomologist J. L. FoRSBERG, M.S., Associate Plant Patholo gist C. J. Weinman, Ph.D., Entomologist S. C. Chandler, B.S., Associate Entomologist G.
    [Show full text]
  • Museum of Comparative Zoology
    Museum of Comparative Zoology Harvard University Annual Report 2016. 2017 Director’s Message As I write this message during the fall of 2017, I confess to being more conflicted than in past years regarding the future of the natural world and the ability and inclination of human society to take steps needed to preserve and sustain it. In the last 12 months, the United States discovery and formal description of has formally declared its intention to animal species. This support will allow us withdraw from the United Nations to attract talented young researchers in Paris Agreement on climate change; an essential field and further their career threats to nationally protected lands development at a critical stage. The new are being raised from within the federal program may be launched as early as government; and the ability of the U.S. next academic year, and it is expected Environmental Protection Agency to to expand in subsequent years as the both monitor and protect the air we endowment grows. Melissa Aja Melissa breathe and the water we drink is being Nobody loves parties more than we do, severely curtailed, if not eliminated. James Hanken and we demonstrated this to great effect Director As an organization that is committed last spring with a ceremony to celebrate to exploring and understanding Earth’s the 16 staff members who have achieved biological diversity and sharing what a long-term employment milestone: we learn with professional colleagues, more than 25 years of service at Harvard students and laypeople, these University, with most of those years developments are of great concern.
    [Show full text]
  • Animal Biodiversity: an Outline of Higher-Level Classification and Survey of Taxonomic Richness”
    Order Pseudoscorpiones de Geer, 1778 (2 suborders)1 2 Suborder Epiocheirata Harvey, 1992 (2 superfamilies) Superfamily Chthonioidea Daday, 1888 (4 families) Family Chthoniidae Daday, 1888 (27 genera, 617 species [3 fs3]) † Family Dracochelidae Schawaller, Shear and Bonamo, 1991 (1 fg; 1 fs) Family Lechytiidae Chamberlin, 1929 (1 genus, 23 species [1 fs]) Family Pseudotyrannochthoniidae Beier, 1932 (5 genera, 44 species)4 Family Tridenchthoniidae Balzan, 1892 (15 genera, 71 species [1 fg; 1 fs]) Superfamily Feaelloidea Ellingsen, 1906 (2 families) Family Feaellidae Ellingsen, 1906 (1 genus, 12 species) Family Pseudogarypidae Chamberlin, 1923 (2 genera, 7 species [4 fs]) Suborder Iocheirata Harvey, 1992 (5 superfamilies) Superfamily Neobisioidea Chamberlin, 1930 (7 families) Family Bochicidae Chamberlin, 1930 (12 genera, 41 species) Family Gymnobisiidae Beier, 1947 (4 genera, 11 species) Family Hyidae Chamberlin, 1930 (2 genera, 14 species) Family Ideoroncidae Chamberlin, 1930 (11 genera, 59 species) Family Neobisiidae Chamberlin, 1930 (32 genera, 576 species [4 fs]) Family Parahyidae Harvey, 1992 (1 genus, 1 species) Family Syarinidae Chamberlin, 1930 (17 genera, 109 species) Superfamily Garypoidea Simon, 1879 (6 families) Family Garypidae Simon, 1879 (10 genera, 77 species) Family Garypinidae Daday, 1888 (21 genera, 76 species [2 fs])5 Family Geogarypidae Chamberlin, 1930 (3 genera, 60 species [3 fs]) Family Larcidae Harvey, 1992 (2 genera, 15 species) Family Menthidae Chamberlin, 1930 (5 genera, 12 species) Family Olpiidae Banks, 1895
    [Show full text]
  • Impact of Rainforest Conversion Into Monoculture Plantation Systems on Pseudoscorpion Density, Diversity and Trophic Niches
    93 (2) · August 2021 pp. 83–95 Impact of rainforest conversion into monoculture plantation systems on pseudoscorpion density, diversity and trophic niches Dana Fabienne Liebke1*, Danilo Harms2, Rahayu Widyastuti3, Stefan Scheu1,4 and Anton M. Potapov1,5 1 J. F. Blumenbach Institute of Zoology and Anthropology, Animal Ecology, University of Goettingen, Untere Karspühle 2, 37073 Göttingen, Germany 2 Zoological Museum Hamburg, Leibnitz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany 3 Department of Soil Sciences and Land Resources, Institut Pertanian Bogor-IPB, Damarga Campus, Bogor, 16680, Indonesia 4 Centre of Biodiversity and Sustainable Land Use, Büsgenweg 1, 37077 Göttingen, Germany 5 A. N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky Prospect 33, 119071 Moscow, Russia * Corresponing author, Email: [email protected] Received 18 December 2020 | Accepted 11 May 2021 Published online at www.soil-organisms.de 1 August 2021 | Printed version 15 August 2021 DOI 10.25674/so93iss2id147 Abstract Indonesia’s biodiversity is at risk due to large forest areas being transformed into rubber and oil palm monoculture plantations. The effects of this land-use change on local fauna have been studied in a variety of organisms, including invertebrates from leaf litter and soil habitats. Litter and soil organisms are important drivers of essential ecosystem functions, such as nutrient cycling and carbon sequestration, which are impacted heavily by monoculture plantations. Pseudoscorpions (Arachnida: Pseudoscorpiones) are predatory arthropods in such litter and soil habitats and are an ubiquitous, although typically not very abundant, component of the soil animal food web.
    [Show full text]
  • Order Pseudoscorpiones*
    Zootaxa 3703 (1): 034–035 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Correspondence ZOOTAXA Copyright © 2013 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3703.1.8 http://zoobank.org/urn:lsid:zoobank.org:pub:EABA78D9-83D4-4940-9531-0BD2AFC96FF6 Order Pseudoscorpiones* MARK S. HARVEY Department of Terrestrial Zoology, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia 6986, Australia; [email protected] * In: Zhang, Z.-Q. (Ed.) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013). Zootaxa, 3703, 1–82. The classification and diversity estimates of the order Pseudoscorpiones (Harvey 2011) are here updated. There are 26 Recent families, 454 genera, and 3,533 species; fossil taxa include 1 family, 11 genera, 41 species. Taxon numbers are calculated to the end of 2012. Classification Order Pseudoscorpiones de Geer, 1778 (2 suborders) 1 † Family Dracochelidae Schawaller, Shear and Bonamo, 1991 (1 fg; 1 fs) 2 Suborder Epiocheirata Harvey, 1992 (2 superfamilies) Superfamily Chthonioidea Daday, 1888 (4 families) Family Chthoniidae Daday, 1888 (28 genera, 650 species [3 fs]) Family Lechytiidae Chamberlin, 1929 (1 genus, 23 species [1 fs]) Family Pseudotyrannochthoniidae Beier, 1932 (5 genera, 49 species) Family Tridenchthoniidae Balzan, 1892 (15 genera, 71 species [1 fg; 1 fs]) Superfamily Feaelloidea Ellingsen, 1906 (2 families) Family Feaellidae Ellingsen, 1906 (1 genus, 12 species) Family Pseudogarypidae
    [Show full text]
  • Journal of Natural History a Review of the Pseudoscorpion Genus
    This article was downloaded by: [University of Bath] On: 13 February 2014, At: 11:57 Publisher: Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Journal of Natural History Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/tnah20 A review of the pseudoscorpion genus Ideoblothrus (Pseudoscorpiones, Syarinidae) from western and northern Australia Mark S. Harvey a & Karen L. Edward a a Department of Terrestrial Invertebrates , Western Australian Museum , Welshpool DC, Western Australia, Australia Published online: 28 Mar 2007. To cite this article: Mark S. Harvey & Karen L. Edward (2007) A review of the pseudoscorpion genus Ideoblothrus (Pseudoscorpiones, Syarinidae) from western and northern Australia, Journal of Natural History, 41:5-8, 445-472, DOI: 10.1080/00222930701219123 To link to this article: http://dx.doi.org/10.1080/00222930701219123 PLEASE SCROLL DOWN FOR ARTICLE Taylor & Francis makes every effort to ensure the accuracy of all the information (the “Content”) contained in the publications on our platform. However, Taylor & Francis, our agents, and our licensors make no representations or warranties whatsoever as to the accuracy, completeness, or suitability for any purpose of the Content. Any opinions and views expressed in this publication are the opinions and views of the authors, and are not the views of or endorsed by Taylor & Francis. The accuracy of the Content should not be relied upon and should be independently verified with primary sources of information. Taylor and Francis shall not be liable for any losses, actions, claims, proceedings, demands, costs, expenses, damages, and other liabilities whatsoever or howsoever caused arising directly or indirectly in connection with, in relation to or arising out of the use of the Content.
    [Show full text]
  • William B. Muchmore (1920-2017): His Taxonomic Contributions and Complete Bibliography
    C.D.R. Stephen and M.S. Harvey. William B. Muchmore (1920-2017): His taxonomic contributions and complete bibliography. Journal of Cave and Karst Studies, v. 80, no. 1, p. 39-48. DOI: 10.4311/1700036R1 WILLIAM B. MUCHMORE (1920–2017): HIS TAXONOMIC CONTRIBUTIONS AND A COMPLETE BIBLIOGRAPHY Charles D.R. Stephen1,C and Mark S. Harvey2 Abstract William B. Muchmore (1920–2017) was the most influential worker on cave pseudoscorpions in North America and a globally-recognized expert on this arachnid order. Aside from brief stints in herpetological embryology and isopod taxonomy, he dedicated his 62-year career to the study of pseudoscorpions, focusing on their taxonomy. He described 278 new species, of which 167 were from caves. The majority of North American cave pseudoscorpions were either initially described or revised by him. He also wrote on the phoretic habits of pseudoscorpions, and speculated that cave pseudoscorpion distributions may be influenced by troglophilic rodents and bats. A complete bibliography of his scien- tific papers is provided. William (“Bill”) Breuleux Muchmore (Fig. 1) was born in Cincinnati, Ohio, on May 20, 1920. He was the first child of Oliver Charles Muchmore (1892−1968) and Ruby Breuleux (1895−1958), both of Ohio. His family has a long history in the United States (U.S.), with ancestry on his mother’s side tracing to Philip Frederick Breuleux (born 1830, Busserel, Haute Saône, France; immigrated in 1860 to Sycamore, Ohio), and on his father’s side to John Muchmore (born 1692, Windsor, Connecticut Colony; now the State of Connecticut). He had two children with his wife Marjorie Murrin: Susan Muchmore (born 1947) and Patricia Muchmore (born 1950).
    [Show full text]