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The Taxonomic Status of Gladiolus Illyricus (Iridaceae) in Britain
The Taxonomic Status of Gladiolus illyricus (Iridaceae) in Britain Aeron Buchanan Supervisor: Fred Rumsey, Natural History Museum, London A thesis submitted in partial fulfilment of the requirements for the degree of Master of Science of Imperial College, London Abstract First noticed officially in Britain in 1855, Gladiolus illyricus (Koch) presents an interesting taxonomic and biogeographical challenge: whether or not this isolated northern population should be recognized as a separate sub-species. Fundamental conservation issues rest on the outcome. Here, the investigation into the relationship of the G. illyricus plants of the New Forest, Hampshire, to Gladiolus species across Europe, northern Africa and the middle east is initiated. Two chloroplast regions, one in trnL–trnF and the other across psbA–trnH have been sequenced for 42 speci- mens of G. illyricus, G. communis, G. italicus, G. atroviolaceus, G. triphyllos and G. anatolicus. Phylogenetic and biogeographical treatments support the notion of an east–west genetic gradation along the Mediterranean. Iberia particularly appears as a zone of high hybridization potential and the source of the New Forest population. Alignment with sequences obtained from GenBank give strong support to the classic taxonomy of Gladiolus being monophyletic in its sub-family, Ixioideae. Comments on these chloroplast regions for barcoding are also given. In conclusion, the genetic localization of Britain’s G. illyricus population as an extremity haplotype suggests that it could well deserve sub-species status. Contents 1 Introduction 2 2 Background 4 3 Materials and Methods 8 4 Results and Discussion 15 5 Conclusions 26 Appendices 28 References 56 1. Introduction G. illyricus in Britain Figure 1: G. -
California's Native Ferns
CALIFORNIA’S NATIVE FERNS A survey of our most common ferns and fern relatives Native ferns come in many sizes and live in many habitats • Besides living in shady woodlands and forests, ferns occur in ponds, by streams, in vernal pools, in rock outcrops, and even in desert mountains • Ferns are identified by producing fiddleheads, the new coiled up fronds, in spring, and • Spring from underground stems called rhizomes, and • Produce spores on the backside of fronds in spore sacs, arranged in clusters called sori (singular sorus) Although ferns belong to families just like other plants, the families are often difficult to identify • Families include the brake-fern family (Pteridaceae), the polypody family (Polypodiaceae), the wood fern family (Dryopteridaceae), the blechnum fern family (Blechnaceae), and several others • We’ll study ferns according to their habitat, starting with species that live in shaded places, then moving on to rock ferns, and finally water ferns Ferns from moist shade such as redwood forests are sometimes evergreen, but also often winter dormant. Here you see the evergreen sword fern Polystichum munitum Note that sword fern has once-divided fronds. Other features include swordlike pinnae and round sori Sword fern forms a handsome coarse ground cover under redwoods and other coastal conifers A sword fern relative, Dudley’s shield fern (Polystichum dudleyi) differs by having twice-divided pinnae. Details of the sori are similar to sword fern Deer fern, Blechnum spicant, is a smaller fern than sword fern, living in constantly moist habitats Deer fern is identified by having separate and different looking sterile fronds and fertile fronds as seen in the previous image. -
Checklist of Common Native Plants the Diversity of Acadia National Park Is Refl Ected in Its Plant Life; More Than 1,100 Plant Species Are Found Here
National Park Service Acadia U.S. Department of the Interior Acadia National Park Checklist of Common Native Plants The diversity of Acadia National Park is refl ected in its plant life; more than 1,100 plant species are found here. This checklist groups the park’s most common plants into the communities where they are typically found. The plant’s growth form is indicated by “t” for trees and “s” for shrubs. To identify unfamiliar plants, consult a fi eld guide or visit the Wild Gardens of Acadia at Sieur de Monts Spring, where more than 400 plants are labeled and displayed in their habitats. All plants within Acadia National Park are protected. Please help protect the park’s fragile beauty by leaving plants in the condition that you fi nd them. Deciduous Woods ash, white t Fraxinus americana maple, mountain t Acer spicatum aspen, big-toothed t Populus grandidentata maple, red t Acer rubrum aspen, trembling t Populus tremuloides maple, striped t Acer pensylvanicum aster, large-leaved Aster macrophyllus maple, sugar t Acer saccharum beech, American t Fagus grandifolia mayfl ower, Canada Maianthemum canadense birch, paper t Betula papyrifera oak, red t Quercus rubra birch, yellow t Betula alleghaniesis pine, white t Pinus strobus blueberry, low sweet s Vaccinium angustifolium pyrola, round-leaved Pyrola americana bunchberry Cornus canadensis sarsaparilla, wild Aralia nudicaulis bush-honeysuckle s Diervilla lonicera saxifrage, early Saxifraga virginiensis cherry, pin t Prunus pensylvanica shadbush or serviceberry s,t Amelanchier spp. cherry, choke t Prunus virginiana Solomon’s seal, false Maianthemum racemosum elder, red-berried or s Sambucus racemosa ssp. -
Bracken Management: Ecological, Archaeological and Landscape Issues and Priorities
Natural England Technical Information Note TIN047 Bracken management: ecological, archaeological and landscape issues and priorities For over sixty years research has been carried out on the ecology, management and control of bracken Pteridium aquilinum. Much of this has been aimed at discovering the best methods of control where it has invaded plant communities of higher conservation value, such as heathland and unimproved grassland. More recent research has focussed on the damage it may do to archaeology. Conflicts between different environmental interests can arise where bracken is present. Where they do it is important that any decisions on management are made in the light of the best information available and that the proposals are sustainable. This note identifies some of the issues and suggests priorities in a number of conflict situations. Further information is available in SIN011 Bracken and the TIN048 Bracken management and control. Bracken and Biodiversity Action Plan late summer when other grasslands have (BAP) priority habitats become scorched and less palatable. Grassland Heathland Bracken grows best on deep well drained soils. Both lowland and upland heath are considered It will invade grassland on acid and neutral soils. of international importance and recognised as It is less likely to be a problem on calcareous BAP priority habitats. They both may be invaded grasslands. Where it grows on acid grassland by bracken. In areas of established and healthy around moorland edges, it is of particular heath, bracken may only increase slowly. concern because of its negative impacts on However, in fragmented heath bracken densities valuable grassland communities. In these can increase rapidly. -
Action for Pearl-Bordered Fritillary
FCS Technical Note Support for pearl-bordered fritillary conservation under the Scottish Rural Development Programme (SRDP) Introduction This technical guidance note is aimed at landowners, managers and their advisors who are considering undertaking woodland management for pearl-bordered fritillary under SRDP. It provides information on the most suitable management to benefit the species and how this management might be achieved and assessed under SRDP. Background The pearl-bordered fritillary is a UKBAP Priority species as well as being listed by Scottish Natural Heritage (SNH) as a Species Action Framework (SAF) species. It is also one of the key woodland species identified for action by Forestry Commission Scotland in the Scottish Forestry Strategy 2006. Pearl-bordered fritillary is one of the most rapidly declining butterflies in Britain and Ireland. Scottish populations are declining less severely than those south of the Border and are therefore becoming of increasing significance making up over a third of the British population. It is in need of urgent conservation action. Forestry Commission Scotland has published a Species Action Note setting out conservation needs and priorities in more depth than is included in this note. Species information The pearl-bordered fritillary is widespread but local in Scotland with populations in the glens of the Highlands, Grampian, Argyll, Moray and Perthshire, and an isolated population in South West Scotland around Dumfries. In Scotland the pearl-bordered fritillary is a butterfly of woodland edges or the open spaces within woodlands and it has a one-year life cycle. It requires sunny, sheltered sites, normally south-facing hillsides, as both the adult butterfly and its caterpillar require a very warm micro- climate. -
Cyperaceae of Puerto Rico. Arturo Gonzalez-Mas Louisiana State University and Agricultural & Mechanical College
Louisiana State University LSU Digital Commons LSU Historical Dissertations and Theses Graduate School 1964 Cyperaceae of Puerto Rico. Arturo Gonzalez-mas Louisiana State University and Agricultural & Mechanical College Follow this and additional works at: https://digitalcommons.lsu.edu/gradschool_disstheses Recommended Citation Gonzalez-mas, Arturo, "Cyperaceae of Puerto Rico." (1964). LSU Historical Dissertations and Theses. 912. https://digitalcommons.lsu.edu/gradschool_disstheses/912 This Dissertation is brought to you for free and open access by the Graduate School at LSU Digital Commons. It has been accepted for inclusion in LSU Historical Dissertations and Theses by an authorized administrator of LSU Digital Commons. For more information, please contact [email protected]. This dissertation has been 64—8802 microfilmed exactly as received GONZALEZ—MAS, Arturo, 1923- CYPERACEAE OF PUERTO RICO. Louisiana State University, Ph.D., 1964 B o ta n y University Microfilms, Inc., Ann Arbor, Michigan CYPERACEAE OF PUERTO RICO A Dissertation I' Submitted to the Graduate Faculty of the Louisiana State University and Agricultural and Mechanical College in partial fulfillment of the requirements for the degree of Doctor of Philosophy in The Department of Botany and Plant Pathology by Arturo Gonzalez-Mas B.S., University of Puerto Rico, 1945 M.S., North Carolina State College, 1952 January, 1964 PLEASE NOTE: Not original copy. Small and unreadable print on some maps. Filmed as received. UNIVERSITY MICROFILMS, INC. ACKNOWLEDGMENT The author wishes to express his sincere gratitude to Dr. Clair A. Brown for his interest, guidance, and encouragement during the course of this investigation and for his helpful criticism in the preparation of the manuscript and illustrations. -
Ecological Influences in the Biogeography of the Austral Sedges
ECOLOGICALINFLUENCESINTHEBIOGEOGRAPHYOFTHE AUSTRALSEDGES jan-adriaan viljoen Dissertation presented in fulfillment of the requirements for the degree MSc in Botany Department of Biological Sciences Faculty of Sciences University of Cape Town UniversityFebruary of2016 Cape Town The copyright of this thesis vests in the author. No quotation from it or information derived from it is to be published without full acknowledgement of the source. The thesis is to be used for private study or non- commercial research purposes only. Published by the University of Cape Town (UCT) in terms of the non-exclusive license granted to UCT by the author. University of Cape Town Jan-Adriaan Viljoen. Ecological influences in the biogeography of the aus- tral sedges. MSc dissertation. University of Cape Town. Cape Town. February 2016. supervisors: A. Muthama Muasya G. Anthony Verboom ABSTRACT The biogeographic history of a species is a result of both stochastic processes such as dispersal and habitat filters that determine where a population with a given set of biological requirements can become es- tablished. In this dissertation, I examine the geographical and ecolog- ical distribution of the sedge tribe Schoeneae in conjunction with its inferred speciation history in order to determine the pattern of disper- sal and the environmental factors that have influenced establishment. The biogeographic reconstruction indicates numerous transoceanic dispersal events consistent with random diffusion from an Australian point of origin, but with a bias towards habitats with vegetation type and moisture regime similar to the ancestral conditions of the given subgroup (open and dry habitats in the majority of cases). The global distribution of the tribe also suggests a preference for low-nutrient soils, which I investigate at the local (microhabitat) scale by contrast- ing the distributions of the tribes Schoeneae and Cypereae on the Cape Peninsula along soil fertility axes. -
Buds, Berries & Leaves
Buds, berries & leaves Monitoring moorland plants Bee, butterfly, hare & plant illustrations © Chris Shields Bird illustrations © Mike Langman Housekeeping • Emergency Exits • Fire assembly point • Toilets Today’s Session 1. Presentation • The importance of moorlands • What is phenology? • Conservation works • Species links • Plant ecology, ID & folklore • ID Quiz SHORT BREAK • Upland habitats • How to conduct a survey • Submitting your records • How data will be used 2. Practical session to practice survey methods and field ID 3. Feedback The importance of moorlands • The Peak District & South Pennine moorlands are of great importance, being the most southerly point in the range of some species. • Climate change may affect these population ranges and it will be noticed here first. • Designated as both a Special Protection Area (SPA) for breeding birds and as a Special Area of Conservation (SAC) for internationally important habitats. What is phenology? • Phenology is the study of life history stages, such as leafing, flowering and berry ripening in plants, or migration and breeding in animals. Bees rely on nectar being available at the right • Changes in climate can alter the timing of time and plants rely on the pollinators too phenological events which may cause mismatch between the life stages of different species, for example: • Earlier or later flowering may change the amount of flowers available to pollinators • Changes in the fruiting period may result in reduced food availability for birds • Repeated recording of these crucial stages enables us to identify changes in phenology. Coat colour change in mountain hares is another example of phenology Courtesy of Tim Sparks www.naturescalendar.org.uk Which plants? Field and laboratory experiments have shown changes in the phenology of many moorland species. -
Evaluating the Conservation Significance of Basin
Evaluating the conservation significance of basin wetlands within the Avon Natural Resource Management region: Stage Three Assessment Method Susan Jones, Adrian Pinder, Lien Sim and Stuart Halse MAY 2009 1 2 Evaluating the conservation significance of basin wetlands within the Avon Natural Resource Management region: Stage Three Assessment Method May 2009 Prepared by Science Division Department of Environment and Conservation 3 4 Executive summary: Evaluating the conservation significance of basin wetlands within the Avon Natural Resource Management region: Stage Three Assessment Method . Introduction This publication describes a wetland evaluation and classification methodology for use at the individual wetland scale in the Avon Natural Resource Management (NRM) region. A trial of this method at two example wetlands in each biological wetland type is presented in section 6. Table 1 - Form of wetland inventory Form of wetland inventory Methodology Application Identification Delineation Classification √√√ Evaluation √√√ Publication details This methodology has been developed by the Science Division, Department of Environment and Conservation (DEC), Western Australia. The report was written by Susan Jones, Adrian Pinder, Lien Sim and Stuart Halse (DEC). The authors of this document would like to acknowledge the following people for their important contributions: • Members of the Wetland Status Working Group and Wetlands Coordinating Committee • John Lizamore, Danielle Halliday, Cara Francis, Margaret Collins, Anna Leung, Kirsty Quinlan -
Bracken Control • Want to Prepare a Bracken Management Plan
WHAT IS SEARS? SEARS is a partnership delivering rural and environmental services on behalf of the Scottish Government. We will provide: • a streamlined service; • easy access to information and advice; • a consistent and responsive service; and • a customer focused service WHO ARE THE SEARS PARTNERS? Animal Health Cairngorms National Park Authority (CNPA) Crofters Commission (CC) Deer Commission for Scotland (DCS) Forestry Commission Scotland (FCS) Loch Lomond & The Trossachs National Park Authority (LLTNPA) Scottish Environment Protection Agency (SEPA) Scottish Natural Heritage (SNH) Scottish Government Rural Payments and Inspections Directorate (SGRPID) www.sears.scotland.gov.uk email: [email protected] INTRODUCTION THIS LEAFLET IS INTENDED FOR FARMERS, FORESTERS AND MOORLAND MANAGERS WHO: • WANT TO KNOW MORE ABOUT BRACKEN CONTROL • WANT TO PREPARE A BRACKEN MANAGEMENT PLAN It is an updated version of a booklet originally produced by the Southern Uplands Partnership, which has proved useful to land managers throughout Scotland. Following the general information on these pages, the main steps which should be considered in order to produce a bracken management plan are laid out in Stages 1-5. A management plan will help to ensure that methods are appropriate and will achieve their aims, and is essential if you intend to seek support under the Scotland Rural Development Programme (SRDP) Rural Development Contracts - Rural Priorities. You may want to seek professional help with the production of a management plan. BACKGROUND Bracken ( Pteridium aquilinum ) is an important and natural part of our landscape, and its abundance appears to have fluctuated over thousands of years. Its invasive nature allows it to spread, and it now occurs widely within a variety of habitats throughout Scotland. -
Pteridium: Dennstaedtiaceae) This Separation May Correspond to the Basal Divergence Among in Australia
American Journal of Botany 96(5): 1041–1049. 2009. G LOBAL CHLOROPLAST PHYLOGENY AND BIOGEOGRAPHY OF BRACKEN ( PTERIDIUM ; DENNSTAEDTIACEAE) 1 Joshua P. Der, 2,4 John A. Thomson, 3 Jeran K. Stratford, 2,5 and Paul G. Wolf 2 2 Department of Biology, Utah State University, 5305 Old Main Hill, Logan, Utah 84322 USA; and 3 National Herbarium of New South Wales, Botanic Gardens Trust, Mrs Macquaries Road, Sydney, NSW 2000, Australia Bracken ferns (genus Pteridium ) represent an ancient species complex with a natural worldwide distribution. Pteridium has historically been treated as comprising a single species, but recent treatments have recognized several related species. Phenotypic plasticity, geographically structured morphological variation, and geographically biased sampling have all contributed to taxo- nomic confusion in the genus. We sampled bracken specimens worldwide and used variable regions of the chloroplast genome to investigate phylogeography and reticulate evolution within the genus. Our results distinguish two major clades within Pteridium , a primarily northern hemisphere Laurasian/African clade, which includes all taxa currently assigned to P. aquilinum , and a primar- ily southern hemisphere Austral/South American clade, which includes P. esculentum and P. arachnoideum . All European acces- sions of P. aquilinum subsp. aquilinum appear in a monophyletic group and are nested within a clade containing the African P. aquilinum taxa ( P. aquilinum subsp. capense and P. aquilinum subsp. centrali-africanum ). Our results allow us to hypothesize the maternal progenitors of two allotetraploid bracken species, P. caudatum and P. semihastatum . We also discuss the biogeography of bracken in the context of the chloroplast phylogeny. Our study is one of the fi rst to take a worldwide perspective in addressing variation in a broadly distributed species complex. -
Branch Circus Flora and Fauna Survey PDF Document
FLORA AND VEGETATION SURVEY Branch Circus and Hammond Road, Success Prepared by: Prepared for: RPS MUNTOC PTY LTD AND 290 Churchill Avenue, SUBIACO WA 6008 SILVERSTONE ASSET PTY LTD PO Box 465, SUBIACO WA 6904 C/O Koltasz Smith T: 618 9382 4744 PO Box 127 F: 618 9382 1177 E: [email protected] BURSWOOD WA 6100 W: www.rpsgroup.com.au Report No: L07263 Version/Date: Rev 0, June 2008 RPS Environment Pty Ltd (ABN 45 108 680 977) Document Set ID: 5546761 Version: 1, Version Date: 31/01/2017 Flora and Vegetation Survey Branch Circus and Hammond Road, Success Document Status Review Format RPS Release Issue Version Purpose of Document Orig Review Date Review Approval Date Draft A Draft For Internal Review KelMcC VanYeo 30.04.08 Draft B Draft For Client Review VanYeo KarGod 14.05.08 SN 30.05.08 Rev 0 Final for Issue VanYeo 10.06.08 DC 12.06.08 B. Hollyock 13.06.08 Disclaimer This document is and shall remain the property of RPS. The document may only be used for the purposes for which it was commissioned and in accordance with the Terms of Engagement for the commission. Unauthorised copying or use of this document in any form whatsoever is prohibited. L07263, Rev 0, June 2008 DOCUMENT STATUS / DISCLAIMER Document Set ID: 5546761 Version: 1, Version Date: 31/01/2017 Flora and Vegetation Survey Branch Circus and Hammond Road, Success EXECUTIVE SUMMARY Flora A total of 229 taxa were recorded from the survey area, of which 155 or 68% were native.