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The Taxonomic Status of Gladiolus Illyricus (Iridaceae) in Britain
The Taxonomic Status of Gladiolus illyricus (Iridaceae) in Britain Aeron Buchanan Supervisor: Fred Rumsey, Natural History Museum, London A thesis submitted in partial fulfilment of the requirements for the degree of Master of Science of Imperial College, London Abstract First noticed officially in Britain in 1855, Gladiolus illyricus (Koch) presents an interesting taxonomic and biogeographical challenge: whether or not this isolated northern population should be recognized as a separate sub-species. Fundamental conservation issues rest on the outcome. Here, the investigation into the relationship of the G. illyricus plants of the New Forest, Hampshire, to Gladiolus species across Europe, northern Africa and the middle east is initiated. Two chloroplast regions, one in trnL–trnF and the other across psbA–trnH have been sequenced for 42 speci- mens of G. illyricus, G. communis, G. italicus, G. atroviolaceus, G. triphyllos and G. anatolicus. Phylogenetic and biogeographical treatments support the notion of an east–west genetic gradation along the Mediterranean. Iberia particularly appears as a zone of high hybridization potential and the source of the New Forest population. Alignment with sequences obtained from GenBank give strong support to the classic taxonomy of Gladiolus being monophyletic in its sub-family, Ixioideae. Comments on these chloroplast regions for barcoding are also given. In conclusion, the genetic localization of Britain’s G. illyricus population as an extremity haplotype suggests that it could well deserve sub-species status. Contents 1 Introduction 2 2 Background 4 3 Materials and Methods 8 4 Results and Discussion 15 5 Conclusions 26 Appendices 28 References 56 1. Introduction G. illyricus in Britain Figure 1: G. -
California's Native Ferns
CALIFORNIA’S NATIVE FERNS A survey of our most common ferns and fern relatives Native ferns come in many sizes and live in many habitats • Besides living in shady woodlands and forests, ferns occur in ponds, by streams, in vernal pools, in rock outcrops, and even in desert mountains • Ferns are identified by producing fiddleheads, the new coiled up fronds, in spring, and • Spring from underground stems called rhizomes, and • Produce spores on the backside of fronds in spore sacs, arranged in clusters called sori (singular sorus) Although ferns belong to families just like other plants, the families are often difficult to identify • Families include the brake-fern family (Pteridaceae), the polypody family (Polypodiaceae), the wood fern family (Dryopteridaceae), the blechnum fern family (Blechnaceae), and several others • We’ll study ferns according to their habitat, starting with species that live in shaded places, then moving on to rock ferns, and finally water ferns Ferns from moist shade such as redwood forests are sometimes evergreen, but also often winter dormant. Here you see the evergreen sword fern Polystichum munitum Note that sword fern has once-divided fronds. Other features include swordlike pinnae and round sori Sword fern forms a handsome coarse ground cover under redwoods and other coastal conifers A sword fern relative, Dudley’s shield fern (Polystichum dudleyi) differs by having twice-divided pinnae. Details of the sori are similar to sword fern Deer fern, Blechnum spicant, is a smaller fern than sword fern, living in constantly moist habitats Deer fern is identified by having separate and different looking sterile fronds and fertile fronds as seen in the previous image. -
Checklist of Common Native Plants the Diversity of Acadia National Park Is Refl Ected in Its Plant Life; More Than 1,100 Plant Species Are Found Here
National Park Service Acadia U.S. Department of the Interior Acadia National Park Checklist of Common Native Plants The diversity of Acadia National Park is refl ected in its plant life; more than 1,100 plant species are found here. This checklist groups the park’s most common plants into the communities where they are typically found. The plant’s growth form is indicated by “t” for trees and “s” for shrubs. To identify unfamiliar plants, consult a fi eld guide or visit the Wild Gardens of Acadia at Sieur de Monts Spring, where more than 400 plants are labeled and displayed in their habitats. All plants within Acadia National Park are protected. Please help protect the park’s fragile beauty by leaving plants in the condition that you fi nd them. Deciduous Woods ash, white t Fraxinus americana maple, mountain t Acer spicatum aspen, big-toothed t Populus grandidentata maple, red t Acer rubrum aspen, trembling t Populus tremuloides maple, striped t Acer pensylvanicum aster, large-leaved Aster macrophyllus maple, sugar t Acer saccharum beech, American t Fagus grandifolia mayfl ower, Canada Maianthemum canadense birch, paper t Betula papyrifera oak, red t Quercus rubra birch, yellow t Betula alleghaniesis pine, white t Pinus strobus blueberry, low sweet s Vaccinium angustifolium pyrola, round-leaved Pyrola americana bunchberry Cornus canadensis sarsaparilla, wild Aralia nudicaulis bush-honeysuckle s Diervilla lonicera saxifrage, early Saxifraga virginiensis cherry, pin t Prunus pensylvanica shadbush or serviceberry s,t Amelanchier spp. cherry, choke t Prunus virginiana Solomon’s seal, false Maianthemum racemosum elder, red-berried or s Sambucus racemosa ssp. -
Bracken Management: Ecological, Archaeological and Landscape Issues and Priorities
Natural England Technical Information Note TIN047 Bracken management: ecological, archaeological and landscape issues and priorities For over sixty years research has been carried out on the ecology, management and control of bracken Pteridium aquilinum. Much of this has been aimed at discovering the best methods of control where it has invaded plant communities of higher conservation value, such as heathland and unimproved grassland. More recent research has focussed on the damage it may do to archaeology. Conflicts between different environmental interests can arise where bracken is present. Where they do it is important that any decisions on management are made in the light of the best information available and that the proposals are sustainable. This note identifies some of the issues and suggests priorities in a number of conflict situations. Further information is available in SIN011 Bracken and the TIN048 Bracken management and control. Bracken and Biodiversity Action Plan late summer when other grasslands have (BAP) priority habitats become scorched and less palatable. Grassland Heathland Bracken grows best on deep well drained soils. Both lowland and upland heath are considered It will invade grassland on acid and neutral soils. of international importance and recognised as It is less likely to be a problem on calcareous BAP priority habitats. They both may be invaded grasslands. Where it grows on acid grassland by bracken. In areas of established and healthy around moorland edges, it is of particular heath, bracken may only increase slowly. concern because of its negative impacts on However, in fragmented heath bracken densities valuable grassland communities. In these can increase rapidly. -
Quarterly Volume 34 Number 1 Issue #133 Winter 2011 North American Heather Society
Heather Quarterly Volume 34 Number 1 Issue #133 Winter 2011 North American Heather Society Reflections on David Small Karla Lortz ..................................2 Long-distance collaboration with David Small David Wilson ..........................................................................3 David Small – A Personal Appreciation David Plumridge........4 David Small as educator Bryan Taylor.......................................6 David Small, no Pooh-Bah Judy Wiksten ................................7 A special man Susan Ewalt ......................................................9 David Small: reflections on a giant Donald A. M. Mackay.............................................................10 The irreplaceable David Small Ella May T. Wulff.................12 NAHS Board meeting minutes.....................................................24 Calendar....................................................................................28 North American Heather Society Membership Chair Ella May Wulff, Knolls Drive 2299 Wooded Philomath, OR 97370-5908 RETURN SERVICE REQUESTED issn 1041-6838 Heather News, all rights reserved, is published quarterly by the North American Heather Society, a tax exempt organization. The purpose of The Society is the: The Information Page (1) advancement and study of the botanical genera Andromeda, Calluna, Cassiope, Daboecia, Erica, and Phyllodoce, commonly called heather, and related genera; (2) HOW TO GET THE latest heather INFORMation dissemination of information on heather; and (3) promotion of fellowship -
Journal of the Royal Horticultural Society of London
I 3 2044 105 172"381 : JOURNAL OF THE llopl lortimltoal fbck EDITED BY Key. GEORGE HEXSLOW, ALA., E.L.S., F.G.S. rtanical Demonstrator, and Secretary to the Scientific Committee of the Royal Horticultural Society. VOLUME VI Gray Herbarium Harvard University LOXD N II. WEEDE & Co., PRINTERS, BEOMPTON. ' 1 8 8 0. HARVARD UNIVERSITY HERBARIUM. THE GIFT 0F f 4a Ziiau7- m 3 2044 i"05 172 38" J O U E N A L OF THE EDITED BY Eev. GEOEGE HENSLOW, M.A., F.L.S., F.G.S. Botanical Demonstrator, and Secretary to the Scientific Committee of the Royal Horticultural Society. YOLUME "VI. LONDON: H. WEEDE & Co., PRINTERS, BROMPTON, 1 8 80, OOUITOIL OF THE ROYAL HORTICULTURAL SOCIETY. 1 8 8 0. Patron. HER MAJESTY THE QUEEN. President. The Eight Honourable Lord Aberdare. Vice- Presidents. Lord Alfred S. Churchill. Arthur Grote, Esq., F.L.S. Sir Trevor Lawrence, Bt., M.P. H. J". Elwes, Esq. Treasurer. Henry "W ebb, Esq., Secretary. Eobert Hogg, Esq., LL.D., F.L.S. Members of Council. G. T. Clarke, Esq. W. Haughton, Esq. Colonel R. Tretor Clarke. Major F. Mason. The Rev. H. Harpur Crewe. Sir Henry Scudamore J. Denny, Esq., M.D. Stanhope, Bart. Sir Charles "W. Strickland, Bart. Auditors. R. A. Aspinall, Esq. John Lee, Esq. James F. West, Esq. Assistant Secretary. Samuel Jennings, Esq., F.L S. Chief Clerk J. Douglas Dick. Bankers. London and County Bank, High Street, Kensington, W. Garden Superintendent. A. F. Barron. iv ROYAL HORTICULTURAL SOCIETY. SCIENTIFIC COMMITTEE, 1880. Chairman. Sir Joseph Dalton Hooker, K.C.S.I., M.D., C.B.,F.R.S., V.P.L.S., Royal Gardens, Kew. -
Action for Pearl-Bordered Fritillary
FCS Technical Note Support for pearl-bordered fritillary conservation under the Scottish Rural Development Programme (SRDP) Introduction This technical guidance note is aimed at landowners, managers and their advisors who are considering undertaking woodland management for pearl-bordered fritillary under SRDP. It provides information on the most suitable management to benefit the species and how this management might be achieved and assessed under SRDP. Background The pearl-bordered fritillary is a UKBAP Priority species as well as being listed by Scottish Natural Heritage (SNH) as a Species Action Framework (SAF) species. It is also one of the key woodland species identified for action by Forestry Commission Scotland in the Scottish Forestry Strategy 2006. Pearl-bordered fritillary is one of the most rapidly declining butterflies in Britain and Ireland. Scottish populations are declining less severely than those south of the Border and are therefore becoming of increasing significance making up over a third of the British population. It is in need of urgent conservation action. Forestry Commission Scotland has published a Species Action Note setting out conservation needs and priorities in more depth than is included in this note. Species information The pearl-bordered fritillary is widespread but local in Scotland with populations in the glens of the Highlands, Grampian, Argyll, Moray and Perthshire, and an isolated population in South West Scotland around Dumfries. In Scotland the pearl-bordered fritillary is a butterfly of woodland edges or the open spaces within woodlands and it has a one-year life cycle. It requires sunny, sheltered sites, normally south-facing hillsides, as both the adult butterfly and its caterpillar require a very warm micro- climate. -
Bracken Control • Want to Prepare a Bracken Management Plan
WHAT IS SEARS? SEARS is a partnership delivering rural and environmental services on behalf of the Scottish Government. We will provide: • a streamlined service; • easy access to information and advice; • a consistent and responsive service; and • a customer focused service WHO ARE THE SEARS PARTNERS? Animal Health Cairngorms National Park Authority (CNPA) Crofters Commission (CC) Deer Commission for Scotland (DCS) Forestry Commission Scotland (FCS) Loch Lomond & The Trossachs National Park Authority (LLTNPA) Scottish Environment Protection Agency (SEPA) Scottish Natural Heritage (SNH) Scottish Government Rural Payments and Inspections Directorate (SGRPID) www.sears.scotland.gov.uk email: [email protected] INTRODUCTION THIS LEAFLET IS INTENDED FOR FARMERS, FORESTERS AND MOORLAND MANAGERS WHO: • WANT TO KNOW MORE ABOUT BRACKEN CONTROL • WANT TO PREPARE A BRACKEN MANAGEMENT PLAN It is an updated version of a booklet originally produced by the Southern Uplands Partnership, which has proved useful to land managers throughout Scotland. Following the general information on these pages, the main steps which should be considered in order to produce a bracken management plan are laid out in Stages 1-5. A management plan will help to ensure that methods are appropriate and will achieve their aims, and is essential if you intend to seek support under the Scotland Rural Development Programme (SRDP) Rural Development Contracts - Rural Priorities. You may want to seek professional help with the production of a management plan. BACKGROUND Bracken ( Pteridium aquilinum ) is an important and natural part of our landscape, and its abundance appears to have fluctuated over thousands of years. Its invasive nature allows it to spread, and it now occurs widely within a variety of habitats throughout Scotland. -
Glannau Aberdaron Ac Ynys Enlli/ Aberdaron Coast and Bardsey Island
STANDARD DATA FORM for sites within the ‘UK national site network of European sites’ Special Protection Areas (SPAs) are classified and Special Areas of Conservation (SACs) are designated under: • the Conservation of Habitats and Species Regulations 2017 (as amended) in England and Wales (including the adjacent territorial sea) and to a limited extent in Scotland (reserved matters) and Northern Ireland (excepted matters); • the Conservation (Natural Habitats &c.) Regulations 1994 (as amended) in Scotland; • the Conservation (Natural Habitats, &c) Regulations (Northern Ireland) 1995 (as amended) in Northern Ireland; and • the Conservation of Offshore Marine Habitats and Species Regulations 2017 (as amended) in the UK offshore area. Each SAC or SPA (forming part of the UK national site network of European sites) has its own Standard Data Form containing site-specific information. The information provided here generally follows the same documenting format for SACs and SPAs, as set out in the Official Journal of the European Union recording the Commission Implementing Decision of 11 July 2011 (2011/484/EU). Please note that these forms contain a number of codes, all of which are explained either within the data forms themselves or in the end notes. More general information on SPAs and SACs in the UK is available from the SPA homepage and SAC homepage on the JNCC website. These webpages also provide links to Standard Data Forms for all SAC and SPA sites in the UK. https://jncc.gov.uk/ 1 NATURA 2000 - STANDARD DATA FORM For Special Protection Areas (SPA), Proposed Sites for Community Importance (pSCI), Sites of Community Importance (SCI) and for Special Areas of Conservation (SAC) SITE UK9013121 SITENAME Glannau Aberdaron ac Ynys Enlli/ Aberdaron Coast and Bardsey Island TABLE OF CONTENTS 1. -
Pteridium: Dennstaedtiaceae) This Separation May Correspond to the Basal Divergence Among in Australia
American Journal of Botany 96(5): 1041–1049. 2009. G LOBAL CHLOROPLAST PHYLOGENY AND BIOGEOGRAPHY OF BRACKEN ( PTERIDIUM ; DENNSTAEDTIACEAE) 1 Joshua P. Der, 2,4 John A. Thomson, 3 Jeran K. Stratford, 2,5 and Paul G. Wolf 2 2 Department of Biology, Utah State University, 5305 Old Main Hill, Logan, Utah 84322 USA; and 3 National Herbarium of New South Wales, Botanic Gardens Trust, Mrs Macquaries Road, Sydney, NSW 2000, Australia Bracken ferns (genus Pteridium ) represent an ancient species complex with a natural worldwide distribution. Pteridium has historically been treated as comprising a single species, but recent treatments have recognized several related species. Phenotypic plasticity, geographically structured morphological variation, and geographically biased sampling have all contributed to taxo- nomic confusion in the genus. We sampled bracken specimens worldwide and used variable regions of the chloroplast genome to investigate phylogeography and reticulate evolution within the genus. Our results distinguish two major clades within Pteridium , a primarily northern hemisphere Laurasian/African clade, which includes all taxa currently assigned to P. aquilinum , and a primar- ily southern hemisphere Austral/South American clade, which includes P. esculentum and P. arachnoideum . All European acces- sions of P. aquilinum subsp. aquilinum appear in a monophyletic group and are nested within a clade containing the African P. aquilinum taxa ( P. aquilinum subsp. capense and P. aquilinum subsp. centrali-africanum ). Our results allow us to hypothesize the maternal progenitors of two allotetraploid bracken species, P. caudatum and P. semihastatum . We also discuss the biogeography of bracken in the context of the chloroplast phylogeny. Our study is one of the fi rst to take a worldwide perspective in addressing variation in a broadly distributed species complex. -
News Sheet N 20
Herefordshire Fungus Survey Group o News Sheet N 20: Autumn 2010 Agrocybe rivulosa – Cwmdu, Powys (August 2010) Contents Recorder’s Report, September - December 2008 Page 3 Forget the Weather – Foray in an Art Gallery! Page 5 What’s in a Name? Page 6 Mountains are not just for Walking Page 9 Small Fan-like Fungi Page 11 Bracken and Fungi – an Introduction Page 13 The contents of this newsletter are the copyright property of the Herefordshire Fungus Survey Group. Please do not reproduce material from this publication without prior permission from the Editor. Thank you. another look at the subject. With great erudition he President: Ted Blackwell explores the derivation of some of our common (and not so common!) Latin binomials. Recorder: Jo Weightman Jo has written this time on some of the small, fan-shaped Chairman: Roger Evans fungi, such as those of the Panellus and Melanotus species, that we are inclined to overlook. With her Secretary: Mike Stroud excellent descriptions and the photographs, we might hope that more of these will find their way into our lists in future. Treasurer: Margaret Hawkins One of our newer members, Gareth, reports on a ‘Bracken Welcome to the Autumn 2010 News Sheet Open Day’, given by the Society of Biology and the International Bracken Group in Breconshire. He highlights this important association with fungi and draws our Firstly, my humblest apologies for producing this News attention to the wealth of fungal species that occur in this Sheet so late in the Autumn. I have nobody to blame, environment. except myself! Debbie has told us of how she combines her two passions As you will see from the list of HFSG officers, our (or should it be ‘two of her passions’?) of mountain walking Treasurer has changed from Steve Rolph to Margaret and mycology. -
A Novel Method for Determination of the Natural Toxin Ptaquiloside in Ground and Drinking Water
water Article A Novel Method for Determination of the Natural Toxin Ptaquiloside in Ground and Drinking Water Natasa Skrbic 1,2,* , Ann-Katrin Pedersen 1, Sarah C. B. Christensen 1, Hans Christian Bruun Hansen 2 and Lars Holm Rasmussen 3 1 Greater Copenhagen Utility HOFOR, Parkstien 10, 2450 Copenhagen, Denmark; [email protected] (A.-K.P.); [email protected] (S.C.B.C.) 2 Department of Plant and Environmental Sciences, University of Copenhagen, Thorvaldsensvej 40, 1871 Frederiksberg, Denmark; [email protected] 3 Department of Technology, University College Copenhagen, Sigurdsgade 26, 2200 Copenhagen, Denmark; [email protected] * Correspondence: [email protected]; Tel.: +45-2795-4306 Received: 7 September 2020; Accepted: 8 October 2020; Published: 13 October 2020 Abstract: Ptaquiloside (PTA) is a carcinogenic compound naturally occurring in bracken ferns (Pteridium aquilinum). It is highly water soluble and prone to leaching from topsoil to surface and groundwaters. Due to possible human exposure via drinking water, PTA is considered as an emerging contaminant. We present a sensitive and robust method for analysis of PTA and its degradation product pterosin B (PtB) in groundwater. The method comprises two steps: sample preservation at the field site followed by sample pre-concentration in the laboratory. The preservation step was developed by applying a Plackett–Burman experimental design testing the following variables: water type, pH, filtering, bottle type, storage temperature, transportation conditions and test time. The best sample preservation was obtained by using amber glass bottles, unfiltered solutions buffered at pH 6, transported without ice, stored at 4 ◦C and analysed within 48 h. The recovery was 94% to 100%.