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Schall and Dearing.Pdf
Oecologia (Berlin) (1987) 73:389-392 LxA70u c Springer-Verlag 1987 Malarial parasitism and male competition for mates in the western fence lizard, Sceloporus occidentalis J.J. Schall and M.D. Dearing Department of Zoology, University of Vermont, Burlington, VT 05405, USA Summary. The effect of malarial parasitism on the ability the outcome of male-male interactions and female choice of male western fence lizards, Sceloporus occidentalis, to in the western fence lizard, Sceloporusoccidentalis, in Cali- compete for access to females was assessed experimentally. fornia, USA. The mating system of S. occidentalisis polyga- Pairs of male lizards, one infected with the malarial para- mous. Males interact agonistically toward other males and site, Plasmodium mexicanum, and the other not infected, vigorously court females during the reproductive season. were matched by size and color and placed in large semina- Their conspicuous stereotyped behaviors include bobbing, tural outdoor enclosures along with an adult female lizard. shaking, and display of brightly colored ventral color Infected males displayed to females and to other males less patches (Schall and Sarni 1987; Ressel 1986). At our study often than did noninfected male lizards. Noninfected lizards site approximately 25% of adult male fence lizards are in- were dominant in social interactionsmore often than malar- fected with the malarial parasite, Plasmodiummexicanum ious animals, based on duration and intensity of agonistic (Schall 1983; Bromwich and Schall 1986). This parasite re- encounters toward the other male, and time spent with the duces the ability of the fence lizard to engage in intense female. Thus, malarial infection hinders the ability of male activity; infected males also have smaller testes (Schall fence lizards to compete for mates. -
BIOLOGICAL ASSESSMENT for the LUCAS Creek PROJECT KERN
Biological Assessment- Lucas Creek Project BIOLOGICAL ASSESSMENT For the LUCAS Creek PROJECT KERN RIVER RANGER DISTRICT SEQUOIA NATIONAL FOREST Kern County, California PREPARED By:--J; �ATE: February 5, 2018 Nina Hemphi Forest Fish Biologist/Aquatic Ecologist and Watershed Manager This Biological Assessment analyzes the potential impacts associated with implementation of the Lucas Creek Project on federal endangered and threatened species as identified under the Endangered Species Act. The environmental analysis evaluates the preferred alternative. The Lucas Creek Project includes removal of dead and dying trees on 250 acres on Breckenridge Mountain. The project area is located in sections 23, 24, 25, & 26, township 28 south, range 31 east, Mount Diablo Base Meridian on the Kern River Ranger District of the Sequoia National Forest. The project surrounds the Breckenridge subdivision on Breckenridge Mountain approximately 25 miles southwest of the town of Lake Isabella in Kern County California. The intent of the Lucas Creek Project is to remove hazard trees along roads and properties adjoining the Breckenridge Subdivision. The project would also reduce fuels build-up to protect the community and the Lucas Creek upper and middle watershed from high-intensity fire. This will improve forest resilience and watershed health. This document is prepared in compliance with the requirements of FSM 2672.4 and 36 CFR 219.19. Biological Assessment- Lucas Creek Project I. INTRODUCTION The purpose of this Biological Assessment (BA) is to review the potential effects of Lucas Creek Project on species classified as federally endangered and threatened under the Endangered Species Act (ESA, 1973). Federally listed species are managed under the authority of the Endangered Species Act (ESA) and the National Forest Management Act (NFMA; PL 94- 588). -
California Wildlife Habitat Relationships System California Department of Fish and Wildlife California Interagency Wildlife Task Group
California Wildlife Habitat Relationships System California Department of Fish and Wildlife California Interagency Wildlife Task Group FAIRVIEW SLENDER SALAMANDER Batrachoseps bramei Family: PLETHODONTIDAE Order: CAUDATA Class: AMPHIBIA A073 Written by: T. Kucera, 1997 Updated by: CWHR Staff May 2013 DISTRIBUTION, ABUNDANCE, AND SEASONALITY Yearlong resident in the western slope of the southern Sierra Nevada. Individuals occur along streams and in moist wooded canyons in valley foothill riparian habitats, blue oak woodlands, and Sierra mixed conifer woodlands (Yanev 1978). Brame and Murray (1968) included salamanders from four disjunct regions, including the southern Sierra Nevada, in B. pacificus. Jennings and Hayes (1994) elevated the animals from the southern Sierra Nevada to specific status (B. relictus). Then, on the basis of DNA analyses (Jockusch 1996, Jockusch et al. 1998), the B. relictus complex was split out into four distinct species: B. relictus, B. regius, B. kawia and B. diabolicus. Jockusch et al. (2012) documented morphological and molecular data to support the recognition of B. bramei as a species distinct from B. relictus. SPECIFIC HABITAT REQUIREMENTS Feeding: Feeding probably occurs both above and below ground (Hendrickson 1954). Stebbins (1951) reported that a similar species, the pacific slender salamander (B. pacficus), fed on earthworms, small slugs, a variety of terrestrial arthropods including sowbugs and millipedes, and insects including collembolans, aphids, caterpillars, small beetles, beetle larvae, and ants. The fairview slender salamander probably eats a similar array of prey items. Cover: Members of the genus Batrachoseps do not usually excavate burrows. They rely on passages made by other animals, or produced by root decay or soil shrinkage (Yanev 1978). -
Ventral Coloration and Body Condition Do Not Affect Territorial Behavior in Two Sceloporus Lizards
Ventral coloration and body condition do not affect territorial behavior in two Sceloporus lizards 1Marina Kelada, 1Courtney Moulton, 2Casey Nguyen, 3Griselda Robles Olague 1University of California, Riverside; 2University of California, Irvine; 3University of California, Santa Barbara Sceloporus lizards are known to be defensive of their territories, which provide shelter, food, water, and mates. They often exhibit territorial behavior through visual displays of aggression, specifically with pushups. Along with aggression, this display can also expose the blue coloration on their ventral side, serving as an intraspecific communication between lizards that reveals information about their dominance, sex, and species membership. In this study, we compared how blue coloration and body condition affected territorial behavior in two Sceloporus lizards: the well-studied western fence (Sceloporus occidentalis) and the understudied sagebrush (Sceloporus graciosus), two closely related lizards with overlapping ranges in the San Jacinto Mountains. We also addressed whether these species would exhibit character displacement in their ventral coloration as a result of their sympatry. ImageJ and Adobe Photoshop were used for photo analyses to quantify the proportion and intensity of blue on each lizard’s ventral side. Territoriality was measured through number of pushups displayed and distance traveled. We found that both ventral coloration and body condition had no effect on the territoriality of either species, suggesting that territorial behavior -
Sensitive Species of Snakes, Frogs, and Salamanders in Southern California Conifer Forest Areas: Status and Management1
Sensitive Species of Snakes, Frogs, and Salamanders in Southern California Conifer Forest Areas: Status and Management1 Glenn R. Stewart2, Mark R. Jennings3, and Robert H. Goodman, Jr.4 Abstract At least 35 species of amphibians and reptiles occur regularly in the conifer forest areas of southern California. Twelve of them have some or all of their populations identified as experiencing some degree of threat. Among the snakes, frogs, and salamanders that we believe need particular attention are the southern rubber boa (Charina bottae umbratica), San Bernardino mountain kingsnake (Lampropeltis zonata parvirubra), San Diego mountain kingsnake (L.z. pulchra), California red-legged frog (Rana aurora draytonii), mountain yellow-legged frog (R. muscosa), San Gabriel Mountain slender salamander (Batrachoseps gabrieli), yellow-blotched salamander (Ensatina eschscholtzii croceater), and large-blotched salamander (E.e. klauberi). To varying degrees, these taxa face threats of habitat degradation and fragmentation, as well as a multitude of other impacts ranging from predation by alien species and human collectors to reduced genetic diversity and chance environmental catastrophes. Except for the recently described San Gabriel Mountain slender salamander, all of these focus taxa are included on Federal and/or State lists of endangered, threatened, or special concern species. Those not federally listed as Endangered or Threatened are listed as Forest Service Region 5 Sensitive Species. All of these taxa also are the subjects of recent and ongoing phylogeographic studies, and they are of continuing interest to biologists studying the evolutionary processes that shape modern species of terrestrial vertebrates. Current information on their taxonomy, distribution, habits and problems is briefly reviewed and management recommendations are made. -
New Species of Slender Salamander, Genus Batrachoseps, from the Southern Sierra Nevada of California
Copeia, 2002(4), pp. 1016±1028 New Species of Slender Salamander, Genus Batrachoseps, from the Southern Sierra Nevada of California DAVID B. WAKE,KAY P. Y ANEV, AND ROBERT W. HANSEN Populations of robust salamanders belonging to the plethodontid salamander ge- nus Batrachoseps (subgenus Plethopsis) from the southern Sierra Nevada and adjacent regions represent a previously unknown species here described as Batrachoseps ro- bustus. The new species is robust, with a short trunk (17±18 trunk vertebrae) and well-developed limbs. It differs from its close geographic neighbor, Batrachoseps campi, in lacking patches of dorsal silvery iridophores and in having (typically) a lightly pigmented dorsal stripe, and from Batrachoseps wrighti in being more robust, having more trunk vertebrae, and in lacking conspicuous white spots ventrally. This species is widely distributed on the semiarid Kern Plateau of the southeastern Sierra Nevada and extends along the east slopes of the mountains into the lower Owens Valley; it also is found to the south in the isolated Scodie Mountains. It occurs at high elevations, from 1615±2800 m, in areas of low rainfall and high summer tem- peratures. HE slender salamanders, Batrachoseps, range ci®c. They occur in relatively extreme environ- T from the Columbia River in northern ments for terrestrial salamanders, in areas that Oregon (458339N) to the vicinity of El Rosario, are exceptionally hot and dry in the summer, Baja California Norte (308009N). These sala- and that have a short, unpredictable wet season manders were considered to display mainly in- with little rainfall. Some of them occur at high traspeci®c morphological variation, and for elevations that are under snow cover for several many years only two (Hendrickson, 1954) or months each year. -
Biology 2 Lab Packet for Practical 4
1 Biology 2 Lab Packet For Practical 4 2 CLASSIFICATION: Domain: Eukarya Supergroup: Unikonta Clade: Opisthokonts Kingdom: Animalia Phylum: Chordata – Chordates Subphylum: Urochordata - Tunicates Class: Amphibia – Amphibians Subphylum: Cephalochordata - Lancelets Order: Urodela - Salamanders Subphylum: Vertebrata – Vertebrates Order: Apodans - Caecilians Superclass: Agnatha Order: Anurans – Frogs/Toads Order: Myxiniformes – Hagfish Class: Testudines – Turtles Order: Petromyzontiformes – Lamprey Class: Sphenodontia – Tuataras Superclass: Gnathostomata – Jawed Vertebrates Class: Squamata – Lizards/Snakes Class: Chondrichthyes - Cartilaginous Fish Lizards Subclass: Elasmobranchii – Sharks, Skates and Rays Order: Lamniiformes – Great White Sharks Family – Agamidae – Old World Lizards Order: Carcharhiniformes – Ground Sharks Family – Anguidae – Glass Lizards Order: Orectolobiniformes – Whale Sharks Family – Chameleonidae – Chameleons Order: Rajiiformes – Skates Family – Corytophanidae – Helmet Lizards Order: Myliobatiformes - Rays Family - Crotaphytidae – Collared Lizards Subclass: Holocephali – Ratfish Family – Helodermatidae – Gila monster Order: Chimaeriformes - Chimaeras Family – Iguanidae – Iguanids Class: Sarcopterygii – Lobe-finned fish Family – Phrynosomatidae – NA Spiny Lizards Subclass: Actinistia - Coelocanths Family – Polychrotidae – Anoles Subclass: Dipnoi – Lungfish Family – Geckonidae – Geckos Class: Actinopterygii – Ray-finned Fish Family – Scincidae – Skinks Order: Acipenseriformes – Sturgeon, Paddlefish Family – Anniellidae -
Jockusch Et Al in 2012
Zootaxa 3190: 1–30 (2012) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2012 · Magnolia Press ISSN 1175-5334 (online edition) Morphological and molecular diversification of slender salamanders (Caudata: Plethodontidae: Batrachoseps) in the southern Sierra Nevada of California with descriptions of two new species ELIZABETH L. JOCKUSCH1, IÑIGO MARTÍNEZ-SOLANO1,2, ROBERT W. HANSEN3, & DAVID B. WAKE4 1Department of Ecology and Evolutionary Biology, 75 N. Eagleville Rd., U-3043, University of Connecticut, Storrs, CT 06269, USA. E-mail: [email protected] 2Instituto de Investigación en Recursos Cinegéticos (IREC) (CSIC-UCLM-JCCM), Ronda de Toledo, s/n 13005 Ciudad Real, Spain. E-mail: [email protected] 316333 Deer Path Lane, Clovis, CA 93619-9735, USA. E-mail: [email protected] 4Museum of Vertebrate Zoology, 3101 Valley Life Sciences Building, University of California, Berkeley, CA 94720-3160, USA. E-mail: [email protected] Abstract Slender salamanders of the genus Batrachoseps achieve relatively high diversity in the Kern Canyon region at the southern end of the Sierra Nevada of California through high turnover of species with small geographic ranges. The status of several populations of Batrachoseps in this region is enigmatic, and both morphological and molecular data have suggested that some populations do not belong to any of the currently recognized species. Identification of species in this region is com- plicated by the apparent extinction of Batrachoseps relictus in the vicinity of its type locality in the Lower Kern River Canyon. Here we analyze a comprehensive morphological dataset to evaluate diversity in the Kern River Canyon region. -
1 CWU Comparative Osteology Collection, List of Specimens
CWU Comparative Osteology Collection, List of Specimens List updated November 2019 0-CWU-Collection-List.docx Specimens collected primarily from North American mid-continent and coastal Alaska for zooarchaeological research and teaching purposes. Curated at the Zooarchaeology Laboratory, Department of Anthropology, Central Washington University, under the direction of Dr. Pat Lubinski, [email protected]. Facility is located in Dean Hall Room 222 at CWU’s campus in Ellensburg, Washington. Numbers on right margin provide a count of complete or near-complete specimens in the collection. Specimens on loan from other institutions are not listed. There may also be a listing of mount (commercially mounted articulated skeletons), part (partial skeletons), skull (skulls), or * (in freezer but not yet processed). Vertebrate specimens in taxonomic order, then invertebrates. Taxonomy follows the Integrated Taxonomic Information System online (www.itis.gov) as of June 2016 unless otherwise noted. VERTEBRATES: Phylum Chordata, Class Petromyzontida (lampreys) Order Petromyzontiformes Family Petromyzontidae: Pacific lamprey ............................................................. Entosphenus tridentatus.................................... 1 Phylum Chordata, Class Chondrichthyes (cartilaginous fishes) unidentified shark teeth ........................................................ ........................................................................... 3 Order Squaliformes Family Squalidae Spiny dogfish ........................................................ -
Shedd, Jackson, 2009: Bilateral Asymmetry in Two Secondary
BILATERAL ASYMMETRY IN TWO SECONDARY SEXUAL CHARACTERS IN THE WESTERN FENCE LIZARD (SCELOPORUS OCCIDENTALIS): IMPLICATIONS FOR A CORRELATION WITH LATERALIZED AGGRESSION ____________ A Thesis Presented to the Faculty of California State University, Chico ____________ In Partial Fulfillment of the Requirements for the Degree Master of Science in Biological Sciences ____________ by Jackson D. Shedd Spring 2009 BILATERAL ASYMMETRY IN TWO SECONDARY SEXUAL CHARACTERS IN THE WESTERN FENCE LIZARD (SCELOPORUS OCCIDENTALIS): IMPLICATIONS FOR A CORRELATION WITH LATERALIZED AGGRESSION A Thesis by Jackson D. Shedd Spring 2009 APPROVED BY THE DEAN OF THE SCHOOL OF GRADUATE, INTERNATIONAL, AND INTERDISCIPLINARY STUDIES: _________________________________ Susan E. Place, Ph.D. APPROVED BY THE GRADUATE ADVISORY COMMITTEE: _________________________________ _________________________________ Abdel-Moaty M. Fayek Tag N. Engstrom, Ph.D., Chair Graduate Coordinator _________________________________ Donald G. Miller, Ph.D. _________________________________ Raymond J. Bogiatto, M.S. DEDICATION To Mela iii ACKNOWLEDGMENTS This research was conducted under Scientific Collecting Permit #803021-02 granted by the California Department of Fish and Game. For volunteering their time and ideas in the field, I thank Heather Bowen, Dr. Tag Engstrom, Dawn Garcia, Melisa Garcia, Meghan Gilbart, Mark Lynch, Colleen Martin, Julie Nelson, Michelle Ocken, Eric Olson, and John Rowden. Thank you to Brian Taylor for providing the magnified photographs of femoral pores. Thank you to Brad Stovall for extended cell phone use in the Mojave Desert while completing the last hiccups with this project. Thank you to Nuria Polo-Cavia and Dr. Nancy Carter for assistance and noticeable willingness to help with statistical analysis. Thank you to Dr. Diana Hews for providing direction for abdominal patch measurements and quantification. -
Testing the Persistence of Phenotypic Plasticity After Incubation in the Western Fence Lizard, Sceloporus Occidentalis Christine R
Claremont Colleges Scholarship @ Claremont All HMC Faculty Publications and Research HMC Faculty Scholarship 1-1-2007 Testing the Persistence of Phenotypic Plasticity After Incubation in the Western Fence Lizard, Sceloporus Occidentalis Christine R. Buckley Maddalena Jackson Mina Youssef Duncan J. Irschick University of Massachusetts - Amherst Stephen C. Adolph Harvey Mudd College Recommended Citation Buckley, C. R., M. Jackson, M. Youssef, D. J. Irschick, and S. C. Adolph. "Testing the persistence of phenotypic plasticity after incubation in the Western fence lizard, Sceloporus occidentalis." Evolutionary Ecology Research 9.1 (2007): 169-183. This Article is brought to you for free and open access by the HMC Faculty Scholarship at Scholarship @ Claremont. It has been accepted for inclusion in All HMC Faculty Publications and Research by an authorized administrator of Scholarship @ Claremont. For more information, please contact [email protected]. Evolutionary Ecology Research, 2007, 9: 169–183 Testing the persistence of phenotypic plasticity after incubation in the western fence lizard, Sceloporus occidentalis Christine R. Buckley,1* Maddalena Jackson,2 Mina Youssef,2 Duncan J. Irschick1* and Stephen C. Adolph2 1Department of Ecology and Evolutionary Biology, Tulane University, 310 Dinwiddie Hall, New Orleans, LA 70118 and 2Department of Biology, Harvey Mudd College, 1250 N. Dartmouth Avenue, Claremont, CA 91711, USA ABSTRACT Hypothesis: Phenotypic variation in traits induced by different incubation temperatures does not persist into the lifetime of young lizards, and therefore contributes little to variation in long-term fitness. Organism: Western fence lizard (Sceloporus occidentalis). Methods: Split-clutch laboratory incubation experiment including eggs from two different populations under two different incubation regimes, measurement of morphological traits at hatching, and tracking of morphology and temperature preference behaviour for 7 weeks after hatching. -
Dunes Sagebrush Lizard Petition
1 May 8, 2018 Mr. Ryan Zinke Secretary of the Interior Office of the Secretary Department of the Interior 18th and “C” Street, N.W. Washington DC 20202 Subject: Petition to List the Dunes Sagebrush Lizard as a Threatened or Endangered Species and Designate Critical Habitat Dear Secretary Zinke: The Center for Biological Diversity and Defenders of Wildlife hereby formally petition to list the dunes sagebrush lizard (Sceloperus arenicolus) as a threatened or endangered species under the Endangered Species Act of 1973, as amended (16 U.S.C. 1531 et seq.). This petition is filed under 5 U.S.C. § 553(e) and 50 C.F.R. § 424.14, which grant interested parties the right to petition for the issuance of a rule from the Assistant Secretary of the Interior. The Petitioners also request that critical habitat be designated for S. arenicolus concurrent with the listing, as required by 16 U.S.C. § 1533(b)(6)(C) and 50 C.F.R. § 424.12, and pursuant to the Administrative Procedures Act (5 U.S.C. § 553). The Petitioners understand that this petition sets in motion a specific process, placing defined response requirements on the U.S. Fish and Wildlife Service and specific time constraints on those responses. See 16 U.S.C. § 1533(b). Petitioners The Center for Biological Diversity is a national, non-profit conservation organization with more than 1.6 million members and online activists dedicated to protecting diverse native species and habitats through science, policy, education, and the law. It has offices in 11 states and Mexico.