Ground Truthing Tip-Dating Methods Using Fossil Canidae Reveals Major
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Shape Evolution and Sexual Dimorphism in the Mandible of the Dire Wolf, Canis Dirus, at Rancho La Brea Alexandria L
Marshall University Marshall Digital Scholar Theses, Dissertations and Capstones 2014 Shape evolution and sexual dimorphism in the mandible of the dire wolf, Canis Dirus, at Rancho la Brea Alexandria L. Brannick [email protected] Follow this and additional works at: http://mds.marshall.edu/etd Part of the Animal Sciences Commons, and the Paleontology Commons Recommended Citation Brannick, Alexandria L., "Shape evolution and sexual dimorphism in the mandible of the dire wolf, Canis Dirus, at Rancho la Brea" (2014). Theses, Dissertations and Capstones. Paper 804. This Thesis is brought to you for free and open access by Marshall Digital Scholar. It has been accepted for inclusion in Theses, Dissertations and Capstones by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected]. SHAPE EVOLUTION AND SEXUAL DIMORPHISM IN THE MANDIBLE OF THE DIRE WOLF, CANIS DIRUS, AT RANCHO LA BREA A thesis submitted to the Graduate College of Marshall University In partial fulfillment of the requirements for the degree of Master of Science in Biological Sciences by Alexandria L. Brannick Approved by Dr. F. Robin O’Keefe, Committee Chairperson Dr. Julie Meachen Dr. Paul Constantino Marshall University May 2014 ©2014 Alexandria L. Brannick ALL RIGHTS RESERVED ii ACKNOWLEDGEMENTS I thank my advisor, Dr. F. Robin O’Keefe, for all of his help with this project, the many scientific opportunities he has given me, and his guidance throughout my graduate education. I thank Dr. Julie Meachen for her help with collecting data from the Page Museum, her insight and advice, as well as her support. I learned so much from Dr. -
Estrategias De Subsistencia De Los Primeros Grupos Humanos Que Poblaron Europa
ESTRATEGIAS DE SUBSISTENCIA DE LOS PRIMEROS GRUPOS HUMANOS QUE POBLARON EUROPA ESTRATEGIAS DE SUBSISTENCIA DE LOS PRIMEROS GRUPOS HUMANOS QUE POBLARON EUROPA: EVIDENCIAS CONSERVADAS EN BARRANCO LEÓN Y FUENTE NUEVA-3 (ORCE) M. PATROCINIO ESPIGARES* RESUMEN Varios yacimientos del Pleistoceno inferior de España, Francia e Italia preservan las evidencias de presencia humana más antiguas de Europa. En este contexto, son particularmente interesantes dos localidades ubicadas en las inmediaciones de la villa de Orce (Cuenca de Baza, Granada), Barranco León (BL) y Fuente Nueva-3 (FN-3), datadas en torno a 1,4 Ma. En estos yacimientos se han identificado evidencias de procesado de cadáveres de grandes mamíferos, realizado con herra- mientas líticas de factura Olduvayense. A estos hallazgos hay que sumarle la presencia de un diente de leche atribuido a Homo sp. en Barranco León. En este trabajo se describen en detalle las marcas de origen antrópico localizadas en estos yacimientos, se analizan los patrones de procesado de los cadáveres, y se discute sobre las estrategias de subsistencia de las primeras comunidades humanas que habitaron Europa. Palabras clave: Marcas de corte, Estrategias de subsistencia, Pleistoceno Inferior, Homo sp. ABSTRACT Several Early Pleistocene sites from Spain, France and Italy preserve ancient evidence of human presence. In this context are particularly interesting two localities placed near the town of Orce (Baza Basin, Granada), Barranco León (BL) and Fuente Nueva-3 (FN-3), dated to ~1.4 Ma. At these sites, evidence of processing of large mammal carcasses produced with Oldowan tools have been recovered. These findings are accompanied by the presence of a deciduous tooth, attributed to Homo sp., in Barranco León. -
University of Florida Thesis Or Dissertation Formatting
UNDERSTANDING CARNIVORAN ECOMORPHOLOGY THROUGH DEEP TIME, WITH A CASE STUDY DURING THE CAT-GAP OF FLORIDA By SHARON ELIZABETH HOLTE A DISSERTATION PRESENTED TO THE GRADUATE SCHOOL OF THE UNIVERSITY OF FLORIDA IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF DOCTOR OF PHILOSOPHY UNIVERSITY OF FLORIDA 2018 © 2018 Sharon Elizabeth Holte To Dr. Larry, thank you ACKNOWLEDGMENTS I would like to thank my family for encouraging me to pursue my interests. They have always believed in me and never doubted that I would reach my goals. I am eternally grateful to my mentors, Dr. Jim Mead and the late Dr. Larry Agenbroad, who have shaped me as a paleontologist and have provided me to the strength and knowledge to continue to grow as a scientist. I would like to thank my colleagues from the Florida Museum of Natural History who provided insight and open discussion on my research. In particular, I would like to thank Dr. Aldo Rincon for his help in researching procyonids. I am so grateful to Dr. Anne-Claire Fabre; without her understanding of R and knowledge of 3D morphometrics this project would have been an immense struggle. I would also to thank Rachel Short for the late-night work sessions and discussions. I am extremely grateful to my advisor Dr. David Steadman for his comments, feedback, and guidance through my time here at the University of Florida. I also thank my committee, Dr. Bruce MacFadden, Dr. Jon Bloch, Dr. Elizabeth Screaton, for their feedback and encouragement. I am grateful to the geosciences department at East Tennessee State University, the American Museum of Natural History, and the Museum of Comparative Zoology at Harvard for the loans of specimens. -
Rodentia, Mammalia) in Europe'
Resumen EI yacimiento de Trinchera Oolina es uno de los mas importantes de la Sierra de Atapuerca par su contenido en restos humanos, Homo antecessor en el nivel de Trinchera Dolina 6. Ade mas de esto la enorme riqueza en restos arqueol6gicos y paleontol6gicos hace de Trinchera Dolina un yacimiento unico, de referencia obligada para el Pleistoceno y Paleolitico de Euro pa. Bioestratigraficamente, el yacimiento de Trinchera Oolina (TO) puede dividirse en tres grandes unidades: la que comprende los niveles TD3 a T06; la de los niveles T07 a TOS infe rior y I. de TO 8 superior. T011. Palabras clave: Mamiferos, Pleistoceno. Abstract Gran Dolina is one of the Pleistocene sites located at the Sierra de Atapuerca (Spain). The Gran Dolina deposits belong to different chronological periods of the Early and Middle Pleistocene. The uppermost levels of Gran Dolina (TDII, TDIO and TD8b) contain Middle Pleistocene (post-Cromerian) macro- and micromammal assemblages. The excavation works have overpassed level TDII and have not concluded yet at TDIO: TDII is poor in macromammal remains (carnivores and herbivores) but rich in rodents. The macromammals fossil material from TDtt is very scarce and this enables (for the macromammals) definite conclusions about the chronology and type of community of these levels. The lowermost levels of Gran Dolina (TD3/4, TD5, TD6 and TD8a) contain a different mammal assemblage with typical late Early Pleistocene-Cromerian species. This radical substitution of taxa is placed at TD8 layer probably due to a stratigraphic gap in this level. The level TD6 of the Gran Dolina site contains the earliest fossil human remains of Europe, Homo antecessor, and it has also a rich and diverse micromammal assemblage. -
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1.08 1.19 1.46 Nimravus brachyops Nandinia binotata Neofelis nebulosa 115 Panthera onca 111 114 Panthera atrox 113 Uncia uncia 116 Panthera leo 112 Panthera pardus Panthera tigris Lynx issiodorensis 220 Lynx rufus 221 Lynx pardinus 222 223 Lynx canadensis Lynx lynx 119 Acinonyx jubatus 110 225 226 Puma concolor Puma yagouaroundi 224 Felis nigripes 228 Felis chaus 229 Felis margarita 118 330 227 331Felis catus Felis silvestris 332 Otocolobus manul Prionailurus bengalensis Felis rexroadensis 99 117 334 335 Leopardus pardalis 44 333 Leopardus wiedii 336 Leopardus geoffroyi Leopardus tigrinus 337 Pardofelis marmorata Pardofelis temminckii 440 Pseudaelurus intrepidus Pseudaelurus stouti 88 339 Nimravides pedionomus 442 443 Nimravides galiani 22 338 441 Nimravides thinobates Pseudaelurus marshi Pseudaelurus validus 446 Machairodus alberdiae 77 Machairodus coloradensis 445 Homotherium serum 447 444 448 Smilodon fatalis Smilodon gracilis 66 Pseudaelurus quadridentatus Barbourofelis morrisi 449 Barbourofelis whitfordi 550 551 Barbourofelis fricki Barbourofelis loveorum Stenogale Hemigalus derbyanus 554 555 Arctictis binturong 55 Paradoxurus hermaphroditus Genetta victoriae 553 558 Genetta maculata 559 557 660 Genetta genetta Genetta servalina Poiana richardsonii 556 Civettictis civetta 662 Viverra tangalunga 661 663 552 Viverra zibetha Viverricula indica Crocuta crocuta 666 667 Hyaena brunnea 665 Hyaena hyaena Proteles cristata Fossa fossana 664 669 770 Cryptoprocta ferox Salanoia concolor 668 772 Crossarchus alexandri 33 Suricata suricatta 775 -
Large Mammal Biochronology Framework in Europe at Jaramillo: the Epivillafranchian As a Formal Biochron
Quaternary International 389 (2015) 84e89 Contents lists available at ScienceDirect Quaternary International journal homepage: www.elsevier.com/locate/quaint Large mammal biochronology framework in Europe at Jaramillo: The Epivillafranchian as a formal biochron Luca Bellucci a, Raffaele Sardella a, Lorenzo Rook b, * a Dipartimento di Scienze della Terra, “Sapienza e Universita di Roma”, P.le A. Moro 5, 00185, Roma, Italy b Dipartimento di Scienze della Terra, Universita di Firenze, via G. La Pira 4, 50121, Firenze, Italy article info abstract Article history: European large mammal assemblages in the 1.2e0.9 Ma timespan included Villafranchian taxa together Available online 3 December 2014 with newcomers, mostly from Asia, persisting in the Middle Pleistocene. A number of biochronological schemes have been suggested to define these “transitional” faunas. The term Epivillafranchian, originally Keywords: proposed by Bourdier in 1961 and reconsidered as a biochron by Kahlke in the 1990s, is at present widely Biochronology introduced in the literature. This contribution, after selecting the most representative European large Jaramillo mammal assemblages within this chronological interval, provides a new definition proposal for the Epivillafranchian Epivillafranchian as a biochron included within the Praemegaceros verticornis FO/Bison menneri FO, and Late Villafranchian Crocuta crocuta Galerian FO. Europe © 2014 Elsevier Ltd and INQUA. All rights reserved. 1. Historical background communities of this time span primarily include survivors from the latest Villafranchian, as well as more evolved taxa characteristic of The Villafranchian Mammal Age corresponds, in the Interna- the beginning Middle Pleistocene (Kahlke, 2007; Rook and tional Stratigraphic Scale, to a timespan from Late Pliocene to most Martinez Navarro, 2010). -
Canis Mosbachensis (Canidae, Mammalia) from the Middle Pleistocene of Contrada Monticelli (Putignano, Apulia, Southern Italy)
TO L O N O G E I L C A A P I ' T A A T L E I I A Bollettino della Società Paleontologica Italiana, 56 (1), 2017, 71-78. Modena C N O A S S. P. I. Canis mosbachensis (Canidae, Mammalia) from the Middle Pleistocene of Contrada Monticelli (Putignano, Apulia, southern Italy) Beniamino MECOZZI, Dawid Adam IURINO, Davide F. BERTÉ & Raffaele SARDELLA B. Mecozzi, Dipartimento di Scienze della Terra, Sapienza Università di Roma, Piazzale Aldo Moro 5, I-00185 Roma, Italy; PaleoFactory, Sapienza Università di Roma, Piazzale Aldo Moro 5, I-00185 Roma, Italy; [email protected] D.A. Iurino, Dipartimento di Scienze della Terra, Sapienza Università di Roma, Piazzale Aldo Moro 5, I-00185 Roma, Italy; PaleoFactory, Sapienza Università di Roma, Piazzale Aldo Moro 5, I-00185 Roma, Italy; [email protected] D.F. Berté, Associazione Culturale 3P (Progetto Preistoria Piemonte), Via Lunga 38, I-10099 San Mauro Torinese (Torino, Italy; [email protected] R. Sardella, Dipartimento di Scienze della Terra, Sapienza Università di Roma, Piazzale Aldo Moro 5, I-00185 Roma, Italy; PaleoFactory, Sapienza Università di Roma, Piazzale Aldo Moro 5, I-00185 Roma, Italy; [email protected] KEY WORDS - Carnivorans, Taxonomy, Biochronology, Paleobiogeography. ABSTRACT - Herein we describe for the first time a canid partial cranium from the Contrada Monticelli site. Morphological and biometrical studies allow the fossil remains to be referred to the Middle Pleistocene wolf Canis mosbachensis. Associated taxa include Paleoloxodon antiquus, Stephanorhinus hundsheimensis, cervids, equids and bovids, whose biochronological occurrence allows the site to be referred to the Galerian Mammal Age. -
The Early Hunting Dog from Dmanisi with Comments on the Social
www.nature.com/scientificreports OPEN The early hunting dog from Dmanisi with comments on the social behaviour in Canidae and hominins Saverio Bartolini‑Lucenti1,2*, Joan Madurell‑Malapeira3,4, Bienvenido Martínez‑Navarro5,6,7*, Paul Palmqvist8, David Lordkipanidze9,10 & Lorenzo Rook1 The renowned site of Dmanisi in Georgia, southern Caucasus (ca. 1.8 Ma) yielded the earliest direct evidence of hominin presence out of Africa. In this paper, we report on the frst record of a large‑sized canid from this site, namely dentognathic remains, referable to a young adult individual that displays hypercarnivorous features (e.g., the reduction of the m1 metaconid and entoconid) that allow us to include these specimens in the hypodigm of the late Early Pleistocene species Canis (Xenocyon) lycaonoides. Much fossil evidence suggests that this species was a cooperative pack‑hunter that, unlike other large‑sized canids, was capable of social care toward kin and non‑kin members of its group. This rather derived hypercarnivorous canid, which has an East Asian origin, shows one of its earliest records at Dmanisi in the Caucasus, at the gates of Europe. Interestingly, its dispersal from Asia to Europe and Africa followed a parallel route to that of hominins, but in the opposite direction. Hominins and hunting dogs, both recorded in Dmanisi at the beginning of their dispersal across the Old World, are the only two Early Pleistocene mammal species with proved altruistic behaviour towards their group members, an issue discussed over more than one century in evolutionary biology. Wild dogs are medium- to large-sized canids that possess several hypercarnivorous craniodental features and complex social and predatory behaviours (i.e., social hierarchic groups and pack-hunting of large vertebrate prey typically as large as or larger than themselves). -
Cultural Cannibalism As a Paleoeconomic System in The
539 Cannibalism is by definition the act of consuming tissues of Reports individuals of the same species, and it occurs among a wide variety of living organisms. From an ethological point of view, there are different mechanisms that determine this behavior. Cultural Cannibalism as a Paleoeconomic However, why humans process and consume other humans System in the European Lower Pleistocene is a complex question, and moving away from purely ethologic causes, the answer may encompass nutritional, economic, cos- The Case of Level TD6 of Gran Dolina (Sierra mogonic, social, and political purposes. Because these con- de Atapuerca, Burgos, Spain) ditions can sometimes intermingle, cannibalism must be viewed not as something unitary or simple (Sanday 1986) but Eudald Carbonell, Isabel Ca´ceres, Marina Lozano, rather as a complex activity that has some temporal conti- Palmira Saladie´, Jordi Rosell, Carlos Lorenzo, nuity. Josep Vallverdu´, Rosa Huguet, Antoni Canals, and Human cannibalism has traditionally given rise to extreme Jose´ Marı´a Bermu´dez de Castro feelings ranging from fascination to revulsion—attitudes often Institut Catala` de Paleoecologia Humana i Evolucio´ Social caused by ignorance or cultural bias. Denying the anthro- (IPHES), Unidad Asociada al Consejo Superior de pophagy is a response to the moral issue. Several scientific Investigaciones Cientı´ficas (CSIC). Universitat Rovira I researchers have even taken positions of denial with regard Virgili (URV), Campus Catalunya, Avinguda de Catalunya, to its existence (Arens 1979; Bahn 1992; Salas 1921). Can- 35, 43002 Tarragona, Spain (Carbonell, Ca´ceres, Lozano, nibalism revealed through oral and written sources has led to Saladie´, Rosell, Lorenzo, Vallverdu´ , Huguet, Canals) paradoxical situations that are explained only by traditions (icaceresprehistoria.urv.cat)/Visiting professor, Institute of and customs (Conklin 2001; Sanday 1986). -
(Cuenca De Guadix-Baza, Granada, España): Historia Y Presente
REVISTA DEL CEHGR · núm. 32 · 2020· págs. 23-45 ISSN: 2253-9263 Los yacimientos arqueopaleontológicos de la zona de Orce (cuenca de Guadix-Baza, Granada, España): historia y presente Carmen Luzón, Stefania Titton, Christian Sánchez Bandera, Juha Saarinen, Deborah Barsky, Hugues-A. Blain, Darío Estraviz, Suvi Viranta, Beatriz Azanza, Roberta Sanzi, José A. García Solano, Alexia Serrano Ramos, Daniel DeMiguel, José Yravedra Sainz de los Terreros, José Francisco Reinoso Gordo, Eva Montilla Jiménez, Juan José Rodríguez Alba, Auxiliadora Ruiz Domínguez, José Miguel Cámara Donoso, Oriol Oms, Jordi Agustí, Mikael Fortelius, Juan Manuel Jiménez Arenas Autor de correspondencia: Juan Manuel Jiménez-Arenas Universidad de Granada [email protected] RESUMEN En este trabajo se resumen los estudios llevados a cabo en los distintos yacimientos arqueopa- leontológicos de Orce durante las últimas décadas, haciendo especial hincapié en los resul- tados obtenidos a raíz de las últimas campañas de excavación (2017-2020), enmarcadas en el Proyecto General de Investigación «Primeras ocupaciones humanas y contexto paleoecológico a partir de los depósitos Pliopleistocenos de la cuenca Guadix-Baza. Zona Arqueológica de la cuenca de Orce». Los trabajos realizados en estos últimos años son una buena muestra del enfoque inter e intradisciplinar de la Prehistoria, y evidencian además que, aunque se lleve trabajando en los yacimientos de Orce casi 50 años, estos siguen proporcionando datos muy interesantes acerca del contexto del poblamiento humano de Europa en las etapas -
Dire Wolves Were the Last of an Ancient New World Canid Lineage Angela
Dire wolves were the last of an ancient New World canid lineage Angela R. Perri1,*§, Kieren J. Mitchell2,*§, Alice Mouton3,*, Sandra Álvarez-Carretero4,*, Ardern Hulme-Beaman5,6, James Haile 7, Alexandra Jamieson7, Julie Meachen8, Audrey T. Lin7,9,10, Blaine W. Schubert11, Carly Ameen12, Ekaterina E. Antipina13, Pere Bover14, Selina Brace15, Alberto Carmagnini4, Christian Carøe16, Jose A. Samaniego Castruita16, James C. Chatters17, Keith Dobney5, Mario dos Reis4, Allowen Evin18, Philippe Gaubert19, Shyam Gopalakrishnan16, Graham Gower2, Holly Heiniger2, Kristofer M. Helgen20, Josh Kapp21, Pavel A. Kosintsev22,23, Anna Linderholm7, 24, Andrew T. Ozga25, 26, 27, Samantha Presslee28, Alexander T. Salis2, Nedda F. Saremi21, Colin Shew3, Katherine Skerry26, Dmitry E. Taranenko29, Mary Thompson30, Mikhail V. Sablin31,Yaroslav V. Kuzmin32, 33, Matthew J. Collins34, 35, Mikkel-Holger S. Sinding16, 36, M. Thomas P. Gilbert16, 37, Anne C. Stone25 ,26, Beth Shapiro21, 38, Blaire Van Valkenburgh3, Robert K. Wayne3, Greger Larson7, and Alan Cooper39, Laurent A. F. Frantz4, 40§. 1Department of Archaeology, Durham University, Durham, UK 2Australian Centre for Ancient DNA, School of Biological Sciences, University of Adelaide, Australia 3Department of Ecology and Evolutionary Biology, University of California, Los Angeles, CA, USA 4School of Biological and Chemical Sciences, Queen Mary University of London, London, UK 5Department of Archaeology, Classics and Egyptology, University of Liverpool, Liverpool, UK 6School of Natural Sciences and Psychology, -
Human Origin Sites and the World Heritage Convention in Eurasia
World Heritage papers41 HEADWORLD HERITAGES 4 Human Origin Sites and the World Heritage Convention in Eurasia VOLUME I In support of UNESCO’s 70th Anniversary Celebrations United Nations [ Cultural Organization Human Origin Sites and the World Heritage Convention in Eurasia Nuria Sanz, Editor General Coordinator of HEADS Programme on Human Evolution HEADS 4 VOLUME I Published in 2015 by the United Nations Educational, Scientific and Cultural Organization, 7, place de Fontenoy, 75352 Paris 07 SP, France and the UNESCO Office in Mexico, Presidente Masaryk 526, Polanco, Miguel Hidalgo, 11550 Ciudad de Mexico, D.F., Mexico. © UNESCO 2015 ISBN 978-92-3-100107-9 This publication is available in Open Access under the Attribution-ShareAlike 3.0 IGO (CC-BY-SA 3.0 IGO) license (http://creativecommons.org/licenses/by-sa/3.0/igo/). By using the content of this publication, the users accept to be bound by the terms of use of the UNESCO Open Access Repository (http://www.unesco.org/open-access/terms-use-ccbysa-en). The designations employed and the presentation of material throughout this publication do not imply the expression of any opinion whatsoever on the part of UNESCO concerning the legal status of any country, territory, city or area or of its authorities, or concerning the delimitation of its frontiers or boundaries. The ideas and opinions expressed in this publication are those of the authors; they are not necessarily those of UNESCO and do not commit the Organization. Cover Photos: Top: Hohle Fels excavation. © Harry Vetter bottom (from left to right): Petroglyphs from Sikachi-Alyan rock art site.