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Downloaded from https://www.cambridge.org/core. IP address: 170.106.39.241, on 03 Oct 2021 at 23:52:42, subject to the Cambridge Core terms of use, available at https://www.cambridge.org/core/terms. https://doi.org/10.1017/S0025315419000092 Downloaded from https://www.cambridge.org/core. IP address: 170.106.39.241, on 03 Oct 2021 at 23:52:42, subject to the Cambridge Core terms of use, available at https://www.cambridge.org/core/terms. https://doi.org/10.1017/S0025315419000092 00253154_99-2_00253154_99-2 11/03/19 12:12 PM Page 2 JMBA THE MARINE BIOLOGICAL ASSOCIATION JOURNAL OF THE MARINE BIOLOGICAL ASSOCIATION OF THE UNITED KINGDOM OF THE UNITED KINGDOM TERMS OF MEMBERSHIP FOR 2019 EDITOR IN CHIEF: PROFESSOR JANE LEWIS MBA MEMBERSHIP Price from 1st March 2019 School of Life Sciences, University of Westminster, London, UK (prices per annum) EDITORIAL BOARD Young Marine Biologist (YMB) £12.50 M. COCK, Roscoff Marine Station, France C. HAUTON, University of Southampton, UK N. DE VOOGD, Netherlands Biodiversity Center, The Netherlands D. M. PATERSON, University of St Andrews, UK Student Member £25.00 P. G. H. EVANS, Sea Watch Foundation, UK D. POWER, University of the Algarve, Portugal M. FROST, Marine Biological Association of the United Kingdom, UK H. SOLO-GABRIELE, University of Miami, Florida, USA Associate Member £40.00 (direct debit discount £5) ASSOCIATE EDITORS Professional Member, Mem.MBA £45.00 (direct debit discount £5) P. A NDERWALD, Swiss National Park, Switzerland D.A. MCHUGH, Department of Biology, Colgate University, USA M. ARVEDLUND, University of Copenhagen, Denmark T. M OENS, University of Ghent, Belgium MBA Fellow, FMBA £120.00 (direct debit discount £5) J.A. BAEZA, Smithsonian Marine Station, USA P.D.R. MØLLER, Natural History Museum of Denmark C.N. BIANCHI, University of Genova, Italy P. J. PALSBØLL, GELIFES, University of Groningen, The Netherlands Institutional Member £340.00 J. CARTES, Institut de Ciències del Mar de Barcelona, Spain N.D. POPE, Marine Biological Association of the United Kingdom, UK B.K.K. CHAN, Biodiversity Research Centre, Academia Sinica, Taiwan N. QUEIROZ, CIBIO, Vairão, Portugal M. CUNLIFFE, Marine Biological Association of the United Kingdom, UK J.A. RAVEN, FRS, University of Dundee, UK JOURNAL SUBSCRIPTION J. ELLIS, Centre for Environment, Fisheries & Aquaculture Science (Cefas), UK M.M. RIVADENEIRA, Universidad Católica del Norte, Chile (MBA price added to the above membership fees, with the exception of YMB as no Journal option for this category) M.R. ELPHICK, Queen Mary University of London, UK J.M. RUIZ, Universidade da Coruña, Spain C. FRID, Griffith University, Australia M. RUSSELL, Villanova University, PA, USA Online Journal £20.00 N. HICKS, Scottish Association for Marine Science, UK M. STUMPP, Helmholtz Centre for Ocean Research Kiel, Germany J.G. HIDDINK, Bangor University, UK D. THIELTGES, NIOZ Royal Netherlands Institute for Sea Research; Print & Online Journal £80.00 M.Y. HU, University of Kiel, Germany Utrecht University, Netherlands D. JOLLIVET, Roscoff Marine Station, France P. T HOR, Norwegian Polar Institute, Norway W. J. L ANGSTON, Marine Biological Association of the United Kingdom, UK O. S. WANGENSTEEN, University of Barcelona, Spain The Association is a learned society founded in 1884 and from private members, universities, learned societies, the D. WILCOCKSON, Aberystwyth University, UK V. L APTIKHOVSKY, Falklands Islands Fisheries Department, UK incorporated by Royal Charter in 2013. The Marine Biological Fishmongers’ Company and other public bodies. M. LOUIS, University of St Andrews, UK N. WHITELEY, School of Biological Sciences, Bangor, UK Association (MBA) provides a unified, clear, independent voice on The Marine Biological Association is a very active scientific F. M ALTAGLIATI, University of Pisa, Italy behalf of the marine biological community. In 1888, the MBA learned society with a large international membership. The MBA opened the Plymouth laboratory at Citadel Hill. also manages an internationally renowned research programme. Address all correspondence to: Journal of the Marine Biological Association, The Laboratory, Citadel Hill, Plymouth, PL1 2PB, UK. (Tel: +44 (0) 1752 426493; Subscriptions can be paid by cash/cheque, direct debit from a e-mail: [email protected]) The MBA is also a registered charity with its key aims being to promote scientific research into all aspects of life in the sea, UK bank account or credit card. *Subscriptions are reduced by £5 per annum if paying by direct debit. EDITORIAL POLICY USA, Canada and Mexico, to Cambridge University Press, 1 Liberty Plaza, Floor 20, including the environment on which it depends, and to disseminate The Journal of the Marine Biological Association UK (JMBA) is an international New York, NY 10006, USA. EU subscribers (outside the UK) who are not registered for to the public the knowledge gained. The Patron of the Association Members of the Association have a wide range of rights and journal, publishing original research and reviews on all aspects of marine biology, to VAT should add VAT at their country’s rate. VAT registered subscribers should provide is His Royal Highness the Duke of Edinburgh, who is also an privileges including the use of post-nominals (FMBA for MBA support the aims of the MBA. The topics of present interest for JMBA are: their VAT registration number. Japanese prices for institutions are available from: Honorary Fellow. The other Honorary Fellows are His Serene Fellows, Mem.MBA for Professional Members) and being entitled • Marine ecology, behaviour and fisheries; Kinokuniya Company Ltd, PO Box 55, Chitose, Tokyo 156, Japan. Highness Prince Albert of Monaco, Dr Sylvia Earle, Sir Tim Hunt to receive The Marine Biologist Magazine. The Commissioners of Periodical postage paid at New York, NY, and at additional mailing offices. • Biodiversity and population studies of marine ecosystems, especially potential and Professor James Lovelock. Inland Revenue have approved the Association for the purposes of impacts of global warming, ocean acidification and climate change on ecosystem Postmaster: send address changes in USA, Canada and Mexico to Journal of the Marine Biological Association, Cambridge University Press, 100 Brook Hill Drive, West Nyack, New The work of the Association is controlled by the Council elected Section 16, Finance Act, 1958, and that the whole of the annual resilience; subscriptions paid by a member who qualifies for relief under the • Physiology, biochemistry, genetics and molecular biology of marine organisms, York 10994–2133, USA. annually by its subscribing members. including microbiology, particularly related to the integrative ecophysiology of Claims for missing issues should be made immediately on receipt of the subsequent The Marine Biological Association, under the direction of its section will be allowable as a deduction from his emoluments issue. marine organisms; Council, has undertaken research in all branches of marine science assessable to income tax. • Taxonomic syntheses, including molecular phylogenies; All correspondence should be addressed to: The Membership COPYRIGHT/PHOTOCOPYING • Morphology, life history and developmental biology of marine organisms, and many of the results have been published in this Journal. An The journal is registered with the Copyright Clearance Center, 222 Rosewood Secretary, Marine Biological Association, The Laboratory, Citadel especially model systems; account of the history of the first hundred years of the Association Drive, Danvers, MA 09123, USA. Organizations in the USA which are also registered Hill, Plymouth, PL1 2PB, UK (Tel. +44 (0) 1752 426493; e-mail: • Chemical and physical oceanography, as directly relevant to marine biology; and Journal may be found in Volume 67 (pp. 463 and 465). with CCC may therefore photocopy material (beyond the limits permitted by sections • Marine environmental health, and in particular its relationship to human health [email protected]). 107 and 108 of US copyright law) subject to payment to CCC of the per-copy fee of Since its foundation the Association has been supported by and well-being. $16.00. This consent does not extend to multiple copying for promotional or grants from Research Councils and by subscriptions and donations In addition to papers describing original research, the journal will also publish commercial purposes. Code 0025-3154/2019. short reviews and original viewpoints on topics of current marine biological interest. ISI Tear Sheet Service, 3501 Market Street, Philadelphia, PA 19104, USA, is authorized Papers describing work involving experimental procedures on living animals will to supply single photocopies of separate articles for private use only. Organizations be published only if they comply with the legal requirements of the UK Home Office. authorized by the UK Copyright Licensing Agency may also photocopy material subject PUBLISHED BY THE PRESS SYNDICATE OF THE UNIVERSITY OF CAMBRIDGE In the case of species not covered by the current law, papers will be published only if to the usual conditions. For all other use, permission should be sought from Cambridge or The Pitt Building, Trumpington Street, Cambridge, CB2 1RP, United Kingdom it is judged that the experiments have been conducted in a humane manner and the American branch of Cambridge University Press. Microfilm copies of back numbers adequate precautions taken to minimize stress. are available from University Microfilms International, 300 North Zeeb Road, MI This journal issue