Issn 0972- 1800

ISSN 0972- 1800

VOLUME 22, NO. 1 QUARTERLY JANUARY-MARCH, 2020

Date of Publication: 28th March, 2020

BIONOTES A Quarterly Newsletter for Research Notes and News On Any Aspect Related with Life Forms

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From Volume 21 Published by the Entomological Society of India (ESI), New Delhi (Nodal Officer: V.V. Ramamurthy, ESI, New Delhi) And Butterfly Research Centre, Bhimtal Executive Editor: Peter Smetacek Assistant Editor: Shristee Panthee Butterfly Research Trust, Bhimtal

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Cover Photo by Parixit Kafley of Samia canningi ejecting fluid from tip of abdomen. Vol.22 (1), March, 2020 BIONOTES

TABLE OF CONTENTS

SEVERE INFESTATION OF PODAGRICA FUSCICORNIS (CHEVROLAT, 1837) (CHRYSOMELIDAE) ON A NEW HOST PLANT ACALYPHA INDICA (L.) (EUPHORBIACEAE) FROM ODISHA, INDIA by Ashirwad Tripathy 2 SAMIA CANNINGI (INSECTA: LEPIDOPTERA: SATURNIIDAE) HAS A FUNCTIONAL PROBOSCIS AND ALIMENTARY CANAL by Parixit Kafley & Peter Smetacek 4 A NEW REPORT OF PARTIAL ALBINISM IN A HIMALAYAN BULBUL PYCNONOTUS LEUCOGENYS FROM UTTARAKHAND, INDIA by Paramjit Singh, Rajshekhar Singh, Devanshi Singh & Shankar Kumar 6 NEW RECORD OF ILLEIS INDICA TIMBERLAKE, 1943 (COLEOPTERA: COCCINELLIDAE) FROM ODISHA, INDIA by Ashirwad Tripathy 9 A COMPENDIUM ON MUSHROOM MITES IN INDIA by Reshma Parveen & Salil Kumar Gupta 11 FOUR NEW BUTTERFLY SPECIES FOR NEPAL: ABISARA CHELA, TAGIADES JAPETUS, LETHE DURA & LETHE DISTANS by Piet Van Der Poel, Colin Smith, Mahendra Singh Limbu & Surendra Pariyar 21 EDESSENA GENTIUSALIS (INSECTA: LEPIDOPTERA: EREBIDAE: HERMININAE): A NEW RECORD FOR INDIA by Shristee Panthee, Ambica Agnihotri & Peter Smetacek 24 FIRST RECORD OF JOKER BUTTERFLY BYBLIA ILITHYIA (INSECTA: LEPIDOPTERA: NYMPHALIDAE) FROM PAKISTAN by Muhammad Akram Awan, Wali Nohrio & Dileep Permar 26 CONFIRMATION OF THE EXTRA LASCAR PANTOPORIA SANDAKA IN ODISHA, INDIA by Sandeep Mishra & Daya Shanker Sharma 28 PRELIMINARY OBSERVATIONS ON VISITOR SPECTRUM OF RHODODENDRON ARBOREUM IN THE KUMAON HIMALAYA, INDIA by Ambica Agnihotri, Alfred Daniel & Piet Van Der Poel 29

Vol. 22 (1), March, 2020 BIONOTES A COMPENDIUM ON MUSHROOM MITES IN INDIA RESHMA PARVEEN1 & SALIL KUMAR GUPTA2 Medicinal Plants Research & Extension Centre, RK Mission, Narendrapur, Kolkata-700103 *[email protected] [email protected]

Reviewer: Peter Smetacek

Abstract cause economic loss to the mushroom The present compendium provides a list of growers. So far as mites occurring on mites occurring both on edible (4 spp.) and mushrooms are concerned, not much has been wild mushrooms (26 spp.), collected mainly explored from most parts of India. Some of the from West Bengal and a few from Kerala. A important publlications are Das (1986), total of 106 species, under 68 genera, 43 Somchoudhury et al. (1987) and Das et al. families and 4 orders are reported. Of those, (1987, 1987a, 1988, 1989, 1993). Gupta 64 species of mites are represented on edible (2012) provided summarized information of mushrooms and 57 species of mites are 17 species under 9 genera known till that time represented on wild mushrooms. This list from India. Thereafter, Gupta & Pal (2017), includes 6 spp. as likely new to science, to be Aiswarya et al. (2018), Parveen & Gupta described later, 4 spp. which are hitherto (2019, 2020), Mondal & Gupta (2019) unreported from India and 8 spp. as new provided additional information. Since most occurrence on mushrooms from India. All the of the available information regarding mites of species are arranged taxonomic category- mushrooms is scattered and not accessible to wise, giving information regarding their many, it was thought desirable to provide an relative abundance and the nature of updated list of mites on mushrooms in India association with both edible and wild giving all the information available till date. mushrooms. Out of 106 spp., 33 species cause This list includes 106 species and 68 genera damage, 56 are predators and 16 are of under 43 families and 4 orders from both unknown association. edible and wild mushrooms. It includes 6 Introduction species which appear to be new to science (to Mushrooms on one hand are important be described later), 4 species hitherto components for sustainability of ecosystems unreported from India and also 8 species, and, on the other hand, are of economic which were not known to occur on importance both for edible and medicinal mushrooms. Apart from listing these species, purposes. Some are parasitic and there are their relative abundance, the mushroom some which may prove to be fatal, if species on which they had been reported with consumed by human beings. Mushroom the nature of association have also been cultivation is gaining importance day by day provided. because of their manifold uses nowadays. Materials & Methods Many mushroom cultivators in West Bengal, The present compendium of mushroom mites especially women, are earning substantially is based mostly upon collections of mites on through selling them. both edible and wild mushrooms made by the Mushrooms, which can be broadly authors from West Bengal (Parveen & Gupta categorized under 2 groups viz. edible and 2019, 2020; Mondal & Gupta, 2019). Besides, wild, and both are attacked by pests which other published information available to the include insects, mites and nematodes that authors was also included on this list. 11 Vol. 22 (1), March, 2020 BIONOTES Results and Discussion families and 3 orders and all those were fungal Table-1 lists a total of 106 species of mites feeding in nature, belonging to Prostigmata belonging to 68 genera and 43 families under (Raphignathidae, Caligonellidae), 4 orders. Of those, 64 species, 51 genera, 28 Mesostigmata (Uropodidae, Resinacaridae) families, 3 orders have been reported on edible and Oribatida (Galumnidae, Ceratozetidae, mushrooms and the corresponding figures for Trhypochthoniidae, Oppiidae, Oribatulidae wild mushrooms were 57 species, 48 genera, and Austrachipteriidae). It may be mentioned 32 families and 4 orders, respectively. There here that oribatid mites were also predominant were many species which were recorded on in some mushrooms. both types of mushrooms. The present work Through a series of papers, Das (1986), Das et reports 6 species likely to be new viz. al. (1987, 1987a, 1989, 1993) contributed Typhlodromous-4 spp., Neocunaxoides- 1 sp., extensively to knowledge of the diversity of Cheylostigmaeus-1sp.. In addition, 4 species mushroom mites along with their bio-ecology (marked with a single asterisk in Table-1), viz. and control. Somchoudhury et al. through a Charletonia rocciai, Typhlodromous series of papers (1983-1989) also enriched our (Anthoseius) egypticus, Hypoaspis lubrica, knowledge on mushroom mites. Recently, Macrocheles glaber were not hitherto reported Aiswarya et al. (2018) reported 14 species from India and 8 species (indicated by double under 18 genera, 12 families, and 3 orders asterisks in Table-1) were not reported to mostly on wild mushroom from Kerala, while occur on mushrooms in India. Mondal & Gupta (2019) reported 12 species Among the mite species, 27 species under 9 under 10 families on edible mushrooms from families and 3 orders were damage causing in West Bengal. the case of edible mushrooms while Relative abundance: X= Highly abundant (> corresponding figures for wild mushrooms 10 specimens/ gram of sample) were 13 species under 5 families and 2 orders. Y= Occasional occurrence (>5 specimens but Acarus siro and Tyrophagus putrescentiae <10 specimens/ gram of sample) caused damage on edible mushrooms Z= Rare occurrence (<5 specimens/ gram of (Pleurotus spp.) and Suidasia nesbitti and sample) Tyrophagus putrescentiae caused damage on Edible mushrooms: 1= Calocybe indica, 2= wild mushrooms (Pseudohydnum gelatinosum Pleurotus spp. (ostreatus, djamor, sajor-caju), and Chlorophyllum hortense). As a result of 3= Volvariella volvacea, 4= Agaricus infestation, the mushrooms had shown bisporus. damage symptoms like blackening of the Wild mushrooms: 5= Chlorophyllum straw bed, browning of spore-caps and making hortense, 6= Copalandia cyanescens, the stalks hollow, etc. 7=Crepidotus applanatum, 8= Laccaria As far as predatory mites are concerned, the laccata, 9= Ganoderma lucidum, 10= Inocybe most dominant mites belonged to order umbonata, 11= Russula kanadii, l2= Mesostigmata (families Veigaidae, Pseudohydnum gelatinosum, 13= Earliella Parasitidae, Ascidae) on edible mushrooms scabrosa, 14= Auricularia auricular, 15= and Ascidae on wild mushrooms. Among Coriolopsis occidentalis, 16= Russula Prostigmatid mites, the dominant ones albonigra, 17= Undetermined, 18= belonged to Cheyletidae (in case of wild Ternitomyces sp., 19= Lentinus squarrosulus, mushrooms) and Pyemotidae on edible 20= Chlorophyllum molybidites, 21= Stereum mushrooms. The predatory mites mostly sp., 22=Lenzites sp., 23= Marasmius devoured mites belonging to Acaridae and haematocephalus, 24= Scleroderma sp., 25= Suidasiidae. The other 16 mite species Strobilomyces strobilaceus, 26= Phlebopus reported here were under 16 genera, 14 sp., 27= Russula congoana, 28=Mycena sp., 12 Vol. 22 (1), March, 2020 BIONOTES 29= Ganoderma sp., 30= Volvariella assessment of loss caused by mite complex on nigrodisca, 31=Dictyophora sp. some cultivated species of mushroom. *= New report from India Progress in Acarology, 2: 215-221. **= New report on mushroom from India Das, P., A.K. Somchoudhury, A.B. Mukherjee Acknowledgements & P.K. Sarkar. 1993. Some aspects of feeding The authors are thankful to Swami behavior of mushroom mites. In. Acarological Sarvalokananda Maharaj and Swami Researches in India (Eds. A.K. Vasavananda Maharaj, the Secretary and Somchoudhury, A.B.Mukherjee, & Sarkar, Assistant Secretary, respectively of R. K. P.K.): 271-282. Mission, Narendrapur for providing laboratory facilities. Thanks are also due to Gupta, S.K. 2012. Handbook Injuries and Swami Vishwamayananda Maharaj, beneficial mites infesting agri-horticultural Secretary, R. K. Mission, Sargachi and Shri crops in India and their management, Adar Mukherjee, in-charge in mushroom unit Nature Books India, New Delhi. 1-362. of R. K. Mission, Narendrapur for providing Hill, A. & K.L. Deahl. 1978. Description and samples of mushrooms on which this paper is life cycle of a new species of Histiosoma based. (Acari: Histiostomatidae) associated with References commercial mushroom production. Proc. Aiswarya, M., S.K. Gupta & S.P. James. 2018. Entom. Soc. Wash. 80: 317-329. A preliminary report on Acari inhabiting in Mondal, A. & S.K. Gupta. 2019. Insects and different mushrooms of Western Ghats, mites occurring on mushroom in South 24 Calicut, Kerala. Uttar Pradesh J. Zool. 38(1): Parganas district of West Bengal. Int. J. Sci. 15-19. Res. 8(2): 67-69. Das, P. 1986. Bionomics and control of Mukherjee, A.B. & A.K. Somchoudhury. mushroom mites. Ph.D. thesis. B.C. Agr. 1974. Mite pests of mushrooms. FAO Pl. Prot. Univ., Kalyani: 1-197. Bull. 22(2): 51. Das, P., A.K. Somchoudhury & A.B. Parveen, R. & S.K. Gupta. 2019. Some new Mukherjee. 1987. Nature and habitat of records of mites occurring in mushroom in mushroom mites. Env.Ecol. 5(4): 677-680. South Bengal. Int. J. Zool. Studies 4 (5): 8-12. Das, P., A.K. Somchoudhury, A.B. Mukherjee Parveen, R. & S.K. Gupta. 2020. Diversity of & P.K. Sarkar. 1987a. Some aspects of mites (Acari) on wild mushrooms from West feeding behavior of mushroom mites. Abst. Bengal. Int. J.Agri. & Plant Sci. 2(1): 1-07. First Nat. sem. Acar, BCK., Kalyani, 29-31 Oct., 1987: 24. Somchoudhury, A.K. & A.B. Mukherjee. 1988. Source of mite infesting in mushroom Das, P., A.K. Somchoudhury & A.B. cultivation. Env. Ecol. 6(3): 669-671. Mukherjee. 1988. Source of mite infestation in mushroom. Env.Ecol. 6(3): 669-671. Trivedi, T.P. 1988. Occurrence of mites in the beds of cultivated mushroom. Univ. Agri. Sci. Das, P., A.K. Somchoudhury & A.B. Bangalore. Curr. Res. 17(9): 125. Mukherjee. 1989. Seasonal incidence and

13 Vol. 22 (1), March, 2020 BIONOTES Table 1: List of mites both from edible/ wild mushrooms from India with respective hosts/ habitats, relative abundance and nature of association with mushroom species.

31) Mites Species -

References

Family Species

Relative abundance

Sl. No. Order Suborder Edible mushroom (1 4) Wild mushroom (5 Damage causing Predator Others 1 Acaridae Acarus siro Linn. X + + + - - Parveen & Gupta, 2019 Parveen & Gupta, 2020 Aiswarya et al.,2018 2 Acarus gracilis Hughes Z + - + - - Parveen & Gupta, 2019 3 Acarus farris Oudemans Z + + + - - Parveen & Gupta, 2020 4 Tyrophagus dimidiatus Y + - + - - Gupta, 2012 Hermann Das et al., 1987 Somchoudhury &

Mukherjee, 1988 5 Tyrophagus berlesei Z + - + - - Gupta, 2012 Michael 6 Tyrophagus X + + + - - Gupta, 2012

Astigmata - putrescentiae Schrank Parveen & Gupta, 2019 Parveen & Gupta, 2020 Aiswarya et al., 2018

Oribatida: Cohort Oribatida:

Mukherjee &

- Somchoudhury, 1972

7 Tyrophagus longior Z + + + - - Parveen & Gupta, Gervais 2019 Parveen & Gupta, 2020 Aiswarya et al., 2018

Sarcoptiformes: SuborderSarcoptiformes:

14 Vol. 22 (1), March, 2020 BIONOTES 8 Tyrophagus perniciosus Z - + + - - Mukherjee & Zachvatkin Somchoudhury, 1974 9 Rhizoglyphus echinopus X + - + - - Gupta, 2012 Fumouze and Robin Parveen & Gupta, 2019 Somchoudhury & Mukherjee, 1988 Das et al. 1987, 1988, 1989 10 Rhizoglyphus robini Y + - + - - Parveen & Gupta, Claparede 2019 11 Caloglyphus oudemansi Z + - + - - Parveen & Gupta, Zachvatkin 2019 12 Caloglyphus Z + - + - - Gupta, 2012 mycophagus Megnin 13 Caloglyphus berlesei Z + - + - - Parveen & Gupta, Michael 2019 14 Caloglyphus hughesi Z - + + - - Aiswarya et al., Samsinak 2018 15 Histiostomidae Histiostoma heinemanni Z + - + - - Gupta, 2012 Hill & Diahl Das et al., 1987, 1989 Somchoudhury & Mukherjee, 1988 16 Histiostoma ferroniarum X + + + - - Aiswarya et al., Dufour 2018 Parveen & Gupta, 2019 Parveen & Gupta, 2020 17 Histiostoma gracilipis Z + - + - - Gupta, 2012 Banks Hill & Deahl, 1978 18 Histiostoma X + + + - - Parveen & Gupta, sapromyzarum Dufour 2019 Parveen & Gupta, 2020 19 Glycyphagidae Glycyphagus domesticus X + - + - - Aiswarya et al., De Geer 2018 Parveen & Gupta, 2019 20 Glycyphagus bicaudatus Z + - + - - Parveen & Gupta, Hughes 2019 21 Glycyphagus ornatus Z - + + - - Aiswarya et al., Kramer 2018

15 Vol. 22 (1), March, 2020 BIONOTES 22 Austroglycyphagus Z - + + - - Parveen & Gupta, geniculatus Vitzthum 2020 23 Lepidoglyphus destructor X - + + - - Aiswarya et al., Schrank 2018 Parveen & Gupta, 2019 Parveen & Gupta, 2020 24 Suidasiidae Suidasia nesbitti Sasa X + + + - - Parveen & Gupta, 2019 Parveen & Gupta, 2020 25 Tarsonemidae Tarsonemus granarius Z - + + - - Parveen & Gupta, Lindquist 2020

26 Tarsonemus Z + - + - - Gupta, 2012 myceleiphagus Austin & Jary 27 Tarsonemus confusus Z + - + - - Gupta, 2012 Ewing 28 Tarsonemus tarsalis Z + - + - - Gupta, 2012 Canestrini

29 Pygmephoridae Pygmephorus selinicki Z + - + - - New report Krczal

30 Pygmephorus Z + - + - - New report fletchmanni Wicht 31 Dolichocybidae Dolichocybe keiferi Y + - + - - New report Krantz 32 Scutacaridae **Scutacarus Z + - + - - New report baculitarsus Norton & Ide 33 Tydeidae Tydeus collyerae Baker X + - - + - Parveen & Gupta, 2019 34 **Tydeus gosabaensis Z - + - + - New report Gupta 35 **Lorrya stricta Gupta Z + - - + - New report

Prostigmata

- 36 Raphignathidae Raphignathus sp. X + - - - + Parveen & Gupta, 2019 37 Pyemotidae Pyemotes herfsi X + - + - - Parveen & Gupta, Oudemans 2019 38 Caligonellidae Neognathus sp. Z + - - - + Parveen & Gupta, 2019

Trombidiformes: Suborder 16 Vol. 22 (1), March, 2020 BIONOTES 39 Cunaxidae Neocunaxoides sp.n. Z - - + - Parveen & Gupta, + 2019 40 Cunaxoides biscutum Z - + - + - Parveen & Gupta, Nesbitt 2020 41 Tenuiepalpidae Brevipalpus euphorbiae Z - + - - + Parveen & Gupta, Mohanasundarum 2020 42 Cheyletidae Cheyletus eruditus Y + + - + - Parveen & Gupta, Schrank 2020 43 Cheyletus audex Y - + - + - Parveen & Gupta, Oudemans 2020 44 Eucheyletia sinensis Z - + - + - Parveen & Gupta, Volgin 2020 45 Chelacaropsis moorei Z - + - + - Parveen & Gupta, Baker 2020 46 Iolinidae **Pronematus fleschneri Z - + - - + New report Baker 47 Stigmaeidae Cheylostigmaeus sp.n. Z - + - + - Parveen & Gupta, 2020 48 Erythraeidae *Charletonia rocciai Z - + - + - New report Treat & Flechtmann 49 Blattisociidae Lasioseius quadrisetosus X + - - + - Parveen & Gupta, Chant 2019 50 Lasioseius floridensis X - + - + - Parveen & Gupta, Berlese 2020 51 **Lasioseius parberlesei Z - + - + - New report Bhattacharya 52 Lasioseius mcgregori Z - + - + - Parveen & Gupta, Chant 2020 53 Lasioseius formosus Z - + - + - Aiswarya et al., Westerboer 2018 54 Lasioseius penicilliger Y + + - + - Aiswarya et al., Berlese 2018 55 Ascidae Zercoseius spathuliger Z - + - + - Parveen & Gupta, Leonardi 2020 56 **Asca biswasi Z + - - + - New report Bhattacharyya 57 Asca garmani Hurlbutt Z - + - + - Parveen & Gupta, 2020 58 Antennoseius indicus Y + - - + - Parveen & Gupta, Bhattacharyya 2019 59 Gamasellodes bicolor Y + + - + - Parveen & Gupta, Berlese 2020 60 Cheiroseius laelaptoides Z - + - + - Parveen & Gupta, Berlese 2020 61 Platyseius subglaber Z - + - + - Parveen & Gupta, Berlese 2020

Mesostigmata 17 Vol. 22 (1), March, 2020 BIONOTES 62 Melicharidae Proctolaelaps pygmaeus Z - + - + - Aiswarya et al., Muller 2018 63 Parasitidae Parasitus assamensis X + - - + - Parveen & Gupta, Bhattacharyya 2019 64 Parasitus consanguineus X + - - + - Parveen & Gupta, Oudemans & Voigts 2019 Trivedi, 1988 65 Parasitus shillongensis X - + - + - Parveen & Gupta, Bhattacharyya 2020 66 **Pergamasus primitivus Z + - - + - New report Oudemans 67 Pergamasus crassipes Z + - - + - Parveen & Gupta, Berlesei 2019 68 Pseudoparasitus sp. Z - + - + - Aiswarya et al., 2018 69 Phytoseiidae Typhlodromous sp. n. 1 Y + - - + - Parveen & Gupta, 2019 70 Typhlodromous sp. n. 2 Y + - - + - Parveen & Gupta, 2019

71 Typhlodromous sp. n. 3 Y + - - + - Parveen & Gupta, 2019 72 Typhlodromous sp. n. 4 Y - + - + - Parveen & Gupta, 2020 73 *Typhlodromous Z + - - + - New report (Anthoseius) egypticus EL-Badry 74 Neoseiulus fallacies Z - + - + - Parveen & Gupta, (Garman) 2020 75 Amblyseius herbicolus Z + - - + - Mondal & Gupta, (Chant) 2019 76 Laelapidae Cosmolaelaps indicus X + + - + - Parveen & Gupta, Bhattacharyya 2019 Parveen & Gupta, 2020 77 Cyrtolaelaps sp. Z - + - + - Parveen & Gupta, 2020 78 Hypoaspis miles Berlese X + - - + - Somchoudhury & Mukherjee, 1987 Das et al., 1989 Gupta, 2012 79 Hypoaspis berlesei Z + - - + - Mondal & Gupta, (Oudemans) 2019 80 Hypoaspis aculifer Y + + - + - Aiswarya et al., Canestrini 2018

Mesostigmata

18 Vol. 22 (1), March, 2020 BIONOTES 81 *Hypoaspis lubrica Z + - - + - New report Voigts and Oudemans

82 **Androlaelaps casalis Z + - - + - New report (Berlese) 83 Ololaelaps sp. Z + - - + - New report 84 Eviphidae Crassicheles sp. Z + - - + - Parveen & Gupta, 2019 85 Polyaspididae Polyaspis sp. Z + - - + - Parveen & Gupta, 2019 86 Uropolyaspis sp. Z - + - + - Parveen & Gupta, 2020 87 Resinacaridae Resinacarus resinatus Z + - - - + Parveen & Gupta, Vitzthum 2019 88 Rhodacaridae Rhodacarus sp. Z + - - + - Parveen & Gupta, 2019 89 Uropodidae Fuscuropoda marginata X + + - - + Aiswarya et al., C.L.Koch 2018 Parveen & Gupta, 2019 Parveen & Gupta, 2020 90 Trematura sp. Z + - - - + Parveen & Gupta, 2019 91 Leiodenychus krameri X - + - - + Aiswarya et al., (Canestrini) 2018 Parveen & Gupta, 2020 92 Veigaidae Veiga uncata Farrier X + - + - - Parveen & Gupta, 2019 93 Zerconidae Zercon prasadi Blaszak Z - + - + - Parveen & Gupta, 2020 94 Sejiidae Sejus togatus Koch Y + + - + - Parveen & Gupta, 2020 95 Macrocheliidae Macrocheles Y + + - + - Parveen & Gupta, muscaedomesticae 2020 (Scopoli) 96 * Macrocheles glaber Z - + - + - New report Muller 97 Neoparasitidae Gamasiphis Z - + - + - Parveen & Gupta, (Neogamasiphis) 2020 bengalensis Bhattacharyya 98 Ameroseiidae Kleemania plumosus Z - + - - + Aiswarya et al., (Oudemans) 2018

Mesostigmata

19 Vol. 22 (1), March, 2020 BIONOTES 99 Zerconopsidae Zerconopsis sp. Z - + - - + Aiswarya et al., 2018

10 Pachylaelapidae Pachylaelaps dorsalis Z + - - + - Mondal & Gupta, 0 Bhattacharya 2019 10 Galumnidae Galumna flabellifera Y - + - - + Parveen & Gupta, 1 Von Heyden 2020 10 Ceratozetidae Ceratozetes sp. Y - + - - + Parveen & Gupta,

Oribatida Oribatida 2 2020

- 10 Trhypochthoniid Archegozetes sp. Z - + - - + Parveen & Gupta, 3 ae 2020 10 Oppiidae Oppia sp. Z - + - - + Parveen & Gupta, 4 2020 10 Oribatulidae Oribatula sp. Z - + - - + Parveen & Gupta, 5 2020

10 Austrachipteriid Lemellobates sp. Z - + - - + Parveen & Gupta, 6 ae 2020

Sarcoptiformes: SuborderSarcoptiformes: (excluding Astigmata)