A.N.P.S.A. Fern Study Group Newsletter Number 125
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A Taxonomic Revision of Hymenophyllaceae
BLUMEA 51: 221–280 Published on 27 July 2006 http://dx.doi.org/10.3767/000651906X622210 A TAXONOMIC REVISION OF HYMENOPHYLLACEAE ATSUSHI EBIHARA1, 2, JEAN-YVES DUBUISSON3, KUNIO IWATSUKI4, SABINE HENNEQUIN3 & MOTOMI ITO1 SUMMARY A new classification of Hymenophyllaceae, consisting of nine genera (Hymenophyllum, Didymoglos- sum, Crepidomanes, Polyphlebium, Vandenboschia, Abrodictyum, Trichomanes, Cephalomanes and Callistopteris) is proposed. Every genus, subgenus and section chiefly corresponds to the mono- phyletic group elucidated in molecular phylogenetic analyses based on chloroplast sequences. Brief descriptions and keys to the higher taxa are given, and their representative members are enumerated, including some new combinations. Key words: filmy ferns, Hymenophyllaceae, Hymenophyllum, Trichomanes. INTRODUCTION The Hymenophyllaceae, or ‘filmy ferns’, is the largest basal family of leptosporangiate ferns and comprises around 600 species (Iwatsuki, 1990). Members are easily distin- guished by their usually single-cell-thick laminae, and the monophyly of the family has not been questioned. The intrafamilial classification of the family, on the other hand, is highly controversial – several fundamentally different classifications are used by indi- vidual researchers and/or areas. Traditionally, only two genera – Hymenophyllum with bivalved involucres and Trichomanes with tubular involucres – have been recognized in this family. This scheme was expanded by Morton (1968) who hierarchically placed many subgenera, sections and subsections under -
Flora and Fauna of Phong Nha-Ke Bang and Hin Namno, a Compilation Page 2 of 151
Flora and fauna of Phong Nha-Ke Bang and Hin Namno A compilation ii Marianne Meijboom and Ho Thi Ngoc Lanh November 2002 WWF LINC Project: Linking Hin Namno and Phong Nha-Ke Bang through parallel conservation Flora and fauna of Phong Nha-Ke Bang and Hin Namno, a compilation Page 2 of 151 Acknowledgements This report was prepared by the WWF ‘Linking Hin Namno and Phong Nha through parallel conservation’ (LINC) project with financial support from WWF UK and the Department for International Development UK (DfID). The report is a compilation of the available data on the flora and fauna of Phong Nha-Ke Bang and Hin Namno areas, both inside and outside the protected area boundaries. We would like to thank the Management Board of Phong Nha-Ke Bang National Park, especially Mr. Nguyen Tan Hiep, Mr. Luu Minh Thanh, Mr. Cao Xuan Chinh and Mr. Dinh Huy Tri, for sharing information about research carried out in the Phong Nha-Ke Bang area. This compilation also includes data from surveys carried out on the Lao side of the border, in the Hin Namno area. We would also like to thank Barney Long and Pham Nhat for their inputs on the mammal list, Ben Hayes for his comments on bats, Roland Eve for his comments on the bird list, and Brian Stuart and Doug Hendrie for their thorough review of the reptile list. We would like to thank Thomas Ziegler for sharing the latest scientific insights on Vietnamese reptiles. And we are grateful to Andrei Kouznetsov for reviewing the recorded plant species. -
Two Species of Armored Scale Insects (Hemiptera: Diaspididae) Associated with Sori of Ferns Marcelo Guerra Santos¹ & Vera Regina Dos Santos Wolff²
doi:10.12741/ebrasilis.v8i3.492 e-ISSN 1983-0572 Publicação do Projeto Entomologistas do Brasil www.ebras.bio.br Distribuído através da Creative Commons Licence v4.0 (BY-NC-ND) Copyright © EntomoBrasilis Copyright © do(s) Autor(es) Two Species of Armored Scale Insects (Hemiptera: Diaspididae) Associated with Sori of Ferns Marcelo Guerra Santos¹ & Vera Regina dos Santos Wolff² 1. Universidade do Estado do Rio de Janeiro, e-mail: [email protected] (Autor para correspondência). 2. Fundação Estadual de Pesquisa Agropecuária – FEPAGRO, Rio Grande do Sul, e-mail: [email protected]. _____________________________________ EntomoBrasilis 8 (3): 232-234 (2015) Abstract. This note reports the presence of two scale insects species Hemiberlesia palmae (Cockerell) and Pinnaspis strachani (Cooley) (Coccoidea, Diaspididae), associated respectively with Asplenium serratum L. (Aspleniaceae) and Niphidium crassifolium (L.) Lellinger (Polypodiaceae). It is the first record of a fern species as host plant of H. palmae. In both fern species, the diaspidids were found nearby the sori. Keywords: Aspleniaceae; Fern-insect interactions; Polypodiaceae; Pteridophytes; Scale Insect. Duas Espécies de Cochonilhas (Hemiptera: Diaspididae) Associadas com Soros de Samambaias Resumo. A presente comunicação relata a presença de duas espécies de cochonilhas Hemiberlesia palmae (Cockerell) e Pinnaspis strachani (Cooley) (Coccoidea, Diaspididae), associadas respectivamente com Asplenium serratum L. (Aspleniaceae) e Niphidium crassifolium (L.) Lellinger (Polypodiaceae). É o primeiro registro de uma samambaia como planta hospedeira de H. palmae. Nas duas espécies de samambaias, os diaspidídeos encontravam-se concentrados principalmente ao redor dos soros. Palavras-chave: Aspleniaceae; Cochonilhas; Interações samambaia-inseto; Polypodiaceae; Pteridófitas. _____________________________________ nteractions between ferns and insects are more poorly (2003). -
Epilist 1.0: a Global Checklist of Vascular Epiphytes
Zurich Open Repository and Archive University of Zurich Main Library Strickhofstrasse 39 CH-8057 Zurich www.zora.uzh.ch Year: 2021 EpiList 1.0: a global checklist of vascular epiphytes Zotz, Gerhard ; Weigelt, Patrick ; Kessler, Michael ; Kreft, Holger ; Taylor, Amanda Abstract: Epiphytes make up roughly 10% of all vascular plant species globally and play important functional roles, especially in tropical forests. However, to date, there is no comprehensive list of vas- cular epiphyte species. Here, we present EpiList 1.0, the first global list of vascular epiphytes based on standardized definitions and taxonomy. We include obligate epiphytes, facultative epiphytes, and hemiepiphytes, as the latter share the vulnerable epiphytic stage as juveniles. Based on 978 references, the checklist includes >31,000 species of 79 plant families. Species names were standardized against World Flora Online for seed plants and against the World Ferns database for lycophytes and ferns. In cases of species missing from these databases, we used other databases (mostly World Checklist of Selected Plant Families). For all species, author names and IDs for World Flora Online entries are provided to facilitate the alignment with other plant databases, and to avoid ambiguities. EpiList 1.0 will be a rich source for synthetic studies in ecology, biogeography, and evolutionary biology as it offers, for the first time, a species‐level overview over all currently known vascular epiphytes. At the same time, the list represents work in progress: species descriptions of epiphytic taxa are ongoing and published life form information in floristic inventories and trait and distribution databases is often incomplete and sometimes evenwrong. -
The Anticonvulsant Activity of Asplenium Nidus L. (Polypodiaceae) Methanolic Crude Leaf Extract in Chemically Induced Tonic- Clonic Convulsions on Swiss Mice
The STETH, Vol. 12, 2018 The anticonvulsant activity of Asplenium nidus L. (Polypodiaceae) methanolic crude leaf extract in chemically induced tonic- clonic convulsions on Swiss mice Holy May B. Faral*, Ronalyn B. Macaraig, Princess Marian B. Mojares, Reina Jean D. Caramat, Donnabel D. Abando, Laurina M. Balangi, Sheryl C. Aguila, Omar A. Villalobos Pharmacy Department, College of Allied Medical Professions, Lyceum of the Philippines University, Batangas City *[email protected] ABSTRACT: Epilepsy is a chronic non-communicable disorder of the brain that affects people of all ages. Approximately, 50 million people worldwide have epilepsy, making it one of the most common neurological diseases globally. Asplenium nidus L. is a member of family Polypodiaceae which is commonly known as “bird’s nest fern.” It is used in many traditional medicines such as antipyretic, estrogenic, spasmolytic and medicinally as depurative and sedative. The present investigation was designed to evaluate the anticonvulsant activity of the Asplenium nidus L. in pentylnetetrazole and isoniazid-induced convulsions on 30 male Swiss mice equally divided into five groups. After acute toxicity test, oral treatment with Asplenium nidus methanolic extract at varying doses of 500, 750 and 1000 mg/kg BW was given to test animals. The efficacy of the plant extract was compared with diazepam as the standard drug (5 mg/kg BW) and PNSS (10 ml/kg BW) as the control. The significant of differences between groups was determined using Kruskal Wallis Test followed by the Mann Whitney p < 0.05. The data was presented as mean ± SEM in tables. Data were analysed using SPSS v.21 at 95% level of confidence. -
Reprint Requests, Current Address: Dept
American Journal of Botany 88(6): 1118±1130. 2001. RBCL DATA REVEAL TWO MONOPHYLETIC GROUPS OF FILMY FERNS (FILICOPSIDA:HYMENOPHYLLACEAE)1 KATHLEEN M. PRYER,2,5 ALAN R. SMITH,3 JEFFREY S. HUNT,2 AND JEAN-YVES DUBUISSON4 2Department of Botany, The Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, Illinois 60605-2496 USA; 3University Herbarium, 1001 Valley Life Sciences Building #2465, University of California, Berkeley, California 94720-2465 USA; and 4Laboratoire de PaleÂobotanique et PaleÂoeÂcologie, FR3-CNRS ``Institut d'E cologie,'' Universite Pierre et Marie Curie, 12 rue Cuvier, F-75005 Paris, France The ``®lmy fern'' family, Hymenophyllaceae, is traditionally partitioned into two principal genera, Trichomanes s.l. (sensu lato) and Hymenophyllum s.l., based upon sorus shape characters. This basic split in the family has been widely debated this past century and hence was evaluated here by using rbcL nucleotide sequence data in a phylogenetic study of 26 ®lmy ferns and nine outgroup taxa. Our results con®rm the monophyly of the family and provide robust support for two monophyletic groups that correspond to the two classical genera. In addition, we show that some taxa of uncertain af®nity, such as the monotypic genera Cardiomanes and Serpyllopsis, and at least one species of Microtrichomanes, are convincingly included within Hymenophyllum s.l. The tubular- or conical-based sorus that typi®es Trichomanes s.l. and Cardiomanes, the most basal member of Hymenophyllum s.l., is a plesiomorphic character state for the family. Tubular-based sori occurring in other members of Hymenophyllum s.l. are most likely derived independently and more than one time. -
Fern Classification
16 Fern classification ALAN R. SMITH, KATHLEEN M. PRYER, ERIC SCHUETTPELZ, PETRA KORALL, HARALD SCHNEIDER, AND PAUL G. WOLF 16.1 Introduction and historical summary / Over the past 70 years, many fern classifications, nearly all based on morphology, most explicitly or implicitly phylogenetic, have been proposed. The most complete and commonly used classifications, some intended primar• ily as herbarium (filing) schemes, are summarized in Table 16.1, and include: Christensen (1938), Copeland (1947), Holttum (1947, 1949), Nayar (1970), Bierhorst (1971), Crabbe et al. (1975), Pichi Sermolli (1977), Ching (1978), Tryon and Tryon (1982), Kramer (in Kubitzki, 1990), Hennipman (1996), and Stevenson and Loconte (1996). Other classifications or trees implying relationships, some with a regional focus, include Bower (1926), Ching (1940), Dickason (1946), Wagner (1969), Tagawa and Iwatsuki (1972), Holttum (1973), and Mickel (1974). Tryon (1952) and Pichi Sermolli (1973) reviewed and reproduced many of these and still earlier classifica• tions, and Pichi Sermolli (1970, 1981, 1982, 1986) also summarized information on family names of ferns. Smith (1996) provided a summary and discussion of recent classifications. With the advent of cladistic methods and molecular sequencing techniques, there has been an increased interest in classifications reflecting evolutionary relationships. Phylogenetic studies robustly support a basal dichotomy within vascular plants, separating the lycophytes (less than 1 % of extant vascular plants) from the euphyllophytes (Figure 16.l; Raubeson and Jansen, 1992, Kenrick and Crane, 1997; Pryer et al., 2001a, 2004a, 2004b; Qiu et al., 2006). Living euphyl• lophytes, in turn, comprise two major clades: spermatophytes (seed plants), which are in excess of 260 000 species (Thorne, 2002; Scotland and Wortley, Biology and Evolution of Ferns and Lycopliytes, ed. -
Disentangling the Diversity and Taxonomy of Hymenophyllaceae (Hymenophyllales, Polypodiidae) in the Comoros Ahamada H
Disentangling the diversity and taxonomy of Hymenophyllaceae (Hymenophyllales, Polypodiidae) in the Comoros Ahamada H. Saïd, Sabine Hennequin, Germinal Rouhan, Jean-Yves Dubuisson To cite this version: Ahamada H. Saïd, Sabine Hennequin, Germinal Rouhan, Jean-Yves Dubuisson. Disentangling the diversity and taxonomy of Hymenophyllaceae (Hymenophyllales, Polypodiidae) in the Comoros. Eu- ropean Journal of Taxonomy, Consortium of European Natural History Museums, 2017, 313, pp.1-53. 10.5852/ejt.2017.313. hal-01528615 HAL Id: hal-01528615 https://hal.sorbonne-universite.fr/hal-01528615 Submitted on 29 May 2017 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. Distributed under a Creative Commons Attribution| 4.0 International License European Journal of Taxonomy 313: 1–53 ISSN 2118-9773 https://doi.org/10.5852/ejt.2017.313 www.europeanjournaloftaxonomy.eu 2017 · Saïd A.H. et al. This work is licensed under a Creative Commons Attribution 3.0 License. Monograph Disentangling the diversity and taxonomy of Hymenophyllaceae (Hymenophyllales, Polypodiidae) in the Comoros Ahamada H. SAÏD 1,*, Sabine HENNEQUIN 2, Germinal ROUHAN 3 & Jean-Yves DUBUISSON 4 1,3 Institut Systématique Evolution Biodiversité (ISYEB), Sorbonne Universités, Muséum national d’Histoire naturelle, CNRS, UPMC, EPHE, Herbier National, 57 rue Cuvier, CP39, 75005 Paris, France. -
A Revised Family-Level Classification for Eupolypod II Ferns (Polypodiidae: Polypodiales)
TAXON 61 (3) • June 2012: 515–533 Rothfels & al. • Eupolypod II classification A revised family-level classification for eupolypod II ferns (Polypodiidae: Polypodiales) Carl J. Rothfels,1 Michael A. Sundue,2 Li-Yaung Kuo,3 Anders Larsson,4 Masahiro Kato,5 Eric Schuettpelz6 & Kathleen M. Pryer1 1 Department of Biology, Duke University, Box 90338, Durham, North Carolina 27708, U.S.A. 2 The Pringle Herbarium, Department of Plant Biology, University of Vermont, 27 Colchester Ave., Burlington, Vermont 05405, U.S.A. 3 Institute of Ecology and Evolutionary Biology, National Taiwan University, No. 1, Sec. 4, Roosevelt Road, Taipei, 10617, Taiwan 4 Systematic Biology, Evolutionary Biology Centre, Uppsala University, Norbyv. 18D, 752 36, Uppsala, Sweden 5 Department of Botany, National Museum of Nature and Science, Tsukuba 305-0005, Japan 6 Department of Biology and Marine Biology, University of North Carolina Wilmington, 601 South College Road, Wilmington, North Carolina 28403, U.S.A. Carl J. Rothfels and Michael A. Sundue contributed equally to this work. Author for correspondence: Carl J. Rothfels, [email protected] Abstract We present a family-level classification for the eupolypod II clade of leptosporangiate ferns, one of the two major lineages within the Eupolypods, and one of the few parts of the fern tree of life where family-level relationships were not well understood at the time of publication of the 2006 fern classification by Smith & al. Comprising over 2500 species, the composition and particularly the relationships among the major clades of this group have historically been contentious and defied phylogenetic resolution until very recently. Our classification reflects the most current available data, largely derived from published molecular phylogenetic studies. -
Studies in the Systematics of Filmy Ferns VIII C 同phalomanes Presl Subgen
植物研究雑誌 J. J. Jpn. Bo t. 66: 66: 134-146 (1 991) Studies in the Systematics of Filmy Ferns VIII C 同phalomanes Presl subgen. Cephalomanes Kunio IWATSUKI Botanical Botanical Gardens ,Faculty of Science ,University of Tokyo , 3-7-1 ,Hakusan ,Bunkyo ・ku ,Tokyo , 112 JAPAN コケシノブ類の分類に関する研究班 ソテツホラゴケ亜属 岩槻邦男 東京大学理学部附属植物園 112 東京都文京区白山3-7-1 (Received (Received on January 7, 1991) A filmy fern subgenus , Cephalomanes subgen. Cephalomanes , was taxonomically revised. Taxo- nornic nornic characters were surveyed and re-evaluated from systematic viewpoints. Six species including infraspecific infraspecific taxa were recognized and keyed ou t. Enurneration of all taxa was given with notes on distribution distribution and taxonomy. C句phaloman 回 crassum and C. madagascariense are restricted to particular localities localities and are rare with a few collections. C 句phalomanes densinervium and C. singaporianum are also also restricted to narrow areas and are not variable , but C. atrovirens and C. javanicum have wide areas areas of distribution and are variable with many infraspecific taxa. Rheophytic nature , or adaptation to to the habitat along stream , is discussed in relation to habitat and phenetic features. (Continued (Continued from Acta Phytotax. Geobot. 35: 165-179 , 1984.) Cephalomanes was described by Presl in 1843 following to Copeland in its position. Most of the on the basis of C. α trovirens and some ten species species have been ill-defined ,and a revision is have currently been referred to subgen. necessary for this complex group. Cephalomanes (Iwatsuki 1984) from the Old In the present part of this series , subgen. World tropics: East Madagascar , India to con- Cephalomanes will be treated in its strict sense , tinental tinental Southeast Asia ,Hainan ,Taiwan , southern though the generic classification was revised and Ryukyus ,throughout 恥1alesia to Polynesia , south broadly circumscribed leaving Cephalomanes s. -
Hymenophyllaceae) and a New Combination in Trichomanes L
Muelleria 39: 75–78 Published online in advance of the print edition, 21 December 2020 Comparison of modern classifications for filmy ferns (Hymenophyllaceae) and a new combination in Trichomanes L. for the filmy fern Macroglena brassii Croxall, from Queensland, Australia Daniel J. Ohlsen Royal Botanic Gardens Victoria, Birdwood Avenue, Melbourne, Victoria 3004, Australia; email: [email protected] Introduction Abstract Contemporary Hymenophyllaceae The filmy ferns (Hymenophyllaceae) are a distinctive group of treatments typically follow one of leptosporangiate ferns distinguished by a thin membranous lamina that two classifications that recognise is usually one cell thick (or occasionally up to four cells thick in some parts monophyletic genera. One comprises of the lamina) and marginal sori that are protected by an indusium in the nine genera, while the other recognises form of a cup-shaped or bilabiate involucre (Ebihara et al. 2007). Forty nine two genera, Hymenophyllum Sm. and Trichomanes L. Combinations exist for species of this family occur in Australia, of which 15 are probably endemic all Australian species that allow the (Green 1994; Bostock & Spokes 1998; Ebihara & Iwatsuki 2007). Two major former classification to be adopted lineages exist within the Hymenophyllaceae that largely correspond to the in Australia. However, genera of the two original genera recognised within the family: Hymenophyllum Sm. and former classification tend to be poorly Trichomanes L. (Pryer et al. 2001; Hennequin et al. 2003; Ebihara et al. 2004). defined morphologically compared to Numerous other classifications have been proposed that recognise several the latter classification. All Australian species have available combinations in additional genera (e.g. -
Systematics and Biogeography of the Clusioid Clade (Malpighiales) Brad R
Eastern Kentucky University Encompass Biological Sciences Faculty and Staff Research Biological Sciences January 2011 Systematics and Biogeography of the Clusioid Clade (Malpighiales) Brad R. Ruhfel Eastern Kentucky University, [email protected] Follow this and additional works at: http://encompass.eku.edu/bio_fsresearch Part of the Plant Biology Commons Recommended Citation Ruhfel, Brad R., "Systematics and Biogeography of the Clusioid Clade (Malpighiales)" (2011). Biological Sciences Faculty and Staff Research. Paper 3. http://encompass.eku.edu/bio_fsresearch/3 This is brought to you for free and open access by the Biological Sciences at Encompass. It has been accepted for inclusion in Biological Sciences Faculty and Staff Research by an authorized administrator of Encompass. For more information, please contact [email protected]. HARVARD UNIVERSITY Graduate School of Arts and Sciences DISSERTATION ACCEPTANCE CERTIFICATE The undersigned, appointed by the Department of Organismic and Evolutionary Biology have examined a dissertation entitled Systematics and biogeography of the clusioid clade (Malpighiales) presented by Brad R. Ruhfel candidate for the degree of Doctor of Philosophy and hereby certify that it is worthy of acceptance. Signature Typed name: Prof. Charles C. Davis Signature ( ^^^M^ *-^£<& Typed name: Profy^ndrew I^4*ooll Signature / / l^'^ i •*" Typed name: Signature Typed name Signature ^ft/V ^VC^L • Typed name: Prof. Peter Sfe^cnS* Date: 29 April 2011 Systematics and biogeography of the clusioid clade (Malpighiales) A dissertation presented by Brad R. Ruhfel to The Department of Organismic and Evolutionary Biology in partial fulfillment of the requirements for the degree of Doctor of Philosophy in the subject of Biology Harvard University Cambridge, Massachusetts May 2011 UMI Number: 3462126 All rights reserved INFORMATION TO ALL USERS The quality of this reproduction is dependent upon the quality of the copy submitted.