Native Vegetation Condition Assessment and Monitoring Manual for Western Australia

Total Page:16

File Type:pdf, Size:1020Kb

Native Vegetation Condition Assessment and Monitoring Manual for Western Australia Native Vegetation Condition Assessment and Monitoring Manual for Western Australia Prepared by: N. Casson, S. Downes, and A. Harris Prepared for: The Native Vegetation Integrity Project 2009 This project is funded by the Australian Government and the Government of Western Australia CONTENTS ACKNOWLEDGEMENTS PURPOSE AND SCOPE 1) INTRODUCTION 1.1) MAINTENANCE OF VEGETATION - “TYPES OF” CHANGE 1.2) ISOLATING THE SIGNS AND SYMPTOMS OF CHANGE 1.3) A PLETHORA OF FACTORS 2) HOW TO USE THIS MANUAL 2.1) ORIENTATION AND SITE SELECTION 2.2) BASIC ASSESSMENT 2.3) STANDARD ASSESSMENT 2.4) LINKING REMOTE SENSING 2.5) QUANTITATIVE ASSESSMENT FOR VARIOUS PURPOSES 2.6) MONITORING 2.7) LIMITATIONS 2.8) USE OF TERMS 3) ORIENTATION– BACKGROUND, RECONNAISSANCE & OBSERVATION 3.1) LITERATURE-BASED APPROACH - FINDING BACKGROUND INFORMATION 3.2) FIELD APPROACH - FINDING REFERENCE POINTS 3.2.1) TIMING OF FIELDWORK 3.2.2) LOCAL RECONNAISSANCE 3.2.3) OBSERVE THAT COVER VARIES & FLUCTUATES 3.2.4) EVALUATION OF ATTRIBUTES AT REFERENCE AREAS 3.2.5) EVALUATING MORE VEGETATION TYPES 3.2.6) FRAGMENTED OR RESTRICTED VEGETATION TYPES. 3.2.7) USING REFERENCE VALUES TO SUPPORT CONDITION ASSESSMENT. 4) SITE SELECTION 4.1) AIDS TO SITE SELECTION 4.2) LOCATION WITHIN A VEGETATION UNIT 4.3) EDGE EFFECTS 5) CONDITION 5.1) SITE LAYOUT 5.2) RECORDING DATA & SCORING ATTRIBUTES 5.3) SUMMATION OF THE SCORES 5.4) PRESENTATION OF THE SCORES 5.5) ASSESSMENT WITH CONDENSED SCALES OF FEW ATTRIBUTES 5.5.1) CONDITION SCALES – THE NATIONAL “VAST” SCHEME 5.5.2) CONDITION SCALES – THE W.A. “KEIGHERY” SCHEME 5.5.3) CONDITION SCALES - AN ALTERNATE SHORT SCALE 5.5.4) CONDITION SCALES - COLOUR CODE FOR DISPLAY 5.6) ASSESSMENT WITH MORE ATTRIBUTES 5.6.1) CONDITION SCALES - A BASIC VERSION 5.6.2) CONDITION SCALES – A TYPICAL LIST OF SITE ATTRIBUTES 5.6.2.1) NOTES ON SOME UNDERLYING CONCEPTUAL MATTERS 5.6.2.2) NOTES ON RESOURCE DOCUMENTS 5.6.2.3) BACKGROUND & GUIDANCE FOR SITE ATTRIBUTES 5.6.3) BALANCING LANDSCAPE CONTEXT & VEGETATION CONDITION ATTRIBUTES 5.6.3.1) CONDITION SCALES - ADDING LANDSCAPE ATTRIBUTES 5.6.4) CONDITION SCALES - ADDING & ADJUSTING ATTRIBUTES 6) PLANT DIVERSITY 6.1) COORDINATION OF PLANT DIVERSITY & CONDITION 7) CONTEMPORARY REMOTE-SENSING ARRAY 7.1) RATIONALE – REMOTE SENSING PIXEL SIZE & ALIGNMENT 7.2) SITE LAYOUT 7.3) BASIC PROTOCOL FOR MEASURES 8) ADDING OTHER FORMS OF ASSESSMENT 8.1) THREE WAYS OF ASSESSING EACH QUADRAT 8.1.1) WHOLE QUADRAT MEASURES 8.1.2) 1 OR 2 DIAGONAL TRANSECTS 8.1.3) NESTED SUBPLOTS 8.1.4) TRENDS AND STATISTICAL ANALYSES OF TIME SERIES 8.2) LINKING AN ARRAY TO REMOTE ASSESSMENT (INCL. CONDITION) 8.2.1) RECOGNISING CHANGE 8.2.2) ESTABLISHING A SENSE OF PATTERN 8.2.3) RECOGNISING IDENTITY OF VEGETATION 9) GENERAL CONSIDERATIONS ON MONITORING & SAMPLE NUMBERS 9.1) CONSIDERATIONS THAT AFFECT THE NUMBER & LAYOUT OF SAMPLING SITES 9.1.1) CHOICE OF MONITORING OPTION 9.1.2) NUMBER OF VEGETATION UNITS BEING SAMPLED 9.1.3) THE LEVEL OF PLOT REPLICATION REQUIRED 9.2) TIMING OF SAMPLING 9.3) FREQUENCY OF SAMPLING 10) REFERENCES 11) APPENDICES APPENDIX 1: LIST OF ATTRIBUTES FOR A SHORT ASSESSMENT APPENDIX 2: AN EXTENDED LIST OF ATTRIBUTES APPENDIX 3: EXPANSION AND ADAPTATION OF THE KEIGHERY VEGETATION CONDITION SCALE & EQUIVALENTS INTO AN ARRAY OF SEPARATE ATTRIBUTES. APPENDIX 4: BUSHLAND SURVEY SHEETS FROM KEIGHERY 1994. APPENDIX 5: SOME TRIAL FIELD SHEETS APPENDIX 6: CARABAN FIELD TRIAL FINDINGS AND DATA ACKNOWLEDGEMENTS NVI Project members Nick Casson, Stephen Van Leeuwen, Susan Downes, Anne Harris, Judith Harvey, Kristen Bleby, Marc Wohling, Mark Garkaklis, Leah Firth, Sophie Moller, Keith Claymore DEC information, input and support Alan Danks, Allan Willis, Amanda Fairs, Anthea Jones, Barbara Wilson, Carolyn Harding, Chris Dunne, Christine Freegard, Cressida Wilson, David Mickle, Deborah Harding, Deon Utber, Geoff Young, Graeme Behn, Greg Durrell, Janine Kinloch, Kate Brown, Katherine Zdunic, Ken Atkins, Ken Wallace, Kevin Vear, Kim Friedman, Lisa Wright, Maria Lee, Mick Davis, Monica Hunter, Neil Gibson, Nicholas Woolfrey, Paul Cory, Paul Rampant, Paul Van Heurck, Penny Hussey, Richard Robinson, Ryan Vogwill, Sarah Barrett, Sarah Comer, Steve Appleyard, Teagan Johnston, Val English, Vanessa Clarke, Verna Tunsell, Wendy Chow, Wendy Thompson External Cindy Ong, Muriel Davies, Paul Novelly, Ray Froend, Tim Munday PURPOSE AND SCOPE This manual has vegetation condition at its core. It has regard for a number of recognisable factors that may influence vegetation condition, or the perception of vegetation condition. It also has regard for how this inevitably interacts with other, inter-related, facets of vegetation assessment, and with how they may be distilled and incorporated as required. Consequently, it is meant to be used incrementally. The extent to which the various elements are deployed will depend on the task at hand and the intent of the assessor. 1) INTRODUCTION This manual covers the establishment of quadrat-based monitoring to determine native vegetation condition. Vegetation condition is discussed in Literature Review: Vegetation Condition Assessment, Monitoring and Evaluation . 1.1) MAINTENANCE OF VEGETATION - CYCLICAL VERSUS DIRECTIONAL (OFTEN ADVERSE) CHANGE A good introduction is provided in two existing references as follows: Change is a natural feature of native vegetation. “Individual plants grow, flower and shed seed. Plants age and die, to be replaced by others, either from seed or by vegetative growth. Changes in a plant community may be cyclic or directional. Broadly speaking, natural cycles in the plant community may be maintained by recurring events such as fire or flood. After these, species already in the habitat, perhaps as soil-stored seed, are recruited and the vegetation gradually regains its former structure. Depending on the sequence of events and weather conditions, slightly different combinations of species may be favoured, but the plant community is essentially self-maintaining. Directional change may be induced by habitat disturbance which alters soil and water properties, and allows recruitment of species not native to the habitat, in particular, exotic weeds. Such changes generally lead to bushland deterioration.” (Benson and Howell 1990). The extent of the change will be dependent on the nature, extent, and duration, of the disturbance, and the associated pace of change. “Activities that adversely affect the self-maintenance of bushlands are commonly called disturbance factors. Such factors include partial clearing, fragmentation, selective removal of species (for example, timber cutting, wildflower picking, mowing), dieback, fire regime, ‘enrichment planting’ (that is planting of species not found in that plant community), weed invasion, animal impact (horses, foxes, rabbits, cats, dogs), soil movement (both removal and dumping), changes in water regimes (flooding, drainage and watering), rubbish dumping, mining (particularly mining for roadworks, grazing (stock, overgrazing by native mammals), proliferation of tracks (fire-breaks and walk trails), off-road vehicle use, use of service corridors (for electricity, gas, roads and water), fertiliser drift and pesticide or nutrient influx along waterways.” “Not all of these factors have the same level of impact but generally they are interrelated, the presence of one type of disturbance leading to further disturbance. The observed disturbance is most often the cumulative result of a series of compounding disturbance factors.” “An assessment of disturbance, in relation to the ability of the bushland to self- maintain, has been used as the basis of defining condition ratings for plant communities. …” Department of Environmental Protection (2000). 1.2) ISOLATING THE SIGNS AND SYMPTOMS OF CHANGE This sets the stage for the central conundrum/s in condition analysis - ‘How is condition and especially directional change (or an adverse trend) in condition to be separated from characteristics of a vegetation type that have little to do with condition in other words how are symptoms of adverse change to be separated from characteristics of identity (and the usual processes which maintain identity)?’ There is no easy answer to this. It requires due diligence in terms of assembling sufficient background (including the current & historical context) on the vegetation type in question to be able to make an informed judgement of how it fits into a spectrum of states. Where the vegetation type is very small the next closest equivalent type may have to suffice. Overall this can be approached in a number of ways. They all have to do with building a strong context for the vegetation and a strong sense of its identity. The most objective is to assess full plant diversity from many sites roundabout and then look at the balance of native species and weeds in light of disturbance (ideally this would be assessed numerically). There are other ways of establishing context and achieving reference points. Many national systems use a benchmark system with information on reference states for set vegetation types (however, this has its limitations, especially in light of the high diversity of species and communities in W.A. and the logistics of setting up such a system, but also due to an incomplete state of knowledge, flux in taxonomy, and differences in scale and voracity of information). There is much value in many years experience of vegetation survey and assessment which can clearly help inform an understanding of condition (but this does not mean that this comes without individual bias). An intermediate approach shadows the sort of approach that those with long- standing experience of vegetation assessment would use. It employs a mixture of: i) reference material ii) preliminary reconnaissance and observation and iii) cultivating a sense of the polarities of state and whether these are due to resident processes or imposed disturbance. It takes time to develop a sense of ecological pattern and process. Unfortunately the whole exercise is likely to get progressively more difficult for each generation as the drift in condition continues. 1.3) A PLETHORA OF FACTORS This endeavour is like making a diagnosis.
Recommended publications
  • Jeremy Bentham, Werner Stark, and 'The Psychology of Economic Man'
    Jeremy Bentham, ‘The Psychology of Economic Man’, and Behavioural Economics Michael Quinn (Bentham Project, UCL. [email protected]) Résumés English Francais § 1 briefly reviews first the received interpretation of Bentham, which sees him as having had little to do with the development of economics (excepting some passing mentions which recognize his deployment of the concept of utility or his reduction of human motivation to self-interest, and perhaps a note on his discussion of the concept of diminishing marginal utility); and second, the manner in which he applies his concept of rationality to political economy. In § 2, the central thesis of the paper is presented: it is argued that an examination of his insights into the psychology of individual choice supplies good reasons to identify him as an intellectual godfather of behavioural economics. In keeping with the normativity of his concept of rationality, Bentham would maintain that the way in which traditional economics continues to ignore the gulf between its model of human decision-making and the facts of human psychology weakens its usefulness both as a science and as a guide to public policy. Bentham anticipated several modifications to the standard model (for instance loss- aversion, the endowment effect, reference dependence, framing, the desire for cognitive ease, and status-quo bias) which have been introduced later by behavioural economics. § 3 introduces two problems concerning the normativity of economics, the first of which, at least for Bentham, rests upon a false premise,
    [Show full text]
  • Alyogyne Huegelii (Endl.) SCORE: -2.0 RATING: Low Risk Fryxell
    TAXON: Alyogyne huegelii (Endl.) SCORE: -2.0 RATING: Low Risk Fryxell Taxon: Alyogyne huegelii (Endl.) Fryxell Family: Malvaceae Common Name(s): lilac hibiscus Synonym(s): Hibiscus huegelii var. wrayae (Lindl.) HibiscusBenth. wrayae Lindl. Assessor: Chuck Chimera Status: Assessor Approved End Date: 3 May 2018 WRA Score: -2.0 Designation: L Rating: Low Risk Keywords: Shrub, Ornamental, Unarmed, Non-Toxic, Insect-Pollinated Qsn # Question Answer Option Answer 101 Is the species highly domesticated? y=-3, n=0 n 102 Has the species become naturalized where grown? 103 Does the species have weedy races? Species suited to tropical or subtropical climate(s) - If 201 island is primarily wet habitat, then substitute "wet (0-low; 1-intermediate; 2-high) (See Appendix 2) Intermediate tropical" for "tropical or subtropical" 202 Quality of climate match data (0-low; 1-intermediate; 2-high) (See Appendix 2) High 203 Broad climate suitability (environmental versatility) y=1, n=0 n Native or naturalized in regions with tropical or 204 y=1, n=0 y subtropical climates Does the species have a history of repeated introductions 205 y=-2, ?=-1, n=0 y outside its natural range? 301 Naturalized beyond native range y = 1*multiplier (see Appendix 2), n= question 205 n 302 Garden/amenity/disturbance weed n=0, y = 1*multiplier (see Appendix 2) n 303 Agricultural/forestry/horticultural weed n=0, y = 2*multiplier (see Appendix 2) n 304 Environmental weed n=0, y = 2*multiplier (see Appendix 2) n 305 Congeneric weed n=0, y = 1*multiplier (see Appendix 2) n 401 Produces
    [Show full text]
  • Evaluation of Several Pre-Clinical Tools for Identifying Characteristics
    Purdue University Purdue e-Pubs Open Access Theses Theses and Dissertations 12-2016 Evaluation of several pre-clinical tools for identifying characteristics associated with limb bone fracture in thoroughbred racehorses Anthony Nicholas Corsten Purdue University Follow this and additional works at: https://docs.lib.purdue.edu/open_access_theses Part of the Veterinary Medicine Commons Recommended Citation Corsten, Anthony Nicholas, "Evaluation of several pre-clinical tools for identifying characteristics associated with limb bone fracture in thoroughbred racehorses" (2016). Open Access Theses. 841. https://docs.lib.purdue.edu/open_access_theses/841 This document has been made available through Purdue e-Pubs, a service of the Purdue University Libraries. Please contact [email protected] for additional information. EVALUATION OF SEVERAL PRE-CLINICAL TOOLS FOR IDENTIFYING CHARACTERISTICS ASSOCIATED WITH LIMB BONE FRACTURE IN THOROUGHBRED RACEHORSES by Anthony N. Corsten A Thesis Submitted to the Faculty of Purdue University In Partial Fulfillment of the Requirements for the degree of Master of Science in Basic Medical Sciences Department of Basic Medical Sciences West Lafayette, Indiana December 2016 ii THE PURDUE UNIVERSITY GRADUATE SCHOOL STATEMENT OF THESIS APPROVAL Dr. Russell P. Main, Chair Department of Basic Medical Sciences Dr. Timothy B. Lescun Department of Veterinary Clinical Sciences Dr. Joseph M. Wallace Department of Biomedical Engineering Approved by: Dr. Laurie A. Jaeger Head of the Departmental Graduate Program iii TABLE OF
    [Show full text]
  • First Steps Towards a Floral Structural Characterization of the Major Rosid Subclades
    Zurich Open Repository and Archive University of Zurich Main Library Strickhofstrasse 39 CH-8057 Zurich www.zora.uzh.ch Year: 2006 First steps towards a floral structural characterization of the major rosid subclades Endress, P K ; Matthews, M L Abstract: A survey of our own comparative studies on several larger clades of rosids and over 1400 original publications on rosid flowers shows that floral structural features support to various degrees the supraordinal relationships in rosids proposed by molecular phylogenetic studies. However, as many apparent relationships are not yet well resolved, the structural support also remains tentative. Some of the features that turned out to be of interest in the present study had not previously been considered in earlier supraordinal studies. The strongest floral structural support is for malvids (Brassicales, Malvales, Sapindales), which reflects the strong support of phylogenetic analyses. Somewhat less structurally supported are the COM (Celastrales, Oxalidales, Malpighiales) and the nitrogen-fixing (Cucurbitales, Fagales, Fabales, Rosales) clades of fabids, which are both also only weakly supported in phylogenetic analyses. The sister pairs, Cucurbitales plus Fagales, and Malvales plus Sapindales, are structurally only weakly supported, and for the entire fabids there is no clear support by the present floral structural data. However, an additional grouping, the COM clade plus malvids, shares some interesting features but does not appear as a clade in phylogenetic analyses. Thus it appears that the deepest split within eurosids- that between fabids and malvids - in molecular phylogenetic analyses (however weakly supported) is not matched by the present structural data. Features of ovules including thickness of integuments, thickness of nucellus, and degree of ovular curvature, appear to be especially interesting for higher level relationships and should be further explored.
    [Show full text]
  • St Patrick's International College Ltd, November 2015
    Higher Education Review (Alternative Providers) of St Patrick's International College Ltd November 2015 Contents About this review ..................................................................................................... 1 Amended judgements October 2016 ...................................................................... 2 Key findings .............................................................................................................. 3 QAA's judgements about St Patrick's International College Ltd ............................................. 3 Good practice ....................................................................................................................... 3 Recommendations ................................................................................................................ 3 Theme: Digital Literacy ......................................................................................................... 4 About St Patrick's International College Ltd .......................................................... 5 Explanation of the findings about St Patrick's International College Ltd ........... 7 1 Judgement: The maintenance of the academic standards of awards offered on behalf of the awarding organisation ............................................................................... 8 2 Judgement: The quality of student learning opportunities ............................................. 20 3 Judgement: The quality of the information about learning opportunities ......................
    [Show full text]
  • King George VI & Queen Elizabeth Stakes (Sponsored by QIPCO)
    King George VI & Queen Elizabeth Stakes (Sponsored by QIPCO) Ascot Racecourse Background Information for the 65th Running Saturday, July 25, 2015 Winners of the Investec Derby going on to the King George VI & Queen Elizabeth Stakes (Sponsored by QIPCO) Unbeaten Golden Horn, whose victories this year include the Investec Derby and the Coral-Eclipse, will try to become the 14th Derby winner to go on to success in Ascot’s midsummer highlight, the Group One King George VI & Queen Elizabeth Stakes (Sponsored by QIPCO), in the same year and the first since Galileo in 2001. Britain's premier all-aged 12-furlong contest, worth a boosted £1.215 million this year, takes place at 3.50pm on Saturday, July 25. Golden Horn extended his perfect record to five races on July 4 in the 10-furlong Group One Coral- Eclipse at Sandown Park, beating older opponents for the first time in great style. The three-year-old Cape Cross colt, owned by breeder Anthony Oppenheimer and trained by John Gosden in Newmarket, captured Britain's premier Classic, the Investec Derby, over 12 furlongs at Epsom Downs impressively on June 6 after being supplemented following a runaway Betfred Dante Stakes success at York in May. If successful at Ascot on July 25, Golden Horn would also become the fourth horse capture the Derby, Eclipse and King George in the same year. ËËË Three horses have completed the Derby/Eclipse/King George treble in the same year - Nashwan (1989), Mill Reef (1971) and Tulyar (1952). ËËË The 2001 King George VI & Queen Elizabeth Stakes saw Galileo become the first Derby winner at Epsom Downs to win the Ascot contest since Lammtarra in 1995.
    [Show full text]
  • Fire As an Evolutionary Pressure Shaping Plant Traits
    Opinion Fire as an evolutionary pressure shaping plant traits Jon E. Keeley1,2, Juli G. Pausas3, Philip W. Rundel2, William J. Bond4 and Ross A. Bradstock5 1 US Geological Survey, Sequoia-Kings Canyon Field Station, Three Rivers, CA 93271, USA 2 Department of Ecology and Evolutionary Biology, University of California, Los Angeles, CA 90095, USA 3 Centro de Investigaciones sobre Desertificacio´n of the Spanish National Research Council (CIDE - CSIC), IVIA campus, 46113 Montcada, Valencia, Spain 4 Botany Department, University of Cape Town, Private Bag, Rondebosch 7701, South Africa 5 Centre for Environmental Risk Management of Bushfires, University of Wollongong, Wollongong, NSW 2522, Australia Traits, such as resprouting, serotiny and germination by parent adaptive value in fire-prone environments and the heat and smoke, are adaptive in fire-prone environ- extent to which we can demonstrate that they are adapta- ments. However, plants are not adapted to fire per se tions to fire. We also address the question of what fire- but to fire regimes. Species can be threatened when adaptive traits and fire adaptations can tell us about fire humans alter the regime, often by increasing or decreas- management. ing fire frequency. Fire-adaptive traits are potentially the result of different evolutionary pathways. Distinguishing Adaptive traits between traits that are adaptations originating in re- Adaptive traits are those that provide a fitness advantage sponse to fire or exaptations originating in response in a given environment. There are many plant traits that to other factors might not always be possible. However, are of adaptive value in the face of recurrent fire and these fire has been a factor throughout the history of land- vary markedly with fire regime.
    [Show full text]
  • QUINN Barn 40 Hip No. 3761
    Property of Iron County Farms, Inc. Barn Hip No. 40 QUINN 3761 Dark Bay or Brown Mare; foaled 2000 Northern Dancer Nureyev............................. Special Theatrical (IRE) ................. Sassafras (FR) Tree of Knowledge (IRE).... Sensibility QUINN Mill Reef Reference Point................. Home On the Range Checking It Twice (IRE)..... (1989) Key to the Mint Christmas Bonus .............. Sugar Plum Time By THEATRICAL (IRE) (1982). Champion grass horse, black-type winner of $2,840,500, Breeders' Cup Turf [G1], etc. Sire of 19 crops of racing age, 1005 foals, 758 starters, 76 black-type winners, 2 champions, 493 win- ners of 1463 races and earning $73,109,970. Among the leading brood- mare sires, sire of dams of 42 black-type winners, including champions English Channel, Numerous Times, Shillelagh Slew, Mi Amigo Guelo, Mis- sion Possible, Theatrical Award, and of Rail Link, Urban Street, Dublino. 1st dam CHECKING IT TWICE (IRE), by Reference Point. Unraced. Dam of 8 other registered foals, 8 of racing age, 8 to race, 5 winners, including-- Final Decision (g. by Royal Academy). 7 wins, 3 to 8, $79,219. Another King (g. by King Cugat). 3 wins at 3 and 4, 2009, $14,963, in Can- ada. (Total: $13,995). 2nd dam CHRISTMAS BONUS, by Key to the Mint. 7 wins, 3 to 5, $140,557, Poques- sing H. Dam of 15 foals, 12 to race, 11 winners, including-- CHRISTMAS GIFT (f. by Green Desert). 7 wins, 2 to 4, $387,176, Beaugay H. [G3], etc. Dam of CHRISTMAS KID (f. by Lemon Drop Kid, $596,- 877, Ashland S. [G1] (KEE, $310,000), etc.).
    [Show full text]
  • Pedigree Insights
    Andrew Caulfield, October 17, 2006–Teofilo (Ire) P EDIGREE INSIGHTS The enormity of the task ahead of Teofilo mustn=t be under-estimated, as only two colts have managed to BY ANDREW CAULFIELD win even two legs of the Triple Crown since Nijinsky (Reference Point added the St Leger to his Derby DARLEY DEWHURST S.-G1, ,250,000, Newmarket, success in 1987, while Nashwan took the 2000 10-14, 2yo, c/f, 7fT, 1:26.12, gd/sf. Guineas and Derby two years later). The Triple Crown 1--TEOFILO (IRE), 127, c, 2, by Galileo (Ire) remains an exciting possibility, though, and Teofilo=s 1st Dam: Speirbhean (Ire) (SW-Ire), by Danehill record suggests that he could develop into that 2nd Dam: Saviour, by Majestic Light one-in-a-million colt blessed with the necessary 3rd Dam: Victoria Queen, by Victoria Park brilliance, versatility and toughness. Like Nijinsky, his O-Mrs J Bolger; B/T-J Bolger; J-K Manning; ,141,950. juvenile record stands at five wins from five starts and Lifetime Record: G1SW-Ire, 5-5-0-0, ,349,515. both colts gained their fifth success in the G1 Dewhurst S. However, the signs are that Jim Bolger, who also Click for the Racing Post chart or the free brisnet.com bred Teofilo, isn=t looking too far ahead, as he is now catalogue-style pedigree. considering sending his star colt to France for the Criterium International on October 29. If American racing thinks it has been hard done by, Nijinsky was a member of the second crop by with only three Triple Crown winners since Citation in Northern Dancer, a winner of the first two legs of the 1948 and none since Affirmed in 1978, spare a thought Triple Crown, whereas Teofilo comes from the second for the British fans.
    [Show full text]
  • Organogenesis and Vasculature of Anaxagorea and Its Implications For
    bioRxiv preprint doi: https://doi.org/10.1101/2020.05.22.111716; this version posted June 19, 2020. The copyright holder for this preprint (which was not certified by peer review) is the author/funder. All rights reserved. No reuse allowed without permission. 1 Organogenesis and Vasculature of Anaxagorea and its Implications for the 2 Integrated Axial-Foliar Origin of Angiosperm Carpel 3 4 Ya Li, 1,† Wei Du, 1,† Shuai Wang,2 Xiao-Fan Wang1,* 5 1 College of Life Sciences, Wuhan University, Wuhan 430072, China 6 2 College of Life Sciences and Environment, Hengyang Normal University, 7 Hengyang 421001, China 8 *Author for correspondence. E-mail: [email protected] 9 † Both authors contribute equally to this work 10 11 Running Title: The Axial-Foliar Origin of Carpel 12 1 bioRxiv preprint doi: https://doi.org/10.1101/2020.05.22.111716; this version posted June 19, 2020. The copyright holder for this preprint (which was not certified by peer review) is the author/funder. All rights reserved. No reuse allowed without permission. 13 Abstract 14 The carpel is the definitive structure of angiosperms, the origin of carpel is of great 15 significance to the phylogenetic origin of angiosperms. Traditional view was that 16 angiosperm carpels were derived from structures similar to macrosporophylls of 17 pteridosperms or Bennettitales, which bear ovules on the surfaces of foliar organs. In 18 contrast, other views indicate that carpels are originated from the foliar appendage 19 enclosing the ovule-bearing axis. One of the key differences between these two 20 conflicting ideas lies in whether the ovular axis is involved in the evolution of carpel.
    [Show full text]
  • Molecular Phylogenetic Relationships Among Members of the Family Phytolaccaceae Sensu Lato Inferred from Internal Transcribed Sp
    Molecular phylogenetic relationships among members of the family Phytolaccaceae sensu lato inferred from internal transcribed spacer sequences of nuclear ribosomal DNA J. Lee1, S.Y. Kim1, S.H. Park1 and M.A. Ali2 1International Biological Material Research Center, Korea Research Institute of Bioscience and Biotechnology, Yuseong-gu, Daejeon, South Korea 2Department of Botany and Microbiology, College of Science, King Saud University, Riyadh, Saudi Arabia Corresponding author: M.A. Ali E-mail: [email protected] Genet. Mol. Res. 12 (4): 4515-4525 (2013) Received August 6, 2012 Accepted November 21, 2012 Published February 28, 2013 DOI http://dx.doi.org/10.4238/2013.February.28.15 ABSTRACT. The phylogeny of a phylogenetically poorly known family, Phytolaccaceae sensu lato (s.l.), was constructed for resolving conflicts concerning taxonomic delimitations. Cladistic analyses were made based on 44 sequences of the internal transcribed spacer of nuclear ribosomal DNA from 11 families (Aizoaceae, Basellaceae, Didiereaceae, Molluginaceae, Nyctaginaceae, Phytolaccaceae s.l., Polygonaceae, Portulacaceae, Sarcobataceae, Tamaricaceae, and Nepenthaceae) of the order Caryophyllales. The maximum parsimony tree from the analysis resolved a monophyletic group of the order Caryophyllales; however, the members, Agdestis, Anisomeria, Gallesia, Gisekia, Hilleria, Ledenbergia, Microtea, Monococcus, Petiveria, Phytolacca, Rivinia, Genetics and Molecular Research 12 (4): 4515-4525 (2013) ©FUNPEC-RP www.funpecrp.com.br J. Lee et al. 4516 Schindleria, Seguieria, Stegnosperma, and Trichostigma, which belong to the family Phytolaccaceae s.l., did not cluster under a single clade, demonstrating that Phytolaccaceae is polyphyletic. Key words: Phytolaccaceae; Phylogenetic relationships; Internal transcribed spacer; Nuclear ribosomal DNA INTRODUCTION The Caryophyllales (part of the core eudicots), sometimes also called Centrospermae, include about 6% of dicotyledonous species and comprise 33 families, 692 genera and approxi- mately 11200 species.
    [Show full text]
  • FW Monitoring Program Method Manual for the Reference Point Survey
    104 TFW -AM9-98-002 TFW Monitoring Program METHOD MANUAL for the REFERENCE POINT SURVEY RP #11 by: Allen E. Pleus Dave Schuett-Hames Indi:an Fisheries Commission May 1998 TFW MOlJitoring Program Manua/- May 1998 Pleus, A.E. and D. Schuett-Hames. 1998. TFW Monitoring Program methods manual for the reference point survey. Prepared for the Washington State Dept. of Natural Resources under the Timber, Fish, and Wildlife Agreement. TFW-AM9-98-002. DNR#104. May. Abstract This manual provides a standard method for establishing stable reference point sites for monitoring stream segments over time. Reference points are established at regular intervals along a previously defined stream segment and monumented to be easily relocated. Stream parameters collected during this survey include: I) segment length; 2) bankfull width; 3) bankfull depth; 4) canopy closure; and 5) optional reference photographs. The manual is divided into pre-survey prepa­ ration, field methods, post-field documentation, and data management sections. An extensive appendix section includes a survey task checklist copy master, a materials and equipment source list, field form copy masters, examples of com­ pleted field forms, a data report example, and a glossary of terms. TFW Monitoring Program Washington Dept. of Natural Resources Northwest Indian Fisheries Commission Forest Practices Division: CMER Documents 6730 Martin Way E. P.O. Box 47014 Olympia, WA 98516 Olympia, WA 98504-7014 Ph: (360)438-1180 Ph: (360)902-1400 Fax: (360)753-8659 Internet: http://www.nwifc.wa.gov Reference Point Survey TFW Monitoring Program Mallual- May 1998 The Authors Acknowledgements Allen E. Pleus is the Lead Training and Quality The development of this document was funded by Assurance Biologist for the TFW Monitoring Pro­ the Timber-Fish-Wildlife (TFW) Cooperative gram at the Northwest Indian Fisheries Commis­ Monitoring, Evaluation, and Research (CMER) sion.
    [Show full text]