SDGG, Heft 65 - Litho- und Biostratigraphie der außeralpihen Kreide von Bayern ^TE0J1^> 11 ] ^-~> L_JL-J Cenomanian-Coniacian ammonoids of the DanubianCretaceous Group (Bavaria, southern Germany) I
MARKUS WILMSEN ', CHISTOPHER J.JTöocT2, BiRGIF^TEBUHR^-ÄT ULRICH KAPLAN^, 1 Scnckenberg~Käturhistorischc SarnmTüngen Dresden/Museum für Mineralogie und Geologie. Königsbrücker Landstr. 159, D-Ol 109 Dresden,
Abstract . — This paper presents a compilation of the ammonoid records of the Danubian Cretaceouos Group, supplemented with new data on their taxonomy and biostratigraphy. The study is based on material of the Museum für Mineralogie und Geologie in Dresden, the Bayerische Staatssammlung für Paläontologie in Munich and the Juramuseum Eichstätt as well as a taxonomic re-evaluation of the ammonoids described and figured by DACQUE (1939). Unfortunately, the ammonoids of the Danubian Cretaceouos Group are generally rare and of low diversity. Furthermore, they arc often poorly preserved and, thus, several specimens have to be kept in open nomenclature. In total, twenty-four Ccnoma- nian, Turonian and Coniacian ammonoid taxa are recorded: Manielliceras inantelli (J. SOWERBY, 1814), Calycoceras {Calycoceras)naviculare (MANTELL, 1822), Metoicoceras geslinianum (D'ORBIONY, 1850), Euomphalocerus septemse- rialum (CKAGIN, 1893), Pseiidocalycoceras angolaense(SPÄTH, 1931),Ps. cf. harpax (SIOLICZKA, 1864), Romaniceras (Romaniceras) kallesi! (ZAZVORKA, 1958), Vascocerasdiartianum (D'ORBIGNY, 1850). Lewesicerasperamplum (MAN- TKU., 1822), he. manteM WRIGHT & WRIGHT, 1951, Collignonic eras'! woollgari1? (MANTELL, 1822), Subprionocyclus sp„ Su. neptuni'! (GEIMTZ, 1849), Peroniceras (Peroniceras) westphaticum (STROMBECK, 1859). Ptacenticeras! sp., Worthoceras vermiculum (SHUMARD, I860), Hemiptyehoceras sp., He. cf. reesidei COBBAN & Scon, 1972, Scipono- ceras sp.. Sc. gracile? (SHUMARD, 1860), Sc. bohemicum? (FRITSCH, 1872), Bacuiites sp., Scaphites sp. and Scaphites kieslingwaldensis doylef! WRIGHT, 1978. Most taxa are illustrated, briefly described and placed within the new litho- and biostratigraphie framework of the Danubian Cretaceouos Group (see NIEBLHR el al. 2009; TROGLR et aL 2009).
Keywords: Southern Germany. Bavaria, ammonoids, taxonomy, biostratigraphy. Upper Cretaceous, Cenomanian, Turonian, Coniacian
Kurzfassung Diese Arbeit kombiniert das Auftreten von Ammonoideen der Danubischen Kreide-Gruppe mit neuen Daten bezüg lich ihrer Taxonomie und Biostratigraphie. Sie basiert auf Material aus dem Museum für Mineralogie und Geologie in Dresden, der Bayerischen Staatssammlung für Paläontologie in München und dem Juramuseum Eichstätt. Auch die von DACÜ-UL (1939) beschriebenen und abgebildeten, spater verloren gegangenen Ammonoideen wurden einer taxonomischen Neubetrachtung unterzogen. Ammonoideen sind in der Danubischen Kreide-Gruppe jedoch generell sehen und gering divers. Weiterhin sind sie oftmals schlecht erhalten, sodass eine nicht geringe Anzahl der Stücke in offener Nomenklatur verbleiben muss. Insgesamt wurden vierundzwanzig Ammonoideen-Taxa aus dem Cenomanium, TuronJum und Coniacium bestimmt: Mantetliceras mantelli (J. SOWERBY, 1814), Calycoceras {Calycoceras) navkulare (MANTELL, 1822), Metoicoceras gestinianum (D'ORBIGNY, 1850), Euomphalocerus septemseriatum (CRAGIN, 1893). Pseudoculycoceras angolaense (SPÄTH; 1931), Ps. cf. harpax (STOLICZKA, 1864), Romaniceras (Romaniceras) kullesi'! (ZAZVORKA, 1958), Vascoceras diartianum (D'ORBIGNY, 1850), Lewesiceras perumplum (MAN ILI.L, 1822),/.f. mantelli WRIGHT & WRIGHT, 1951, Coltignoniceras! woollgari? (MANTELL, 1822), Subprionocyelus sp.. Su, neptum! (GHMI/. 1849), Peroniceras (Peroniceras) westphaticum (STROMBLCK, 1859), Ptaeenticeras'! sp., Worthocems vermiculum (Sm - MARD. 1860), Hemiptyehoceras sp., He. cf. reesidei COBBAN & SCOTT, 1972, Sciponoceras sp.. Sc. gracile! (SHUVIAKD, 1860),Sc. bohemicum'!(FRITSCH, 1872), Bacidiles sp., Scaphites sp. und ScaphiteskieslingwaldensisdoyleP. WKIGIII, 1978. Die meisten Taxa wurden abgebildet, beschrieben und in die neue litho- und biostraiigraphische Gliederung der Danubischen Kreide-Gruppe (siehe NIKHI.HR et al. 2009; TRÖGER et al. 2009) integriert.
Schlüsselworte: Süd-Deutschland, Bayern, Ammoniren, Taxonomie, Biostratigraphie, Überkreide, Cenomanium, Turonium, Coniacium 112 SDGG, Heft 65 - Litho- und Biostratigfaphie der außeralpinen Kreide von Bayern
11ntroduction 1.1 Geological setting Despite nearly 200 years of palaeontological and stra- During the Late Cretaceous transgressions, vast areas of tigraphic research on the Cretaceous strata that crop the northern Tethyan margin were flooded and became out north of the Alpine Molasse in northern Bavaria peri-continental shelf seas. In northern Bavaria (sou (formerly often termed "Regensburger Kreide", now thern Germany), this development is exemplified by the formalized within the Danubian Cretaceous Group; onlap of the Danubian Cretaceous Group (NIEBUHR et al. 2009; WILMSEN & NIEBUHR 2010). The constituent see NIEBUHR et al. 2009), their biostratigraphy is still somewhat crude. This is mainly due to the fact that formations of this group represent non-marine to neritic ammonoids, the classic index fossils, are relatively environments, comprising conglomerates, sands and rare and important collections such as those of TRUS- sandstones, clays, marls and marlstones, calcarenites, HEIM (1935) and DACQUE (1939) were for the most part siliceous opoka and limestones, with a total thickness lost during the World War II. Furthermore, the out of 300 to 500 m. Terrestrial deposits are known from crop conditions are very poor and the preservation of the Lower Cretaceous (Schutzfels Formation) and range the fauna is often poor. Consequently the records of from the Middle Turanian into the Coniacian (or Santo- ammonites from the Danubian Cretaceous Group are nian?, Hessenreuth Formation). Marine units start with sparse and scattered in the literature, and there have the Lower Cenomanian and persist into the Coniacian. been few taxonomic revisions. The complete succession documents a nearly symme trical transgressive/regressive cycle with a maximum The scope of the present paper is a compilation of flooding interval during the late Middle to early Late the available ammonoid records supplemented with Turanian (NIEBUHR et al. 2009). new data on taxonomy and biostratigraphy. For this purpose, collection material of the Museum für Mine From nine formations of the Danubian Cretaceous ralogie und Geologie in Dresden, the Bayerische Group which exclusively consist of marine sediments Staatssammlung für Paläontologie in Munich and the (see NIEBUHR et al. 2009), seven yielded ammonoids: Juramuseum Eichstätt has been examined. For lost The Regensburg Formation (lower Lower Cenomanian specimens such as most of the ammonoids described to Upper Cenomanian in the Regensburg-Kelheim by DACQUE (1939), a taxonomic re-evaluation based on area, Fig 1/1) is characterized by strong terrigenous the illustrations is provided where possible. input and glauconitic-bioclastic mixed sediments with a shallow-water fauna which accumulated mainly in the
. 44|.. 45. ; IK 251 Oanubiau Cretaceous Group Cretaceous below molasse
Fig. 1: Distribution of the Danubian Cretaceous Group in Bavaria, and key areas with different lithological development (according to NIEBUHR et al. 2009): (I) type region Regensburg-Kelheim, (II) proximal Bodenwöhrer Senke, (III) siliceous Wellheim Formation west of Ingol stadt, (IV) Cretaceous of the Franconian Alb northeast of Nürnberg, (V) terrestrial Hessenreuth Formation, (VI) Sandbach Formation west of Passau .in (I) Regensburg-Kelheim (VI) Ortenburg near Passaul •o nodules of siliceous limestone sandstone tilled burrows 0^r* TUM » • conglomerate <> 5 S hioturbation ammonite heteramorph ammonites ^) nautiloids •-s shells bioclasts /Sietoicocerasgeslinianum, Catycocerws (Ca.) navicular?, t Euomphatocerax *eptentseriarnm. Pseudocalyeoctrus angolacnse. jPs cf.harpax, Vascoceras diartianum, IVorihoceras vermicutum. j \ihjmiptychoceras sp., He. cf. reesidvi, Sciponoceras sp., Sc gracile'] "3 S '^Calycoceras naviculare CC 2 ^Acanthoceras rhulomagense ) Siantelticeras mantelli y?MT] (Sch u tzfels Fig. 2: Standard section of the Danubian Cretaceous Group in the Regensburg-Kelheim area (left, see Fig. 1/1) and the Orten burg area west of Passau (right, see Fig. 1/VI) with indication of ammonite horizons. 114 SDGG. Heft 65 - Litho- und Biostratigraphie der außeralpinen Kreide von Bayern northeastern parts of the basin, close to the emergent mannshofer Sandstein (= Jeding Formation). TRUSHIIM Variscan source areas (Fig. 1). The following ammo (1935), in his influential stratigraphic revision of the nite- and planktonic foraminifera-bearing uppermost Lower Cretaceous and Cenomanian of the Danubian Cenomanian-lower Lower Turonian silty marls of Cretaceous Group, listed some ammonites from the the Eibrunn Formation document a first maximum Regensburg and Eibrunn formations, however, without flooding interval. This formation is overlain by the figuring them. DACQUE (1939) presented a monogra spiculitic siltstones (Reinhausen Member) and sand phic revision of the macro-invertebrate fauna of the stones (Knollensand Member) of the Lower Turonian LIpper Cretaceous of the Regensburg- Kelheim area, Winzerbcrg Formation, which grades into coarse including several species of ammonoids. HALNER grained sandstones of the so-called "Hornsand'', and (1971) presented detailed macrofaunal lists from the is capped by a conspicuous sequence boundary. The Upper Cretaceous of the Regensburg-Kelheim area Middle lower Upper Turonian Kagerhöh Formation and mentioned a few species of ammonites. starts with the transgressive "Homsand" facies, rapidly fining-upward into silty-sandy marls and nodular After several decades of scientific stasis in macropa- marly limestones (Eisbuckel Member), followed by laeontological research, FÖRSTER et al. (1983) provi thick-bedded limestones of the Pulverturm Member ded important new data on the Upper Cenomanian and the silty-sandy marls of the Karthaus Member. ammonite fauna of the Eibrunn Formation north of The members are separated by prominent unconformi Regensburg, based on new finds from the type loca ties represented by the so-called "untere" and "obere lity Benberg road-cutting section. A few years later, Glaukonitmergel-Bank". The upper Middle to lower RÖPER & ROTHGAENGER (1995) supplemented this work most Upper Turonian Pulverturm Member is regarded with a description of material collected from the same as the maximum flooding interval of the Danubian horizon in the Bad Abbach area south of Regensburg. Cretaceous Group (NIEBUHR et al. 2009). At a major In the same year, RÖPER & NEUMEIER (1995) mentioned unconformity, the Kagerhöh Formation is truncated the occurrence of Lewesiceras sp. from the Großberg and erosionally overlain by the widespread Großberg Formation of the Eggmühl area south of Regensburg. Formation (upper Upper Turonian). Marine sedimen tation of the Danubian Cretaceous Group ended with the marly Hellkofen Formation and the sandy Jeding Formation (Coniacian). In the southeastern part of the 2 Taxonomy distribution area of the Danubian Cretaceous Group In the following, the ammonite records of the Danubian near Passau (Fig. 1/VI). the sandy marls and marly- Cretaceous Group are listed by formations, in ascen spiculitic limestones of the Sandbach Formation with ding stratigraphical order. It has to be stressed that a lower Marterberg and an upper Buchleiten Member this paper is not systematic palaeontological account were deposited during the upper Upper Turonian to but more a taxonomie list with some comments on basal Middle Coniacian (NILBLUR et al. 2009; TRÖGER the current names and stratigraphic distribution of et al. 2009). the ammonoids. Most of the recorded taxa are well known from Cretaceous successions elsewhere, and thus, discussions and descriptions are short. 1.2 History of ammonite records The following abbreviations apply: Bayerische Staats sammlung für Paläontologie in München (BSP), In contrast to the greater part of the macrobenthic Museum für Mineralogie und Geologie in Dresden fauna of the Danubian Cretaceous Group (see the (MMG), Juramuseum Eichstätt (JME). Open nomen monographs of. e.g., LEHNER 1937a, b; DACQUE 1939), clature follows BENGTSÜN (1988). Part of the fauna was the ammonoids were, unfortunately, not monographed coated with magnesium oxide prior to photography. and only rarely described and figured. The first to Taxonomy follows WRIGHT et al. (1996). mention ammonites from the Danubian Cretaceous Group were STOCKTIFIM (1852), GIMBEI (1854, 1868a, b) and GERSTER (1881). BRUNHLDER (1917), in his impor 2.1 Regensburg Formation tant regional account of the lithological succession Mantelliceras mantelli (J. SÜWERBY, 1814) - Fig. 3 around Regensburg, mentioned several ammonites, e.g. from the "Karthauser Baculitenmergel" (= Kar Remarks: Until the 1940s. the Regensburg Formation thaus Member of the Kagerhöh Formation). LEHNER was regarded as a Cenomanian lithostratigraphic unit (1935, 1937b) recorded ammonites from the Cretace which may extend down into the Lower Cenomanian ous of the Franconian Alb northeast of Nürnberg (Fig. (e.g., TRUSHEIM 1935; DACQUE 1939). However, star 1/V), i.e., the Betzensteiner Kreidekalk (-Betzenstein ting with the mapping survey of TILLMANN (1954) and Member of the Kagerhöh Formation) and the Hart- OSCH MA (1958), an exclusively Late Cenomanian SDGG, lieft 65 - Lilho- und Biostratigraphie der außeralpirien Kreide von Bayern 115 and Saal (WILMSEN & NIEBUHR 2010): Neohiholites became extinct in the middle part of the M iddle Ceno manian (COMBEMOREL et al. 1981: CHRISTIANSEN 1990). TRUSHEIM (1935: 87) mentioned (without figuring) Acanthoceras rhotomagen.se ( BRONHNIART) from 4-5 m below the top of the Regensburg Formation in the Muhlbcrg section near Bad Abbach, a record previously also given by GIMHI.E (1854: 38) for ano ther specimen from the Regensburg Formation of the Regensburg-Kelheim area. Unfortunately, these spe cimens could not be located in the collections. 2.2 Eibrunn Formation Melokoceras geslinianum (n'ORniciNY, 1850) Calycoceras (Calyioceras) navkulare ( MAM EL 1.1822) Euomphaloceras septemseriatum (CKAOIN. 1893) Pseudocalycoceras angolaense(SHAITI; 1931) |listed as Pseudoealycoceras dentonense (MOKEMAN. 1942) by Fig. 3: Mantelliceras mantelli(l. SOWERBY), X 2, Saal Member of the FÖRSTER el al. (1983)] Regensburg Formation, Kapfeiberg near Kelheim. BSP: ASXVII8. Pseudocalycoceras cf. harpax (STIH.IC7.KA, 1864) Vascoceras diartianum (D^ORIIIONY, 1S50) age for the Regensburg Formation became deeply ent Worthoceras vermiculum (SIIUMARD, I860) renched into the literature, ignoring the existing faunal liemiptychoceras cf. reesidei C'OHBAS & Scon, 1972 evidence for an older age (e.g., FÖRSTER et al. 1983; Hemiptychoceras sp. Risen 1983; ROM R& ROTHGAL;NC;ER 1995; Mi M R2000). Seiponoceras graeile'l (SIIUMARD, 1X60) I lowever, the macrofauna of the formation, especially Sciponoceras sp. - Fig. 4 the bivalves and regular echinoids, is typical of Lower to Middle Cenomanian arenaceous formations else where (e.g. the Lower Cenomanian Wilmington Sands of Devonshire in southern England: see MORTIMORE et al.. 2001 and references therein) and already DAC- i.n i (1939) described and figured unequivocal ino- ceramid bivalve evidence for an Early Cenomanian age of the lower pari of the Regensburg Formation in the Kelheim area (see TRÜGIR et al. 2009; WII.MSIA & Nil »IHR 2010). Finally, the Acunthocenis (Mantelli ceras) navkulare of DACQLIE (1939: pi. 4, Fig. 2, from the "Ccnonianer (irünsandstein, Kapfelberg bei Kel heim". BSF": AS XVII 8, also figured by FÖRSTER et al. 1983 as Calycocerus cf". navkulare), re-figured herein on Fig. 3. is clearly a Maiuelliceras mantelli, indicating the lower part of the Lower Cenomanian, rather than an Upper Cenomanian Calycocerus navkulare. The adherent coarse-grained glauconilic sandstone matrix suggests thai it is from the lower part of the Saal Mem ber of the formation, below the level with abundant Inoccranuis ex gr. virgatus Snii.t'TKR (i.e.. the upper Fig. i»: Sciponocerassp. aff. gracM (SWARD, 1860), x 2, Eibrunn lower Cenomanian Maiuelliceras di.xoni Zone). This Formation, Winzerer Höhen near Regensburg. BSP: AS XVII9. is confirmed by DACOTE (1939: 7-8). staling "Bei uns kommt A. navkulare MAM. mit P. asper LAM. in der Remarks: The Eibrunn Formation yields the richest tieferen Cininsandslcinslulc. nicht in der oberen kal ammonoid fauna of the Danubian Cretaceous Group. kigen vor." Supportive evidence for an Early to early All the ammonoids are from a dm-thin interval in the Middle Cenomanian age of the Saal Member of the lower part of the formation below the so-called "Kalk- Regensburg Formation also comes (Vom finds of the mergelbank" which also yields the belemniie Praeuc- belemniie genus Neohiholites STOLEEY from Kelheim tinocamaxplenus(BI.AINVM.I i) (D-u ou. 1939; FORSTI R 116 SDGü, Heft 65 Litho- und Biostratigraphie der außeralpincn Kreide von Bayern et al. 1983; Ron K & ROTHGAENGER 1995; WII.MSF.N et Vascoceras diartianum (FÖRSTER et al. 1983: pi. 3, figs al. in prep.). The fauna characterizes the Metoicoceras 1-5; RÖPER & ROTHGAENGER 1995: 182), however, are gesliniamtm Zone of the upper (but not uppermost) common. Heteromorph ammonites are represented by- Upper Cenomanian (see discussion in FÖRSTER et al. very common Wonhoceras vermiculum (FÖRSTER et al. 1983) and was studied by MW and BN in the BSP. 1983: pi. 1, figs 1-9; RÖPER & ROTHGAENGER 1995: 184). The zonal index, Metoicoceras gesliniamtm (figured Hemiptyehoceras cf. reesidei (FÖRSTER et al. 1983: pi. as Pulclielliagesliniana D'ORU. by DACQUE 1939: pi. 6, 3, figs 9, 10) and rare Hemiptyehoceras sp. as well figs 3, 4 from the Winzerer Höhen near Regensburg) as representatives of the genus Sciponoceras HYATT. was described and illustrated by FÖRSTER et al. (1983: 1894 (see FÖRSTER et al. 1983: 130). Noteworthy is the pi. 3, figs 12-16) from the Benberg section and by specimen figured as Baculites subbaculoides GEIN. by ROPER. & RUMIGAENGER (1995: 183) from Bad Abbach. DACQUE (1939: pi. 6, fig. 6) from the Eibrunn Forma It is associated with fairly common Calycoceras navi- tion of the Winzercr Höhen near Regensburg which is culare (see FÖRSTER et al. 1983: pi. 2, figs 8-12) and less refigured herein in Fig. 4. This specimen may repre common Euumphaloceras sepiemseriatum [already sent Sciponoceras gracile? (SHUMARD, 1860), based figured as Acanthoceras (Metacanthoplites) rhoto- on the presence of well-spaced, shallow, oblique ribs magense DEER, by DACQUE 1939: pi. 6, fig. 5 from the on the flanks. However, due to the poor preservation, Winzerberg section, and by FORSTER et al. 1983: pi. 3, the specific assignment is very doubtful. ligs 6 8 from Benberg]. Pseudocalycoceras angola ense (see FÖRSTER et al. 1983: pi. 2, figs 1-4. and RÖPER The stratigraphic horizon of the ammonoid fauna is & ROTHGAENGER 1995: 183), illustrated as Acanthoce also proved by the occurrence of Inoeeramus pictus ras (Manlelliceras) mantelli Sow. by DACQUE (1939: bohemicus LEONHARD (TROGER et al. 2009), planktonic pi. 2, figs 13. 14), and Pseudocalycoceras cf. harpax foraminifera (Risen 1983) and the Praeactinoeamax (FÖRSTER et al. 1983: pi. 2, figs 6, 7) are rare. Small plenus Event in the nodular Kalkmergelbank (FÖRSTER Fig. 6: Ammonites from the Pulverturm Member of the Kagerhöh Formation. A: Collignonicerasl woollgarP. (MANTELL), X 0.66, Winzerer Höhen near Regensburg. JME. - B: leweskeras peramplum (MANTELL), X 0.66, Galgenberg in Regensburg. MMG: BaK 2k. SDGG, Heft 65 - Litho- und Biostratigraphie der außeralpinen Kreide von Bayern 119 et al. 1983: WLLMSEN et al. in prep.). Even if the Eibrunn Member. The ammonoid fauna of the middle and Formation spans the Cenomanian-Turonian boundary upper members will be treated separately below. and ranges into the lower Lower Turonian (proved by inoceramid occurrences, see TRÖGER et al. 2009), Pulverturm Member: no uppermost Cenomanian or lowermost Turonian Cotlignoniceras'! woollgarp. (Mantell, 1822) Fig. 6A ammonites have been recorded so far. Lewesiceras peramplum (M antei l. 1822) Figs 6B. 7A Romaniceras (Romaniceras) kallesi? (Zazvorka, 1958) 2.3 Winzerberg Formation - Fig. 7B Lewesiceras peramplum (MANTELL, 1822) -Fig. 5 Remarks: The Pulverturm Member appears to be fairly rich in ammonoids (at least by Danubian Cretaceous Remarks: The spiculitic siltstones and sandstones of Group standards) and it also yields several nautiloids the Winzerberg Formation contain several inoceramid (representatives of the genera Eutrephoceras and De.l- taxa (TRÖGER et al. 2009). However, ammonoids from this unit are very rare. The only record known to us is tocymatoceras). the specimen described and illustrated as Pachydis- Cotlignoniceras? \\ oollgari? (MANTELL, 1822) is repre cus peramplus MANT. by DACQUE (1939: pi. 5, fig. 1). sented by a single, poorly preserved, large specimen According to WRIGHT & KENNEDY (1981), Lewesice (295 mm in diameter) in the collection of the JME ras peramplum occurs in the upper Lower Turonian (Fig. 6) from the Winzerer Höhen near Regensburg. Mammites nodosoides and the Middle Turonian Cotli It is characterized by distant, slightly flexuous ribs on gnoniceras woollgari zones. The record from the Win the outer whorl which are bent forward towards the zerberg Formation is from Reinhausen. in the present- venter, bearing umbilical, ventrolateral and slightly day urban area of Regensburg. Based on superposition clavate siphonal tubercles. As the poor preservation of the Winzerberg Formation by the Middle to lower does not allow a safe determination, the specimen is Upper Turonian Kagerhöh Formation (NIEBUHR et al. kept in open nomenclature. However, it shows some 2009) and inoceramid bivalve evidence (TRÖGER et al. collignoniceratine characters (see KENNEDY et al. 2001) 2009), Le. peramplum from the Winzerberg Formation and Co. woollgari was already mentioned by GI.'MBEI. must be of late Early Turonian age. This is in accord (1868) from the Pulverturm beds and large forms of with the record of two large, non-septate fragments of Co. woollgari with similar ribbing pattern on the outer Mammites from the same locality, questionably attribu whorls were described by FRITSCH (1872: 30, pi. 3, fig. ted to Ma. michelobensis LAUBE & BRUDER by DACQUE 1, pi. 4, fig. 1) from the Bohemian Cretaceous. Thus, (1939: 110). According to WRIGHT & KENNEDY (1981: the specimen is questionably assigned to Co. woollgari 75), Ma. michelobensis LAUBE & BRUDER, 1887 is a (albeit with great reservation). The species characteri younger subjective synonym of Mammites nodosoides zes the eponymous Middle Turonian Cotlignoniceras (SCHLÜTER, 1871). Unfortunately, DACQUE (1939) did woollgari Zone (WRIGHT & KENNEDY 1981). not figure these fragments but from his description it is clear that he was dealing with adult representatives Lewesiceras peramplum appears to be fairly common of the genus Mammites LAUBE & BRUDER. 1887 (see and is represented by several specimens (see Figs 6. LAUBE & BRUDER 1887: 229). 7). According to WRIGHT & KENNEDY (1981), it ranges from the Lower Turonian Mammites nodosoides Zone to the Middle Turonian Cotlignoniceras woollgari IM Kagerhöh Formation Zone. The highest strafigraphically identifiable in situ The Middle to lower Upper Turonian Kagerhöh For occurrence is from below the Ogbourne Hardground at mation of the Regensburg-Kelheim area (see Fig. 1/1) Fognam Quarry, together with Cotlignoniceras wooll- is subdivided into three members, a lower Eisbuckel gari ( WRIGHT & KENNEDY 1981: text-fig. 6). The species Member, a middle Pulverturm Member, and an upper may range into the lower part of the Upper Turonian, Karthaus Member, separated by thin, glauconitic marl for example, the Brantingham Dale specimen cited by horizons ("untere" and "obere Glaukonitmergel- WRIGHT & KENNEDY (1981: 32). However, all higher Bank"; see NIEBUHR et al. 2009). No ammonoids are Upper Turonian Lewesiceras are Le. mantelli WR IGI IT & recorded from the lower Middle Turonian Eisbuckel WRIGHT 1951 or the much rarer species with 11 tubercles. Fig. 7: Ammonoids from the Pulverturm and Karthaus members of the Kagerhöh Formation. A: Lewesiceras peramplum (MANTELL), X 1, Pulverturm Member, Galgenberg in Regensburg. BSP. - B: Romaniceras (Romaniceras) kallesi! (ZAZVUKKA), 13 cm in diameter, untere Glaukonitmergel-Bank, basal Pulverturm Member, Regensburg area. Figured by DA(QIIS (1939: pi. 10, fig. 8), original lost. - C, E, F: Sciponoceras bohemicum7. (FRITSCH), all x l, Karthaus Member. C = Unterdeggenbach, BSP: AS XVII53^, TRUSHEIM collection; E = Regensburg-Karthaus, BSP, DACQUE collection; f = Unterdeggenbach, BSP: AS XVII535, TFWSHEIK collection. - D: undertermined baculitid fragment, x 1, Karthaus Member, Regensburg-Karthaus. BSP, DACQUE collection. 120 SDGG, Heft 65 Litho- und Biostratigfaphie der außeralpinen Kreide von Bayern DACQUE (1939: pl. 10, figs 8,8a) described and illustra Remarks: The "Karthauser Baculitenmergel" (BRUN- ted Acanthoceras deveriamtm D'ORB. from the untere HUBF.R 1917) seems to be fairly rich in ammonoids, Glaukoniimergel-Bank of the Pulverturm Member baculitid and scaphitid heteromorphs being particu of the Regensburg area (re-figured herein in Fig. 7B). larly common. The Lewesiceras figured by DACQUE However, Romaniceras (Ro.) deveriamtm (D'ORBIGNY, (1939, pl. 15, figs 20, 21) as Pachydiseus (Sonnera- 1841) is a lower Upper Turanian index (e.g., BENUTSON tial) peramphis MANT. is re-figured herein (Fig. 8) 1996) and in the U.K. it is found in situ in the lower as Lewesiceras mantelli WRIGHT & WRIGHT, 1951. Upper Turanian (neptuni Zone?) Caburn Sponge Bed DACQUE also illustrated and described Prionotropis below the Caburn Marl / bentonite = TD of northern carolimis D'ORB. from Karthaus (1939: 190, pl. 14, Germany (see MORTIMORE et al. 2001: fig. 1.12). Its fig. 29), associated with Scaphites geinitzi D'ORBIGN Y, occurrence at the base of the Pulverturm Member is in 1850 (DACQUE 1939: pl. 14, fig. 30). Both specimens contradiction to the rich Middle Turanian inoceramid are reproduced in Fig. 9. bivalve fauna from these strata (TRÖGER et al. 2009). Prionotropis carolinus is Collignoniceras caroli- Actually, the DACQUE specimen is fairly poorly pre num (D'ORBIGNY, 1841), a relatively rare species that served, only three rows of originally not too strong, is restricted to the Middle Turanian Collignoniceras slightly clavate tubercles are visible on the venter woollgari Zone (see KENNEDY et al. 1980b). Based on (outer ventrolateral and siphonal ones), whereas the its morphology and straligraphical position (i.e., lower Hanks do not preserve any details of the former orna Upper Turanian), it is more likely that DACQUE'S speci ment. Thus, the specimen may yet represent another men represents a member of the Upper Turanian col- species of the genus Romaniceras SPÄTH, 1923. Its lignoniceratine genus Subprionocyclus SHIMIZU, 1932. rather evolute coiling and the convex trace of the ribs across the venter resulting in a somewhat triangular Sit. hitchinensis (BILLINGHURST, 1927) and Su. branneri arrangement of the three central tubercles are more (ANDERSON, 1902) differ from DACQUE'S specimen by their coarser ornament or denser ribbing, respectively. compatible with Ro. (Ro.) kallesi (ZAZVORKA, 1958), a mid-Middle Turanian species (see KENNEDY et al. Su. neptuni (GEINITZ, 1849) and Su. normalis (ANDER SON, 1958), however, are similar, both being characte 1980a). Ro. (Yubariceras) ornatissimum (STOLICZK-A, rised by clavate inner and outer ventrolateral tubercles, 1865) is slightly more involute and characterized by a crenulated or entire keels, compressed whorl sections coarser ornament. However, based on its poor preser with flat to fastigate venters and relatively strong, more- vation and unavailability for study, DACQUE'S specimen or-less straight prosiradiate ribs which bend forward is kept in open nomenclature. at the inner ventrolateral clavi (which may be missing in certain growth stages). Slight differences arise in whorl-section, degree of involution and ribbing pattern Karthaus Member: (Su. normalis is more compressed and involute, with Lewesiceras mantelli WRIGHT & WRIGHT, 1951 - Fig. 8 steep to vertical, undercut umbilical wall and maximum Siibprionocylus iiepmni'! (GEINITZ, 1849) - Fig. 9A width below mid-flank, and has Hatter ribs; WRIGHT Scaplules sp. - Fig. 9B 1979; KAPLAN 1988). DACQUE'S specimen certainly Sciponoceras bohemicum'l (FRITSCH. 1872) - Fig. 7C, represents either GEINITZ'S or ANDERSONS'S species. It F-I- is in fact very similar to the Subprionocyclus sp. from undetermined baculitid Fig. 7D the Upper Turanian Chalk Rock of the Reed quarry near Royston, England, figured by WRIGHT (1979: pl. 5, fig. 11) which was provisionally included in Su. neptuni by this author (WRIGHT 1979: 320). In support, DACQUE'S specimen is slightly too evolute and has too sharp ribs for S. normalis. Furthermore, there seem to be weak inner ventrolateral tubercles evident from his figure even if DACQUE (1939: 190) described only outer ventrolateral ones. However, the inner ventrolaleral tubercles arc also very weak in the lectotype of Ammonites neptuni (GEINITZ, 1849: 114, pl. 3, fig. 3; see also KAPLAN 1988: 17, pl. 4, fig.4 ) studied by MW in the collection of the Fig. 8: Lewesiceras mantelli\Nmm & WRIGHT, approx. x 1, "Bacu- MMG. Thus, we prefer to include DACQUE'S specimen litenmergel. Karthaus" = Karthaus Member of the Kagerhöh inSubprionocylusneptunp. (GEINIIZ, 1849). Su. neptuni Formation. Figured by DACQUE (1939: pl. 15, figs 20,21), original characterizes the eponymous ammonite biozone of the lost. lower Upper Turanian which fits well w7ith the inoce ramid fauna (cf. TRÖGER et al. 2009). An assignment SDGG, Heft 65 - Litho- und Biostratigraphie der außeralpinen Kreide von Bayern 121 to Sit. normalis or even Prionocyclus germari (REUSS, 2.5 Sandbach Formation 1845) would indicate a horizon in the higher part of the neptuni Zone (above the Hyphantoceras events in Subprionocyclus sp. - Fig. 10A northern Germany and the Czech Republic, e.g., WIESE Scaphites kieslingwaldensis doylei? WRIGHT, 1978 - et al. 2004) or in the uppermost Upper Turonian germari Fig. 10G Peroniceras (Peroniceras) westphalicum (Stromheck. Zone, equivalent to the Mytiloides scupini inoceramid 1859) - Fig. 10B-D, F zone. This is, however, excluded by the inoceramid Placenticeras? sp. - Fig. 10F. evidence (TRÖGER et al. 2009). Baculites sp. - Fig. 10H The figured specimen of Scaphites does not match any Remarks: The upper Upper Turonian-Middlc Conia of the various morphotypes of this species from the cian Sandbach Formation crops out in the southeastern Upper Turonian of Germany and England as figured distributional area of the Danubian Cretaceous Group by WRIGHT (1979) and KAPLAN et al. (1987). In fact, near Passau (see Fig. 1/VI). It consists of a lower Mar- the ribbing of the specimen illustrated in Fig. 9B looks terberg Member (micaceous, sandy-silty marls) and an pathological. In the view of this observation and the upper Buchleiten Member (marls and spiculitic marly fact that the original is lost, we keep this specimen in limestones). The ammonoids from this formation are open nomenclature, without specific determination. illustrated in Fig. 10. Seven hamitids were figured as Baculites baculoi- The Marterberg Member yielded Subprionocyclus sp., des MANT. by DACQUE (1939: pi. 15, figs 9-15) from Scaphites kieslingwaldensis doylei? WRIGHT, 1978 and Karthaus which all seem to represent Sciponoceras Baculites sp. (Fig. 10A, G, H).The poor preservation of bohemicum (FRITSCH, 1872) based on the absence of the small fragment of Subprionocyclus unfortunately tubercles. There are 13 additional fragmentary baculi- prevents a safe specific identification. It represents one tid specimens of DACQUE in the BSP (two of them illus item of the important historical collection of STOCK- trated in Fig. 7D, E) and also the TRUSHEIM collection HEIM (1852) which was transferred to Dresden around (BSP material) yields five baculitids (two of which are 1850, and H.B. GEINITZ determined it as Ammonites illustrated in Fig. 7C, F). However, considering their neptuni GEINITZ. It is in fact similar to illustrations of fragmentary preservation, the specific assignment is Su. neptuni, e.g., the specimen from the Upper Turo uncertain. If tubercles could be proved, also Baculites nian Chalk Rock of Hitch Wood (England) of WRIGHT undulatus D'ORBIGNY, 1850 could be present. (1979: pi. 5, fig.3) . If it is conspecific to GEINITZ spe Lewesiceras mantelli, Sciponoceras bohemicum and cies', it would represent a level very high in the neptuni Scaphites sp. are all relatively long-ranging (Middle Zone or lowermost scupini Zone (KAPLAN & KENNEDY Turonian woollgari Zone to Upper Turonian neptuni 1996). Scaphites kieslingwaldensis doylei? likewise and germari ammonite zones) according to the data in indicates a position high in the Upper Turonian and KAPLAN & KENNEDY (1996) for Westphalia. The record ranges into the Lower Coniacian (KAPLAN & KENNEDY of these three taxa from the Karthaus Member is there 1996). This range fits well with the inoceramid bivalve fore biostratigraphically uninformative. However, the fauna which indicates the upper Upper Turonian Myti occurrence of a Subprionocyclus neptuni? supports loides scupini Zone and the lower part of the Lower a neptuni zonal position which is also supported by Coniacian for the Marterberg Member (TRÖGER et al. the inoceramid fauna of both the Kagerhöh and the 2009). The small baculitid fragment illustrated in Fig. overlying Großberg Formation (TRÖGER et al. 2009). 10H was determined as Baculites incurvatus DUJAR- DIN by GEINITZ. There are some oblique ribs and traces of bullate tubercles, but we refrain from a specific determination due to the poor preservation. GERSTER (1881) mentioned Ammonites tri- carinatus D'ORB. [= Peroniceras (Pero niceras) sublricarinatum (D'ORBIGNY, 1850)] from the Buchleiten Member. The collection material from that member in the MMG (STOCKHEIM collection) yiel Fig. 9: Ammonites from the Karthaus Member of the Kagerhöh Formation. ded several fragmentary specimens of A: Subprionocylus neptuni1. (GEIWN), X 1, "Baculitenmergel. Karthaus". Figured Peroniceras (Pe.). However, we prefer to by DACQUE (1939: pi. 14, fig. 29), original lost. - B: Scaphites sp. (pathological include these specimens in Peroniceras specimen), x 1, ''Baculitenmergel. Karthaus". Figured by DAtguf. (1939: pi. 14, (Pe) westphalicum (STROMBECK, 1859) fig. 30), original lost. based on their generally compressed 122 SLXICI, HELL 65 - LITHO- UND BIOSTRATIGRAPHIC DER AUßERALPINEN KREIDE VON BAYERN Fig. 10: Ammonoids from (he Sandbach formation near Passau. A: Subprionocyclus sp., x 2, Marterberg Member. MMG: BaK 8. - B, C, D, F: Peroniceras (Peroniceras) westphalicum (STROMBCCX), x 1, Buchleiten Member. B, F = MMG: BaK 6; C, D, = MMG: BaK 5. - E: Placenticeras? sp., x 1, Buchleiten Member. MMG: BaK I. - G: Scaphites kieslingwaldvnsis doyleP. WRIGHT, X 2, Marterberg Member. MMG: BaK 9. - H: Baculites sp., x 1, Marterberg Member. MMG: BaK 3. SDGG, Heft 65 - Litho- und Biostratigraphie der außeralpinen Kreide von Bayern 123 whorl-section, low umbilical walls, the predominance (REGCNSBURGER KREIDE, NORDOSTBAYERN). ZITTELIANA, 10: of"relatively straight, slightly proradiate single ribs on 123-141; MÜNCHEN. the outer whorls ending in moderately clavate vent FRITSCH, A. (1972); CEPHALOPODEN DER BÖHMISCHEN KREIDE rolateral tubercles and the more-or-less monocarinate FORMATION. - 52 PP.; PRAGUE (THE AUTHOR). venter (see KLINULR & KENNEDY 1984). Peroniceras GEINITZ, 11.B. (1849-50): DAS QUADERSANDSLEINGEBIRGE ODER (Pe.) subtricarinaium has much more bifurcations and DAS KREIDEGEBIRGE IN DEUTSCHLAND. - 293 PP.; FREIBERG (VON CRAZ & GERLACH). a conspicuously tricarinate venter. Peroniceras (Pe.) westphalicum is a Middle Coniacian species (KIINGER GERSTER. C. (1881): DIE PLANCRBILDUNGEN URN ORTCNBURG BEI PASSAU. - NOV. ACTA KAISERL 1.COP-CAROL. DEUTSCH. & KENNEDY 1984). The fragmentary Placenticeras? AKAD. NATURF., 42 (1): 1-59: MÜNCHEN. sp. (Fig. 10E) from the Buchleiten Member is not con- GCMBEI. C.W. VON (1854): ÜBERSICHT ÜBER DIE GEOGNOSTISCHEN specific to Placenticeras d orbignianum known from VERHÄLTNISSE DER OBERPFALZ. - KORRESP.-BLATT ZOOL.-MINE the Coniacian Jeding Formation of the Franconian area RAL. VCR. REGENSBURG. S IL): 32. 37-43. REGENSBURG. (I.NINER 1935). A specific determination, however, is GLMBEL, C.W. VON (1868A): VERZEICHNIS DER IN DER SAMMLUNG impossible. The ammonoids of the Buchleiten Mem DES ZOOL.-MINERAL. VEREINS IN REGENSBURG VORTINDLICHEN ber are associated with an inoceramid bivalve fauna VERSTEINERUNGEN AUS DEN SCHICHTEN DER PROCÄN- ODER of late Early Coniacian age (TRÖGER et al. 2009). The KREIDEFORMATION AUS DER UMGEGEND VON REGENSBURG. - presence of Peroniceras (Pe.) westphalicum suggests KORRESP.-BL. ZOOL.-MINERAL. VER, REGENSBURG, 22: 51-80: that the member also includes the (lower parts of the) REGENSBURG. Middle Coniacian. GLMBEL, C.W, VON (1868B): ÜEOGNOSTISCHE BESCHREIBUNG DES KOENIGREICHS BAYERN. 2. ABIHEILUNG, ÜEOGNOSTISCHE BE SCHREIBUNG DES OSLBAYERISCHEN GREN/GCBIRGES ODER DES BAYERISCHEN UND OBERPFÄLZER WALDGEBIRGES. - 968 PP.: Acknowledgements GOTHA (JUSTUS PERTHES). HACKER, U. (1971): ZUR FOSSILFÜHRUNG DER REGCNSBURGER We thank M. RÖTUEL and R. WINKLER (Senckenberg OBERKREIDESCHICHTEN. - ACTA ALBERTINA RATIBONENSIA. 31: Naturhistorische Sammlungen Dresden, Museum für 135-150; REGENSBURG. Mineralogie und Geologie), M. EUERT, M. KÖLBL-EBERT KAPLAN, U. (1988): DIE AMMONILCN-SUBFAMILIE COLLIGNO- and Ci. VIOHL (Juramiiseum Eichstätt) as well as A. NICERATINAE WRIGHT & WRIÜHT 1951 AUS DEM TURON (OBER- NU TZ EL. S. SCHNEIDER und W. WERNER (Bayerische KREIDE) VON WESTFALEN UND NIEDERSACHSEN (N W-DCUTSCH- Staatssammlung für Paläontologie. Munich) for the LAND). GEOL. PALÄONT. WESTFALEN, 12: 5-45; MÜNSTER. opportunity to study ammonoid material from their KAPLAN, U. & KENNEDY. W.J. (1996V UPPER TURONIAN AND collections. CONIACIAN AMMONITE STRATIGRAPHY OF WESTPHALIA, NW- GERMANY. - ACTA GCOL. POLON.. 46: 305-352; WARSZAWA. KAPLAN, U„ KENNEDY, W.J. & WHICH I, C.W. (1987): TURONIAN AND CONIACIAN SCAPHITIDAE FROM ENGLAND AND NORTHWES References TERN GERMANY. GEOL. JB.. A 103: 5-39: HANNOVER. BI V.ISOV P. (1988): OPEN NOMENCLATURE. - PALAEONTOLOGY, KENNEDY, W.J., WRIGHT. C.W, & HANCOCK, J M. (1980A): THE 31 (1): 223-227: LONDON. EUROPEAN SPECIES OF THE CREIACEOUS AMMONITE Rumti- HI V ISOS. P. (1496): THE TURONIAN STAGE AND SUBSTAGE BOUN nkeras WITH A REVISION OF THE GENUS. PALAEONTOLOGY. 23: DARIES. BULL, DE I' INSL. ROYAL SEI. NAT. BELGIQUE (SEI. DE 325-362; LONDON. LA TERRC): 66: 69-74; BRUSSELS. KENNEDY, W.J., WRIGHT, C.W. & HANCOCK. J.M. (1980B): COL- BRI NIIUDHK, A. (1917): DIE GEOLOGISCHEN VERHÄLTNISSE VON LIGNONICERATID AMMONITES FROM IHC MID-TURONIAN OF ENG REGENSBURG UND UMGEBUNG. NATURWISS. VER. REGENS LAND AND NORTHERN FRANCE. PALAEONTOLOGY. 23: 557-603. BURG: 107 PP.; REGENSBURG. LONDON. CIIRLI.INEUN. P. (2008): LAND-KÜSTE MEER: STRATIGRAPHIC, KENNEDY, W.J., COUBAN, W.A. & LANDMAN. N.I I. (2001): A PALÄONTOLOGIE UND SCDIMENLOLOGIE DER DANUBISCHEN REVISION OF THE TURONIAN MEMBERS OF THE AMMONITE SUBFA KREIDE-GRUPPE IN DER ÖSTLICHEN BODENWÖHRER SENKE MILY COLLIGNONICERATINAE FROM THE UNITED STATES WESTERN (OBERPFALZ, BAYERN). DIPLOMA THESIS, JULIUS-MAXIMILI- INTERIOR AND GULF COAST. - BULL. AMERICAN MUS. NAT. HIST.. ANS-UNIVEISITÄT: 104 PP.; WÜRZBURG. [UNPUBLISHED] 267: 148 PP.; NEW YORK. (TIKISII SSIN. W,K. (1990): UPPER CRETACEOUS BELEMNITE STRA KLINOFR, H.C. & KENNEDY, W.J. (1984): CREIACEOUS FAUNAS TIGRAPHY OF EUROPE. CRET. RESEARCH, 11: 37-386; LONDON. FROM ZULULAND AND NATAL, SOUTH AFRICA: THE AMMONITE OIMHF.MOREI., R., CHRIÜIEXSEN, W.K., NAIDIN, D.P. & SPAETH, SUBFAMILY PERONIEERATINAE HYATT, 1900. ANNALS SOUTH C. (1981): [.ES BCLCMNITES. - CRCT. RESEARCH, 2: 383-286; AFRICAN MUS., 92: 113-294; CAPE TOWN. LONDON. LAUBE, G.C. & BRUDER. G. (1887): AMMONITEN DER BÖHMI D\( VI I. E. (1939): DIE FAUNA DER REGENSBURG-KEHLHEIMER SCHEN KREIDE. - PALÄONTOGRAPHICA. 33: 217-239, STUTTGART. OBERKRCIDC. ABH. BAYER. AKAD. WISS., N.F., 45: 218 PP.; LEUNER, L. (1935): ÜBER DAS TURON AUF DEM FRÄNKISCHEN JURA. MÜNCHEN (C.II. BECK). STUDIEN ÜBER DIE FRÄNKISCHE ALBÜBERDECKENDC KREIDE. FORSILR, R., MEYER. R. & RISEN, H. (1983): AMMUNITEN UND VIII. ZW. MIN. GCOL. PAL., ABT. B. 11: 423-438; STUTT PLANKTONISCHE FORAMINIFEREN AUS DEN EIBRUNNER MERGELN GART. 124 SDGG, Heft 65 - Litho- und Biostratigfaphie der außeralpinen Kreide von Bayern LEHMER. L. (1937a): Fauna und Flora der fränkischen alb- und Böhmens. CorTesp.-Bl. zool.-mineral. Ver. Regens überdeckenden Kreide. I. Die Lamellibranchiaten (ohne burg, 6: 83-93; Regensburg. Inoceramen), herausgegeben von R. DEHM. - Palaconto- TILLMANN, H. (1954): Die Kreide in Nordostbayern. - In. graphica, A 85: 115-228; München. Erläuterungen zur Geologischen Karte von Bayern LF.HNER, L, (1937b): Fauna und Flora der fränkischen alb- 1:500000: 85-94; München (BGL). überdeekenden Kreide. II. Fauna 2. Teil und Flora, her TRÖGER, K.-A., NIEBUHR, B. & WILMSES. M. (2009): Inoce ausgegeben von R. DfcHM. - Palaeontographica, A 87: ramen aus dem Cenomanium bis Coniacium der Danu- 158-230: München. bischen Kreide-Gruppe (Bayern, Süd-Deutschland). - MFYFR, R.K.F. (2000): Außeralpine Kreide in Süddeutschland SDGG, 65: 59-110; Hannover. (Regensburger Kreide). - In: Stratigraphische Kommis TRUSHEIM, F. (1935): Die geologische Geschichte Südost sion Deutschland (Ed.), Stratigraphie von Deutschland deutschlands während der Unterkreide und des Ccno- III. Die Kreide der Bundesrepublik Deutschland. Courier mans. - N. Jb. Min., GeoL, Paläont.. Beil.. 75: 1-109; Forschungsinst. Senckenberg. 226: 141-147; Frankfurt. Stuttgart. MoFTivioKh. R.N., WOOD, C.J. & GALLOIS. R.W. (2001): British WIESE, F., CECH, S„ EKRT, B„ KOST'AK, M„ MAZLCH. M. & Upper Cretaceous stratigraphy. - Geol. Conserv. Rcv. VOIGT, S. (2004): The Upper Turanian of the Bohemian Ser.. 23: 558 pp.: Peterborough. Cretaceous Basin (Czech Republic) exemplified by the Nir.BUHR, B., PÜRNER, T. & WILMSF.N, M. (2009): Lithostrati- Upohlavy working quarry: integrated stratigraphy and graphie der außeralpinen Kreide Bayerns. - SDGG, 65: palaeoceanography of a gaieway to the Tethys. - Cret. 7-58: Hannover. Research, 25, 329-352; Amsterdam. OSCHMANN. F. (1958): Geologische Karte von Bayern WILMSEN, M. & NIEBUHR, B. (2010): On the age of the Upper Cretaceous transgression between Regensburg and Neu- 1:25000, Erläuterungen zum Blatt Nr. 7038 Bad Abbach. burg an der Donau (Bavaria, southern Germany). - N. Jb. 184 pp.; München (BGL). Geol. Paläont.; Stuttgart. [In press] RISCH, H. (1983): Zur Mikrobiostratigraphie der Regensbur WRIGHT, C.J. & KENNEDY, W.J. (1981): The Ammonoidea of ger Kreide. Zitteliana, 10: 143-153: München. the Plenus Marls and the Middle Chalk. -Palaeont. Soc. RÜPER, M. & NELMEIER. F. (1995): Neue Fossilfundc aus der Re Monogr., 560 (134): 148 pp.; London. gensburger Oberkreide Teil 2: Der Großberger Sandstein WRIGHT, C.W. (1979): The ammonites of the English Chalk von Eggmühl (Oberturon). - Fossilien, 6/1995: 367-372; Rock (Upper Turonian). - Bull. British Mus. (Natural Korb. History), Geol. Ser.. 31: 281-332; London. RÖPBR, M. & ROTHUAF.SCIER, M. (1995): Neue Fossilfunde aus WRIGHT, C.W., CALLOMON. JH., HOWARTH. M.K. (1996): der Regensburger Oberkreide Teil 1: Eibrunner Mergel. - Cretaceous Ammonoidea. - In: KAFSLFR, R.L. (Ed.), Fossilien, 3/1995: 180-184; Korb. Treatise on Invertebrate Paleontology. Part L: Mollusca, STOCKHEIM, F. VON (1852): Vergleichende Übersicht der Krci- revised, Volume 4. 362 pp.: Geol. Soc. America, Inc., degebildc Regensburg* mit jenen derGegend von Passau Boulder and The University of Kansas Press, Lawrence.