Delbert Hunter Arboretum and Botanical Garden
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Outline of Angiosperm Phylogeny
Outline of angiosperm phylogeny: orders, families, and representative genera with emphasis on Oregon native plants Priscilla Spears December 2013 The following listing gives an introduction to the phylogenetic classification of the flowering plants that has emerged in recent decades, and which is based on nucleic acid sequences as well as morphological and developmental data. This listing emphasizes temperate families of the Northern Hemisphere and is meant as an overview with examples of Oregon native plants. It includes many exotic genera that are grown in Oregon as ornamentals plus other plants of interest worldwide. The genera that are Oregon natives are printed in a blue font. Genera that are exotics are shown in black, however genera in blue may also contain non-native species. Names separated by a slash are alternatives or else the nomenclature is in flux. When several genera have the same common name, the names are separated by commas. The order of the family names is from the linear listing of families in the APG III report. For further information, see the references on the last page. Basal Angiosperms (ANITA grade) Amborellales Amborellaceae, sole family, the earliest branch of flowering plants, a shrub native to New Caledonia – Amborella Nymphaeales Hydatellaceae – aquatics from Australasia, previously classified as a grass Cabombaceae (water shield – Brasenia, fanwort – Cabomba) Nymphaeaceae (water lilies – Nymphaea; pond lilies – Nuphar) Austrobaileyales Schisandraceae (wild sarsaparilla, star vine – Schisandra; Japanese -
Plant Introductions
Plant Introductions Promoting the garden use of native plants is a core part of Mt. Cuba Center’s mission, and since 1988, the Plant Introduction Program has brought many outstanding cultivars to market. Selected for strong ornamental appeal and broad regional adaptability, many of these introductions have risen to become some of the most popular cultivars of native plants available. Plant evaluation and introduction continues today, with several bright prospects for additional introductions in the future. Plant Introductions Actaea pachypoda ‘Misty Blue’ Misty Blue white baneberry Misty Blue white baneberry is an herbaceous perennial, selected for its unique and highly attractive soft, bluish-green foliage. It was discovered in a planting of typical green-leaved plants of unknown origin growing at Mt. Cuba Center. In spring 1" to 2" tall bottlebrush-like clusters of white flowers are borne on stems above the foliage. By September, large, white fruit with dark purple to black spots (“doll’s eyes”) mature on reddish pedicels. It is a carefree, long-lived, 24" to 36" tall plant, growing best in partial to filtered shade in evenly moist, well-drained soils with a pH from slightly acidic to neutral. Introduced 2009 Ageratina altissima ‘Chocolate’ (formerly Eupatorium rugosum) Chocolate white snakeroot Chocolate white snakeroot is an herbaceous perennial selected by Dr. Richard W. Lighty and descended from a plant found at Winterthur Gardens. It had the darkest burgundy foliage of many seedlings grown over a ten year period. ‘Chocolate’ grows up to 3' tall and has white inflorescences along with dark burgundy leaves which color best in full sun. -
Northwest Plant Names and Symbols for Ecosystem Inventory and Analysis Fourth Edition
USDA Forest Service General Technical Report PNW-46 1976 NORTHWEST PLANT NAMES AND SYMBOLS FOR ECOSYSTEM INVENTORY AND ANALYSIS FOURTH EDITION PACIFIC NORTHWEST FOREST AND RANGE EXPERIMENT STATION U.S. DEPARTMENT OF AGRICULTURE FOREST SERVICE PORTLAND, OREGON This file was created by scanning the printed publication. Text errors identified by the software have been corrected; however, some errors may remain. CONTENTS Page . INTRODUCTION TO FOURTH EDITION ....... 1 Features and Additions. ......... 1 Inquiries ................ 2 History of Plant Code Development .... 3 MASTER LIST OF SPECIES AND SYMBOLS ..... 5 Grasses.. ............... 7 Grasslike Plants. ............ 29 Forbs.. ................ 43 Shrubs. .................203 Trees. .................225 ABSTRACT LIST OF SYNONYMS ..............233 This paper is basicafly'an alpha code and name 1 isting of forest and rangeland grasses, sedges, LIST OF SOIL SURFACE ITEMS .........261 rushes, forbs, shrubs, and trees of Oregon, Wash- ington, and Idaho. The code expedites recording of vegetation inventory data and is especially useful to those processing their data by contem- porary computer systems. Editorial and secretarial personnel will find the name and authorship lists i ' to be handy desk references. KEYWORDS: Plant nomenclature, vegetation survey, I Oregon, Washington, Idaho. G. A. GARRISON and J. M. SKOVLIN are Assistant Director and Project Leader, respectively, of Paci fic Northwest Forest and Range Experiment Station; C. E. POULTON is Director, Range and Resource Ecology Applications of Earth Sate1 1 ite Corporation; and A. H. WINWARD is Professor of Range Management at Oregon State University . and a fifth letter also appears in those instances where a varietal name is appended to the genus and INTRODUCTION species. (3) Some genera symbols consist of four letters or less, e.g., ACER, AIM, GEUM, IRIS, POA, TO FOURTH EDITION RHUS, ROSA. -
Trees, Shrubs, and Perennials That Intrigue Me (Gymnosperms First
Big-picture, evolutionary view of trees and shrubs (and a few of my favorite herbaceous perennials), ver. 2007-11-04 Descriptions of the trees and shrubs taken (stolen!!!) from online sources, from my own observations in and around Greenwood Lake, NY, and from these books: • Dirr’s Hardy Trees and Shrubs, Michael A. Dirr, Timber Press, © 1997 • Trees of North America (Golden field guide), C. Frank Brockman, St. Martin’s Press, © 2001 • Smithsonian Handbooks, Trees, Allen J. Coombes, Dorling Kindersley, © 2002 • Native Trees for North American Landscapes, Guy Sternberg with Jim Wilson, Timber Press, © 2004 • Complete Trees, Shrubs, and Hedges, Jacqueline Hériteau, © 2006 They are generally listed from most ancient to most recently evolved. (I’m not sure if this is true for the rosids and asterids, starting on page 30. I just listed them in the same order as Angiosperm Phylogeny Group II.) This document started out as my personal landscaping plan and morphed into something almost unwieldy and phantasmagorical. Key to symbols and colored text: Checkboxes indicate species and/or cultivars that I want. Checkmarks indicate those that I have (or that one of my neighbors has). Text in blue indicates shrub or hedge. (Unfinished task – there is no text in blue other than this text right here.) Text in red indicates that the species or cultivar is undesirable: • Out of range climatically (either wrong zone, or won’t do well because of differences in moisture or seasons, even though it is in the “right” zone). • Will grow too tall or wide and simply won’t fit well on my property. -
Dimensions of Biodiversity
Dimensions of Biodiversity NATIONAL SCIENCE FOUNDATION CO-FUNDED BY 2010–2015 PROJECTS Introduction 4 Project Abstracts 2015 8 Project Updates 2014 30 Project Updates 2013 42 Project Updates 2012 56 Project Updates 2011 72 Project Updates 2010 88 FRONT COVER IMAGES A B f g h i k j C l m o n q p r D E IMAGE CREDIT THIS PAGE FRONT COVER a MBARI & d Steven Haddock f Steven Haddock k Steven Haddock o Carolyn Wessinger Peter Girguis e Carolyn g Erin Tripp l Lauren Schiebelhut p Steven Litaker b James Lendemer Wessinger h Marty Condon m Lawrence Smart q Sahand Pirbadian & c Matthew L. Lewis i Marty Condon n Verity Salmon Moh El-Naggar j Niklaus Grünwald r Marty Condon FIELD SITES Argentina France Singapore Australia French Guiana South Africa Bahamas French Polynesia Suriname Belize Germany Spain Bermuda Iceland Sweden Bolivia Japan Switzerland Brazil Madagascar Tahiti Canada Malaysia Taiwan China Mexico Thailand Colombia Norway Trinidad Costa Rica Palau United States Czech Republic Panama United Kingdom Dominican Peru Venezuela Republic Philippines Labrador Sea Ecuador Poland North Atlantic Finland Puerto Rico Ocean Russia North Pacific Ocean Saudi Arabia COLLABORATORS Argentina Finland Palau Australia France Panama Brazil Germany Peru Canada Guam Russia INTERNATIONAL PARTNERS Chile India South Africa China Brazil China Indonesia Sri Lanka (NSFC) (FAPESP) Colombia Japan Sweden Costa Rica Kenya United Denmark Malaysia Kingdom Ecuador Mexico ACKNOWLEDGMENTS Many NSF staff members, too numerous to We thank Mina Ta and Matthew Pepper for mention individually, assisted in the development their graphic design contribution to the abstract and implementation of the Dimensions of booklet. -
Native Plants for Wildlife Habitat and Conservation Landscaping Chesapeake Bay Watershed Acknowledgments
U.S. Fish & Wildlife Service Native Plants for Wildlife Habitat and Conservation Landscaping Chesapeake Bay Watershed Acknowledgments Contributors: Printing was made possible through the generous funding from Adkins Arboretum; Baltimore County Department of Environmental Protection and Resource Management; Chesapeake Bay Trust; Irvine Natural Science Center; Maryland Native Plant Society; National Fish and Wildlife Foundation; The Nature Conservancy, Maryland-DC Chapter; U.S. Department of Agriculture, Natural Resource Conservation Service, Cape May Plant Materials Center; and U.S. Fish and Wildlife Service, Chesapeake Bay Field Office. Reviewers: species included in this guide were reviewed by the following authorities regarding native range, appropriateness for use in individual states, and availability in the nursery trade: Rodney Bartgis, The Nature Conservancy, West Virginia. Ashton Berdine, The Nature Conservancy, West Virginia. Chris Firestone, Bureau of Forestry, Pennsylvania Department of Conservation and Natural Resources. Chris Frye, State Botanist, Wildlife and Heritage Service, Maryland Department of Natural Resources. Mike Hollins, Sylva Native Nursery & Seed Co. William A. McAvoy, Delaware Natural Heritage Program, Delaware Department of Natural Resources and Environmental Control. Mary Pat Rowan, Landscape Architect, Maryland Native Plant Society. Rod Simmons, Maryland Native Plant Society. Alison Sterling, Wildlife Resources Section, West Virginia Department of Natural Resources. Troy Weldy, Associate Botanist, New York Natural Heritage Program, New York State Department of Environmental Conservation. Graphic Design and Layout: Laurie Hewitt, U.S. Fish and Wildlife Service, Chesapeake Bay Field Office. Special thanks to: Volunteer Carole Jelich; Christopher F. Miller, Regional Plant Materials Specialist, Natural Resource Conservation Service; and R. Harrison Weigand, Maryland Department of Natural Resources, Maryland Wildlife and Heritage Division for assistance throughout this project. -
Errata Flora of Virginia, First Edition, First Printing
Errata Flora of Virginia, first edition, first printing Page Correction viii 8th paragraph down; 2nd sentence should read “They include Sally Anderson, Woody Bousquet, Barbara Dudley Davis, Dwayne Estes, Irene Frentz, Jolie Harrison, Jan Hodges, Ryan Huish, Ryan Klopf, Wesley Knapp, Sara Koropchak, Ron Lance, Richard LeBlond, Rachel Ludwig, Carol Ann McCormick, Zack Mur- rell, Robert Peet, Derick Poindexter, Milo Pyne, Megan Rollins, Rosalind Rowe, Bruce Sorrie, Mo Stevens, Catharine Tucker, Allison Weakley, and Andrea Weeks.” ix Between 2nd and 3rd paragraphs, insert this paragraph: “Several botanists developed or co-developed regional taxonomic treatments and keys for Weakley (2012), which then formed the basis for the taxonomy and keys of generic treatments in the Flora of Virginia: Bruce Sorrie (Sisyrinchium, Juncus, Eleocharis, Viola, Lechea, Sabatia, Amsonia, Scutellaria, Lobelia, Coreopsis, Pityopsis, Viburnum), Richard LeBlond (Rhynchospora, Scleria, Coleataenia, Dichanthelium, Panicum, Paspalum, Rhexia), Milo Pyne (Physalis, Solanum), Derick Poindexter (Abies, Taenidia), Wesley Knapp (Juncus), Robert Peet (Carya), Zack Murrell (Cornus), and Dwayne Estes (Penstemon).” xi Entry under J. Christohper Ludwig; 4th line down should read “vegetation communities” (rather than “vege- tative communities”) xv Eliminate entry here for Paul and Eunice Hyer xv Add entry for NatureServe and entry for Rivanna Garden Club xvi Under Honorary and Memorial Gifts add two entries: In memory of Dorothy M. Edwards, from Paul and Eunice Hyer In memory -
Azalea Or Rhododendron (Ericaceae)
Rhododendron - Azalea or Rhododendron (Ericaceae) --------------------------------------------------------------------------------------------- Azaleas and Rhododendrons are very popular small- on short petioles, and subject to chlorosis (yellowing to medium-sized shrubs that flower prolifically in due to a deficiency of iron and other micronutrients) spring, but are slow growing and require acidic, in alkaline soils organic, moist, and well-drained soils in shady -for most Azaleas, leaves are either deciduous or semi- situations for establishment and optimum evergreen, about 1-2" long, generally with some performance. pubescence, and have either a subtle mixed autumn color, or virtually no change in color before FEATURES abscission Form -for most Rhododendrons, leaves are generally -small- to medium-sized ornamental shrubs lustrous evergreen, from 2-6" long, and with subtle -Azaleas generally mature from 2-4' in height and scales and/or with dots on their lower surfaces width, and Rhododendrons generally mature from 4- Flowers 10' in height and width -for both Azaleas and -upright clump growth habit in youth, becoming Rhododendrons, shades rounded to spreading with age of white, salmon, -slow growth rate pink, red, crimson, Culture violet, lavender, or -partial shade to full shade purple are the most -performs best in partial shade in moist, well-drained, common floral colors, acidic soils, with the keys to successful growth being but yellows and the placement of the shrub in a well-drained soil that oranges also exist; is moderately -
1 Updates Required to Plant Systematics: A
Updates Required to Plant Systematics: A Phylogenetic Approach, Third Edition, as a Result of Recent Publications (Updated June 13, 2014) As necessitated by recent publications, updates to the Third Edition of our textbook will be provided in this document. It is hoped that this list will facilitate the efficient incorporation new systematic information into systematic courses in which our textbook is used. Plant systematics is a dynamic field, and new information on phylogenetic relationships is constantly being published. Thus, it is not surprising that even introductory texts require constant modification in order to stay current. The updates are organized by chapter and page number. Some require only minor changes, as indicated below, while others will require more extensive modifications of the wording in the text or figures, and in such cases we have presented here only a summary of the major points. The eventual fourth edition will, of course, contain many organizational changes not treated below. Page iv: Meriania hernandii Meriania hernandoi Chapter 1. Page 12, in Literature Cited, replace “Stuessy, T. F. 1990” with “Stuessy, T. F. 2009,” which is the second edition of this book. Stuessy, T. F. 2009. Plant taxonomy: The systematic evaluation of comparative data. 2nd ed. Columbia University Press, New York. Chapter 2. Page 37, column 1, line 5: Stuessy 1983, 1990;… Stuessy 1983, 2009; … And in Literature Cited, replace “Stuessy 1990” with: Stuessy, T. F. 2009. Plant taxonomy: The systematic evaluation of comparative data. 2nd ed. Columbia University Press, New York. Chapter 4. Page 58, column 1, line 5: and Dilcher 1974). …, Dilcher 1974, and Ellis et al. -
Ericaceae A. L. De Jussieu (Heath Family) Trees, Shrubs, Lianas
Ericaceae A. L. de Jussieu (Heath Family) Trees, shrubs, lianas, sometimes epiphytic, occasionally Floral formula: mycoparasitic herbs lacking chlorophyll, strongly associ- ated with mycorrhizal fungi. Hairs simple, usually mul- capsule, berry, drupe ticellular and unicellular, sometimes dendritic, gland- Distribution and ecology: Cosmopolitan, but especially headed, or peltate scales, but not stellate. Leaves alternate, common in tropical montane habitats, southern Africa, sometimes opposite or whorled, simple, entire to serrate, sometimes revolute, with pinnate, ± parallel, or palmate eastern North America, and eastern Asia; usually light- loving shrubs of acid soils. venation, blade reduced in mycoparasites; stipules lack- ing. Inflorescences various, flowers usually bisexual, rarely unisexual (then plants usually dioecious), radial to Genera/species: 130/2700. Major genera: Rhododendron slightly bilateral, usually ± pendulous. Sepals usually 4 (800 spp.), Erica (600), Vaccinium (400), Gaultheria (150), or 5, distinct to slightly connate. Petals usually 4 or 5 and Leucopogon (140), Cavendishia (100), and Arctostaphylos connate, often cylindrical to urn-shaped, with small to (50). Noteworthy genera (in addition to most of the above) in the continental United States and/or Canada large, imbricate to valvate lobes, but sometimes ± bell-shaped are Andromeda, Arbutus, Bejaria, Ceratiola, Chamaedaphne, or funnel-like, occasionally distinct (a reversal). Perianth Chimaphila, Corema, Empetrum, Gaylussacia, Kalmia, Leu- reduced to 2 or 3 sepals and petals, or 3 or 4 tepals in a cothoe, Lyonia, Monotropa, Monotropsis, Oxydendrum, few genera that are wind-pollinated. Stamens 8-10, but Pieris, Pterospora, and Pyrola. reduced to 2 or 3 in wind-pollinated species; filaments free or adnate to corolla, sometimes connate, sometimes Economic plants and products: The edible fruits of Vac- with paired projections (spurs) near or at junction with cinium (blueberries, cranberries) are economically impor- anther; anthers becoming inverted, 2- or 1-locular, usually tant. -
Dimensions of Biodiversity: 2010-2014 Projects (Nsf15030
Dimensions of Biodiversity 2010–2014 NATIONAL SCIENCE FOUNDATION PROJECTS CO-FUNDED BY Introduction 4 Project Abstracts 2014 6 Project Updates 2013 32 Project Updates 2012 46 Project Updates 2011 60 Project Updates 2010 76 a FRONT COVER IMAGES f g h b c i k l j m n o p q r d e IMAGE CREDIT THIS PAGE FRONT COVER a Jon G. Sanders d Karen E. Sears f Anothny R. Ives k Karen E. Sears o Michael N. Dawson b E.M. Rivkina e Jenny Xiang g Piotr Łukasik l Ivan Prates p Jon G. Sanders c Klaus Nüsslein h Walter S. Judd m John Wertz & q Ryan McMinds & i Robert Brucker & Alicia Withrow Jerome Payet Seth Bordenstein n Fabian A. r Richard Lankau j Olle Pellmyr Michelangeli FIELD SITES Argentina France Singapore Australia French Guiana South Africa Bahamas French Polynesia Spain Belize Germany Sweden Bermuda Iceland Switzerland Bolivia Japan Tahiti Brazil Madagascar Taiwan Canada Malaysia Thailand China Mexico Trinidad Colombia Norway United States Costa Rica Palau United Kingdom Czech Republic Panama Venezuela Dominican Peru Labrador Sea Republic Philippines North Atlantic Ecuador Poland Ocean Finland Puerto Rico North Pacific Ocean Russia Saudi Arabia COLLABORATORS Argentina Finland Palau Australia France Panama Brazil Germany Peru Canada Guam Russia INTERNATIONAL PARTNERS Chile India South Africa China Brazil China Indonesia Sri Lanka (NSFC) (FAPESP) Colombia Japan Sweden Costa Rica Kenya United Ecuador Malaysia Kingdom Mexico ACKNOWLEDGMENTS Many NSF staff members, too numerous to We thank Mina Ta, Design Specialist, for her mention individually, assisted in the development important contributions to the abstract booklet. -
The Ericaceae of California
THE ERICACEAE OF CALIFORNIA. THE NOW DIVERSE HEATHER FAMILY The Ericaceae is a worldwide family of great importance for its many beautiful ornamentals. • In addition, the genus Vaccinium (and a few others) provide delicious, edible berries including huckleberries, blueberries, and cranberries • The family typically favors moist, shaded, humusy environments with acid soils, but there are several important exceptions • The family is particularly diverse in the Himalayas and other parts of Eastern Asia, in the Cape Province of South Africa, and in California • Two genera stand out for living in hot, dry situations: the genus Arctostaphylos (manzanita) centered in California, and the genus Erica (heather) centered in South Africa At one time, most of the Ericaceae was considered to embrace woody shrubs and trees, many with simple, evergreen leaves • Currently some related groups, including the southern heaths in the former Epacridaceae, and the semi- to fully fungus parasitic, herbaceous plants once placed in the Pyrolaceae, are now lumped into the Ericaceae • The family features urn- to bell-shaped flowers, with some also being saucer shaped or trumpet shaped as in the case of the genus Rhododendron • Typical flower traits include (generally) 5 sepals, 5 petals joined together at the base, 5 or 10 stamens that are not attached to the petals, and a compound pistil of (mostly) 5 carpels • The fruits are either fleshy berries or dry capsules, and… • The stamens’ anthers open by pores or holes at the top Because the Ericaceae has over 2,000