Sieve Plates and Habitat Adaptation in the Foraminifer Planulina Ornatal

Total Page:16

File Type:pdf, Size:1020Kb

Sieve Plates and Habitat Adaptation in the Foraminifer Planulina Ornatal Sieve Plates and Habitat Adaptation in the Foraminifer Planulina ornatal Johanna M. Resig2 and Craig R. Glenn 2 Abstract: Planulina ornata (d'Orbigny), a coarsely perforate species of forami­ nifera having a low trochospiral test, was recovered attached to phosphatic hardgrounds from the lower oxygen-minimum zone off Peru. Above the base of individual pores are calcified, perforate sieve plates, the largest so far described. Structure of the pores suggests a possible association with mitochondria and respiratory function. These large pores may facilitate extraction of the severely limited amount of oxygen from the ambient bottom waters at that locale. IN THE COURSE of investigating foraminifera Jahn (1953) illustrated and described the mi­ encrusting phosphoritic hardgrounds from croporous pore plates and termed them sieve the lower part of the oxygen-minimum zone plates. In some of the studied species, the of the Peru continental slope (Resig and pore plates occur singly at the base of each Glenn 1997), Planulina ornata (d'Orbigny), a pore (Be et al. 1980), but other species have species in which the calcified pore plates and multiple plates, one with each growth lamina their perforations are unusually large, was (LeCalvez 1947, Jahn 1953, Sliter 1974). recovered. Documentation of this occurrence Some pore-sieve plate and micropore diame­ is presented here to supplement the small ters that may be typical offinely and coarsely body of literature on pore microstructure and perforate benthic species and planktonic spe­ function in foraminifera and to lend support cies are as follows: I-j..lm sieve plate, O.I-j..lm to the proposed association of these struc­ micropores in a nonionid (Jahn 1953); 2- to tures with respiratory organelles (Berthold 6-j..lm sieve plate, 0.2- to 0.3-j..lm micropores 1976, Leutenegger and Hansen 1979). in Lamellodiscorbis aguayoi (Arnold 1954b); and An increasing number of perforate fora­ 3.3-j..lm sieve plate, 0.1- to 0.6-j..lm micropores minifera representing various benthic and in Globigerinoides sacculifer (Be et al. 1980). planktonic genera are shown to have their Planulina ornata, in contrast, has 6- to lO-j..lm pores closed offfrom the chambers by organic diameter sieve plates and 0.5- to 1.0-j..lm membranes or plates (Table 1), a condition diameter micropores. These well-defined that may exist for most perforate foraminifera structures are presumably functional within (Boltovskoy and Wright 1976). The plates the species' habitat. are reported to be nonporous in some fora­ minifera such as Ammonia (Banner and Wil­ HABITAT OF Planulina ornata liams 1973), but the majority of those studied are microporous and either calcified or not. Planulina ornata was originally described by d'Orbigny (1839) as Truncatulina ornata from off the Port ofValparaiso, Chile, where it was 1 Partial support of this research was provided by a rare. Its geographic range continues north­ Nation-ai-Science Foundation grant (OCE-9201305) to ward to Alaska (Buzas and Culver 1990). c.R.G. This is University of Hawai'i School of Ocean and Earth Science and Technology Contribution No. Specimens occur free in the sediment and 6005. Manuscript accepted 1 March 2002. are reported to be most frequent on the outer 2 Department of Geology and Geophysics; University continental shelf (Bandy and Arnal 1957 of Hawai'i at Manoa, Honolulu, Hawai'i 96822. [Central America], Bandy 1961 [Gulf of California]) or on the upper continental slope Pacific Science (2003), vol. 57, no. 1:103-110 between 160 and 260 m (Boltovskoy and © 2003 by University ofHawai'i Press Theyer 1970 [Chile]). Their bathymetric All rights reserved range generally extends deeper along the 103 104 PACIFIC SCIENCE· January 2003 TABLE 1 western American margin, to about 1860 m Reported Occurrence of Pore Plates in Foraminifera off Chile (Ingle et al. 1980) or 1200 m off Peru (Resig 1981). Dchio (1960) found Superfarnily/Species Reference protoplasm-bearing tests identified through rose bengal stain from 38 to 946 m and empty Spirillinacea tests from 10 to 1260 m off San Diego, Cali­ Patellina corrugata Berthold (1976) fornia. Specimens are also present in Qua­ Nodosariacea Robulus midwayensis* Hay et aI. (1963) ternary sediment of four cores from the Bolivinacea Peruvian margin taken in water depths be­ Bolivina argentea Leutenegger and tween 307 and 447 m (Resig 1990). In none Hansen (1979) ofthese reports did the species compose more Bolivina cf. excavata Leutenegger and Hansen (1979) than 4% of the benthic foraminiferal as­ Bolivina sp. Sliter (1974) semblages. "Loxostomum" pseudobeyrichi Leutenegger and The extent to which sedimentary speci­ Hansen (1979) mens reflect the life habitat of P. ornata is not Cassidulinacea known. Specimens found free in the sediment Cassidulinoides cornuta Leutenegger and Hansen (1979) may have lived as epi- or infauna; they may Buliminacea have experienced pre- or postmortem dis­ Buliminella tenuata Leutenegger and lodgement or transport. The adherent speci­ Hansen (1979) mens reported here are the only ones so far to Globobulimina pacifica Leutenegger and Hansen (1979) have been collected with certainty in their life Discorbacea positions. Discorbis erecta LeCalvez (1947) Several specimens of P. ornata were recov­ Discorinopsis aguayoi** Arnold (1954a,b) ered from off the coast of Peru at' 11° 59.89' Rosalina floridana Angell (1967) S, 77° 50.91' W in a water depth of 517 m, Planorbulinacea Cibicides cassivellauni Wood and during operations of the Johnson Sea Link Haynes (1957) I submersible (Figure 1). The specimens ad­ Cymbaloporetta squammosa Lynts and hered on their umbilical sides to phosphatic Pfister (1967) crusts (samples 3363 no. 1 and no. 3) that Asterigerinacea Amphistegina madagascariensis*** Hansen (1972) formed in the lowermost portion of the Amphistegina lobifera Leutenegger and impinging Peruvian oxygen-minimum zone, Hansen (1979) under the following oceanographic parame­ Nonionacea ters: O2 = 0.06 ml/liter (2.7 lJ-M), temper­ Nonionella stella Leutenegger and ature 9.7°C, and salinity 34.7%0. Current Hansen (1979) = = Nonionidae (unspecified) Jahn (1953) velocity averaged 7 em/sec. Large encrusting Rotaliacea agglutinated foraminifera were dispersed over Ammonia spp. Banner and the surface of the crusts at a density of ap­ Williams proximately 46 specimens per square centi­ (1973) Nummulitacea meter on 3363 no. 3 (Resig and Glenn 1997), Nummulitidae (unspecified) Jahn (1953) their food presumably drawn from benthic Globigerinacea microbes and phytodetritus settling from the Planktonic species (unspecified) Towe (1971) high-productivity surface waters associated Globigerinoides sacculifer Anderson and with Peruvian upwelling. Of the total fora­ Be (1976) Globorotalia menardii Hemleben et al. minifera occupying the habitat, less than 1% (1977) were adherent forms; the remainder were permanently affixed to the hard phosphatic­ ,.., Lenticulina 11lidwayensis; **, Lo11le/lodiscorbis aguayoi; ***, A11lphistegi1lll lessonii. crust substrate. The bathymetric range exhibited by P. ornata may relate directly to the dissolved oxygen profile off the western Americas, where oxygen depletion begins on the outer Sieve Plates and Habitat of Planulina Resig and Glenn 105 6 8 10 12 14 16 18 20 0 12.0°5 Transect ~ 7cmls 3372'1 200 :I> 23 cmls < Cfl III 0 10 cmls S 3 ~ (') c .r: :: ii. 400 CD GI no data 0 a s.. ;;: '" 11 cmls .,0" == ~ 7cmls 6cmls 100 200 300 Dissolved O2 (flM) 34.5 34.6 34.7 34.8 34.9 35.0 35.1 I ! I ! I I I Salinity (%.) FIGURE 1. Location of samples 3363 no. 1 and no. 3 containing Planulina ornata at 11" 59.89' S, 77" 50.91' W, water depth 517 m. Oceanographic parameters as well as other samples examined from along a depth traverse are also shown. (After Resig and Glenn 1997.) shelf (Bandy 1961, Ingle et al. 1980, Resig geous epifaunal adaptation for attachment in and Glenn 1997) and where specimens of P. turbulent water and for stability in traveling ornata are most frequent. Bandy (1963) re­ on the surface of the seafloor (Corliss 1985). ported unusually large (1 to 2 mm diameter) The aperture, an equatorial arch at the base specimens of the species from the bottom of of the final chamber (Figure 2A), is sur­ three California Borderland basins with dis­ mounted by a lip and extends beneath an um­ solved oxygen values of 0.2-0.4 ml/liter. bilical folium onto the umbilical side. Large pores are densely and evenly distributed over the chamber surfaces on both sides ofthe test. TEST MORPHOLOGY AND PORE The pores of P. ornata are 6 to 10 11m in MICROSTRUCTURE diameter and are closed by calcified sieve Details of the morphology of Planulina ornata plates in which the rnicropore diameters are are shown in Figure 2A-M. Planulina ornata 0.5 to 1.0 11m (Figure 2D and F). Because has a compressed, keeled, trochospiral test in these sieve plates are formed at the base of which the flat umbilical side is only slightly the pore only when a new chamber is added, less evolute than the spiral side, which is of they are no longer visible in the early cham­ very low convexity (Figure 2A-C). Plano­ bers after new laminae added to the test convex shapes are considered to be advanta- deepen the pore canals (Figure 2G). Some of Sieve Plates and Habitat of Planulina . Resig and Glenn 107 these sieve plates appear to be in direct and sieve plates (LeCalvez 1947, Jahn 1953), contact with the interior, whereas others are which effectively block the passage of pseu­ covered by the test's organic lining (Figure dopodia (Berthold 1976). In some instances, 2E, H, and M). The absence of the lining however, thin cytoplasmic strands have been may be an artifact of preservation or may seen to penetrate microporous sieve plates provide some contact of the endoplasm with (Febvre-Chevalier 1971, Hansen 1972, An­ the test's exterior.
Recommended publications
  • Recent and Quaternary Foraminifera Collected Around New Caledonia
    Plates 6/1 & 6/2 Recent and Quaternary foraminifera collected around New Caledonia Jean-Pierre DEBENAY 1 & Guy CABIOCH 2 1 UPRES EA, Universite d'Angers, 2 Bd Lavoisiere, 49045, Angers cedex, France 2 Institut de Recherche pour Developpement, UR 055, Paliotropique, Centre IRD, BPA5, 98848 Noumea cedex, Nouvelle-Caledonie [email protected] Abstract The compilation of the works carried out on Recent and Quaternary foraminifera collected in the waters surrounding New Caledonia allowed us to identify 574 species. These species are listed according to the classification of Loeblish & Tappan (1988), updated for the Recent species by Debenay et al. (1996). Their affinity with microfaunas from other regions is briefly discussed. Resume La compilation des travaux sur les foramini:feres actuels et quaternaires recoltes dans les eaux entourant la Nouvelle-Caledonie nous a permis de repertorier 574 especes. Ces especes sont presen­ tees selon la classification de Loeblish & Tappan (1988), mise ajour pour les especes actuelles par Debenay et al. (1996). Leur affinite avec les microfaunes d'autres regions est discutee brievement. Introduction The first study about foraminifera from the southwestern Pacific near New Caledonia was carried out by Brady (1884) during the voyage of H.M.S. Challenger (1873-1876), updated by Barker (1960), The nearest station was station 177, near Vanuatu (16°45'S-168°5'E). However, studies concerning directly New Caledonia began much later, with partial and local inventories in coastal samples (Gambini,1958, 1959; Renaud-Debyser, 1965; Toulouse, 1965, 1966). Samples of recent and fossil sediments collected during the Singer-Polignac mission (1960-1965) were further used for several studies of foraminiferal assemblages (Coudray & Margerel, 1974; Coudray, 1976; Margerel, 1981).
    [Show full text]
  • 389. Distribution and Ecology of Benthonic Foraminifera in the Sediments of the Andaman Sea W
    CO Z':TRIB UTIONS FROM THE CUSHMAN FOUN DATION FOR FORAMIN IFE!RAL RESEARCH 123 CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH VOLUM E XXI, PART 4, OCTOBER 1970 389. DISTRIBUTION AND ECOLOGY OF BENTHONIC FORAMINIFERA IN THE SEDIMENTS OF THE ANDAMAN SEA W. E. FRERICHS University of Wyoming, Laramie, Wyoming ABSTRACT the extreme southern and southwestern parts of the Fo raminiferal a..<Jtiemblages in sediments of the Andaman sea (text fig. 1). Sea characterize five fauna l provinces. each of w hich Is de fined by ecologic factors, S lightly euryhallne conditions Cores were split routinely in the laboratory, and and a rela tively coarse grained substrate chal'acteri ze the upper 5 cm of each was sampled for the faunal the delta-front faunal province. Extremely hig h ra tes of analyses. These core sections and representative sedimentation. euryhaline co ndition~. and clay substl'ate are typical of the Gulf o f Mal'taban province. Extre me ly fractions of the grab samples were dried and s low rates of sedimentation a nd a coarse-grained s ub­ weighed and then washed on a 250-mesh Tyler strate characterize the Mergul platform province. Normal screen (0.061 mm openings). salinities and average rates of sedimentation characterize the Andaman-N lcobar R idge faunal province, Sediments Samples used to determine the rel ative abun­ having a h igh organic content and Indicating active solu­ dance of species at the tops of the cores and in the tion of calcium carbonate occur in the basin fa unal PI'ovlnce.
    [Show full text]
  • Tsunami-Generated Rafting of Foraminifera Across the North Pacific Ocean
    Aquatic Invasions (2018) Volume 13, Issue 1: 17–30 DOI: https://doi.org/10.3391/ai.2018.13.1.03 © 2018 The Author(s). Journal compilation © 2018 REABIC Special Issue: Transoceanic Dispersal of Marine Life from Japan to North America and the Hawaiian Islands as a Result of the Japanese Earthquake and Tsunami of 2011 Research Article Tsunami-generated rafting of foraminifera across the North Pacific Ocean Kenneth L. Finger University of California Museum of Paleontology, Valley Life Sciences Building – 1101, Berkeley, CA 94720-4780, USA E-mail: [email protected] Received: 9 February 2017 / Accepted: 12 December 2017 / Published online: 15 February 2018 Handling editor: James T. Carlton Co-Editors’ Note: This is one of the papers from the special issue of Aquatic Invasions on “Transoceanic Dispersal of Marine Life from Japan to North America and the Hawaiian Islands as a Result of the Japanese Earthquake and Tsunami of 2011." The special issue was supported by funding provided by the Ministry of the Environment (MOE) of the Government of Japan through the North Pacific Marine Science Organization (PICES). Abstract This is the first report of long-distance transoceanic dispersal of coastal, shallow-water benthic foraminifera by ocean rafting, documenting survival and reproduction for up to four years. Fouling was sampled on rafted items (set adrift by the Tohoku tsunami that struck northeastern Honshu in March 2011) landing in North America and the Hawaiian Islands. Seventeen species of shallow-water benthic foraminifera were recovered from these debris objects. Eleven species are regarded as having been acquired in Japan, while two additional species (Planogypsina squamiformis (Chapman, 1901) and Homotrema rubra (Lamarck, 1816)) were obtained in the Indo-Pacific as those objects drifted into shallow tropical waters before turning north and east to North America.
    [Show full text]
  • The Benthic Foraminifer Stomatorbina Binkhorsti (Reuss, 1862): Taxonomic Review and Ecological Insights
    Austrian Journal of Earth Sciences Vienna 2019 Volume 112/2 195-206 DOI: 10.17738/ajes.2019.0011 The benthic foraminifer Stomatorbina binkhorsti (Reuss, 1862): Taxonomic review and ecological insights Felix SCHLAGINTWEIT1,* & Sylvain RIGAUD2 1) Lerchenauerstr. 167, 80935 München, Germany 2) Asian School of the Environment, 62 Nanyang Drive, 637459 Singapore *) Corresponding author: [email protected] KEYWORDS Benthic foraminifera, composite wall, Palaeocene, Juvenarium, Kambühel Formation, Lower Austria Abstract The benthic foraminifer Rosalina binkhorsti Reuss, 1862, was cosmopolitan in Late Cretaceous to early Paleogene shal- low-water seas. It possesses a distinctive composite wall made of a continuous, agglutinated layer discontinuously covered by secondary hyaline outer deposits. Its taxonomic position, phylogeny, morphology, wall structure, and compo- sition have been debated for a long time. Based on abundant, well-preserved material from the Danian of the Kambühel Formation in the Northern Calcareous Alps, Austria, we identify elements in the here emended species Stomatorbina binkhorsti which support a strong affinity to the order Textulariida, within the genus Stomatorbina Dorreen, 1948. Usually regarded as free (non-fixing), S. binkhorsti is here illustrated attached to small bioclasts in high-energy carbonate settings. The attached specimens are juvenile forms with a wall covered by massive hyaline deposits. This observation suggests that secondary lamellar parts added to the wall may have served for stabilisation or fixation to the substrate. Rosalina binkhorsti Reuss, 1862, war eine in den Flachwassermeeren der Oberkreide und des frühen Paläogens kos- mopolitische benthonische Foraminifere. Sie besitzt eine zusammengesetzte Wand, bestehend aus einer kontinuierlichen agglutinierten Lage welche diskontinuierlich von äusseren sekundär-hyalinen Abschnitten bedeckt ist.
    [Show full text]
  • Neogene Benthic Foraminifera from the Southern Bering Sea (IODP Expedition 323)
    Palaeontologia Electronica palaeo-electronica.org Neogene benthic foraminifera from the southern Bering Sea (IODP Expedition 323) Eiichi Setoyama and Michael A. Kaminski ABSTRACT This study describes a total of 95 calcareous benthic foraminiferal taxa from the Pliocene–Pleistocene recovered from IODP Hole U1341B in the southern Bering Sea with illustrations produced with an optical microscope and SEM. The benthic foramin- iferal assemblages are mostly dominated by calcareous taxa, and poorly diversified agglutinated forms are rare or often absent, comprising only minor components. Elon- gate, tapered, and/or flattened planispiral infaunal morphotypes are common or domi- nate the assemblages reflecting the persistent high-productivity and hypoxic conditions in the deep Bering Sea. Most of the species found in the cores are long-ranging, but we observe the extinction of several cylindrical forms that disappeared during the mid- Pleistocene Climatic Transition. Eiichi Setoyama. Earth Sciences Department, Research Group of Reservoir Characterization, King Fahd University of Petroleum & Minerals, Dhahran, 31261, Saudi Arabia current address: Energy & Geoscience Institute, University of Utah, 423 Wakara Way, Suite 300, Salt Lake City, Utah 84108, USA [email protected] Michael A. Kaminski. Earth Sciences Department, Research Group of Reservoir Characterization, King Fahd University of Petroleum & Minerals, Dhahran, 31261, Saudi Arabia [email protected] Keywords: Bering Sea; biostratigraphy; foraminifera; palaeoceanography; Pliocene-Pleistocene; taxonomy Submission: 19 February 2014. Acceptance: 1 July 2015 INTRODUCTION the foraminiferal assemblages and palaeoceano- graphic proxies in continuously-cored sections in The Bering Sea is a large, permanently the deeper, southern part of the Bering Sea, with hypoxic deep basin that has a well-developed oxy- an aim toward assessing the effects of climate gen-minimum zone (Takahashi et al., 2011).
    [Show full text]
  • This Article Was Published in an Elsevier Journal. the Attached Copy Is Furnished to the Author for Non-Commercial Research
    This article was published in an Elsevier journal. The attached copy is furnished to the author for non-commercial research and education use, including for instruction at the author’s institution, sharing with colleagues and providing to institution administration. Other uses, including reproduction and distribution, or selling or licensing copies, or posting to personal, institutional or third party websites are prohibited. In most cases authors are permitted to post their version of the article (e.g. in Word or Tex form) to their personal website or institutional repository. Authors requiring further information regarding Elsevier’s archiving and manuscript policies are encouraged to visit: http://www.elsevier.com/copyright Author's personal copy Available online at www.sciencedirect.com Marine Micropaleontology 66 (2008) 233–246 www.elsevier.com/locate/marmicro Molecular phylogeny of Rotaliida (Foraminifera) based on complete small subunit rDNA sequences ⁎ Magali Schweizer a,b, , Jan Pawlowski c, Tanja J. Kouwenhoven a, Jackie Guiard c, Bert van der Zwaan a,d a Department of Earth Sciences, Utrecht University, The Netherlands b Geological Institute, ETH Zurich, Switzerland c Department of Zoology and Animal Biology, University of Geneva, Switzerland d Department of Biogeology, Radboud University Nijmegen, The Netherlands Received 18 May 2007; received in revised form 8 October 2007; accepted 9 October 2007 Abstract The traditional morphology-based classification of Rotaliida was recently challenged by molecular phylogenetic studies based on partial small subunit (SSU) rDNA sequences. These studies revealed some unexpected groupings of rotaliid genera. However, the support for the new clades was rather weak, mainly because of the limited length of the analysed fragment.
    [Show full text]
  • Taxonomy and Distribution of Meiobenthic Intertidal Foraminifera in the Coastal Tract of Midnapore (East), West Bengal, India
    ISSN: 2347-3215 Volume 2 Number 3 (2014) pp. 98-104 www.ijcrar.com Taxonomy and distribution of meiobenthic intertidal foraminifera in the coastal tract of Midnapore (East), West Bengal, India D.Ghosh1, S. Majumdar2 and S.K.Chakraborty1* 1Department of Zoology, Vidyasagar University, Midnapore- 721102, West Bengal, India 2S.D. Marine biological research institute, Sagarisland, West Bengal, 743372, India *Corresponding author KEYWORDS A B S T R A C T Taxonomy and distribution of recent meiobenthic intertidal foraminifera in Rotaliida, the coastal tract of Midnapore District have been studied for a period of two Miliolida, Lituolida, years (March, 2009- February, 2011). A total of 44 meiobenthic foraminiferal Lagenida. species belonging to 22 genera, 17 families, 14 super families and 7 orders has been recorded from the intertidal belts of this coastal environment. Faunal assemblages revealed a dominance of the order Rotaliida (20 species) followed by the order Miliolida (9 species), order Lituolida (7 species), Lagenida (3 species), Trochamminida (2 species), Buliminida (2 species) and order Textulariida (1 species). Asterorotalia trispinosa, A. multispinosa, A. dentata, Ammonia beccarii, A. tepida, Miliammina fusca, Quinqueloculina seminulum, Trochammina inflata, Ammobaculites agglutinans, Elphidium hispidulum, and E. crispum were found to be the most abundant foraminifera recorded from different study sites during different seasons. Introduction Studies on recent benthic foraminifera absolutely rare (Ghosh, 1966; Majumdar et. along the intertidal beach sediments of al., 1996, 1999; Majumdar, 2004). Murray, India were very few. Occurances of recent 2006 had reported intertidal recent benthic benthic foraminifera along the west coast of foraminifera from the West Bengal coast. India has been reported by Bhalla et.
    [Show full text]
  • Protista (PDF)
    1 = Astasiopsis distortum (Dujardin,1841) Bütschli,1885 South Scandinavian Marine Protoctista ? Dingensia Patterson & Zölffel,1992, in Patterson & Larsen (™ Heteromita angusta Dujardin,1841) Provisional Check-list compiled at the Tjärnö Marine Biological * Taxon incertae sedis. Very similar to Cryptaulax Skuja Laboratory by: Dinomonas Kent,1880 TJÄRNÖLAB. / Hans G. Hansson - 1991-07 - 1997-04-02 * Taxon incertae sedis. Species found in South Scandinavia, as well as from neighbouring areas, chiefly the British Isles, have been considered, as some of them may show to have a slightly more northern distribution, than what is known today. However, species with a typical Lusitanian distribution, with their northern Diphylleia Massart,1920 distribution limit around France or Southern British Isles, have as a rule been omitted here, albeit a few species with probable norhern limits around * Marine? Incertae sedis. the British Isles are listed here until distribution patterns are better known. The compiler would be very grateful for every correction of presumptive lapses and omittances an initiated reader could make. Diplocalium Grassé & Deflandre,1952 (™ Bicosoeca inopinatum ??,1???) * Marine? Incertae sedis. Denotations: (™) = Genotype @ = Associated to * = General note Diplomita Fromentel,1874 (™ Diplomita insignis Fromentel,1874) P.S. This list is a very unfinished manuscript. Chiefly flagellated organisms have yet been considered. This * Marine? Incertae sedis. provisional PDF-file is so far only published as an Intranet file within TMBL:s domain. Diplonema Griessmann,1913, non Berendt,1845 (Diptera), nec Greene,1857 (Coel.) = Isonema ??,1???, non Meek & Worthen,1865 (Mollusca), nec Maas,1909 (Coel.) PROTOCTISTA = Flagellamonas Skvortzow,19?? = Lackeymonas Skvortzow,19?? = Lowymonas Skvortzow,19?? = Milaneziamonas Skvortzow,19?? = Spira Skvortzow,19?? = Teixeiromonas Skvortzow,19?? = PROTISTA = Kolbeana Skvortzow,19?? * Genus incertae sedis.
    [Show full text]
  • Mikropaläontologie (Foraminiferen, Ostrakoden), Biostratigraphie Und
    abhandlungen Band 1 - Teil 1 Mikropaläontologie (Foraminiferen, Ostrakoden), Biostratigraphie und fazielle Entwicklung der Kreide von Nordsomalia mit einem Beitrag zur geodynamischen Entwicklung des östlichen Gondwana im Mesozoikum und frühen Känozoikum Micropalaeontology (Foraminiferida, Ostracoda), biostratigraphy and facies development of the Cretaceous of Northern Somalia including a contribution concerning the geodynamic development of eastern Gondwana during the Cretaceous to basal Paleocene Peter LUGER (†) TEXTBAND Landshut, 06. Dezember 2018 ISSN 2626-4161 (Print) ISSN 2626-9864 (Online) ISBN 978-3-947953-00-4 (Gesamtausgabe) ISBN 978-3-947953-01-1 (Band 1 - Teil 1) ISBN 978-3-947953-02-8 (Band 1 - Teil 2) Die Zeitschrift "documenta naturae abhandlungen" ist die Fortsetzung der Sonderband-Reihe der "Zeitschrift Documenta naturae", begründet 1976 in Landshut. Copyright © 2018 amh-Geo Geowissenschaftlicher Dienst, Aham bei Landshut Alle Rechte vorbehalten. - All rights reserved. Der/die Autor(en) sind verantwortlich für den Inhalt der Beiträge, für die Gesamtgestaltung Herausgeber und Verlag. Das vorliegende Werk einschließlich aller seiner Teile ist urheberrechtlich geschützt. Jede Verwendung, auch auszugsweise, insbesondere Übersetzungen, Nachdrucke, Vervielfältigungen jeder Art, Mikroverfilmungen, Einspeicherungen in elektronische Systeme, bedarf der schriftlichen Genehmigung des Verlages. ISSN 2626-4161 (Print) ISSN 2626-9864 (Online) ISBN 978-3-947953-00-4 (Gesamtausgabe) ISBN 978-3-947953-01-1 (Band 1 - Teil 1) ISBN 978-3-947953-02-8
    [Show full text]
  • Foraminiferal Distribution Off the Southern Tip of India to Understand
    Foraminiferal distribution off the southern tip of India to understand its response to cross basin water exchange and to reconstruct seasonal monsoon intensity during the Late Quaternary Thesis submitted to the Goa University School of Earth, Ocean, and Atmospheric Sciences for the award of degree of Doctor of Philosophy by Dharmendra Pratap Singh Goa University School of Earth, Ocean, and Atmospheric Sciences, Goa University (Micropaleontology Laboratory, Geological Oceanography Division CSIR- National Institute of Oceanography, Dona Paula, Goa) April 2019 i Declaration As required under the university ordinance OA.19, I hereby state that the present thesis entitled “Foraminiferal distribution off the southern tip of India to understand its response to cross basin water exchange and to reconstruct seasonal monsoon intensity during the Late Quaternary” is my original contribution and the same has not been submitted on any pervious occasion. To the best of my knowledge, the present study is the first comprehensive work of its kind from the area mentioned. Literature related to the scientific objectives has been cited. Due acknowledgments have been made wherever facilities and suggestions have been availed of. Dharmendra Pratap Singh ii Certificate As required under the university ordinance OA.19, I certify that the thesis entitled “Foraminiferal distribution off the southern tip of India to understand its response to cross basin water exchange and to reconstruct seasonal monsoon intensity during the Late Quaternary” submitted by Mr. Dharmendra Pratap Singh for the award of the degree of Doctor of Philosophy in the School of Earth, Ocean, and Atmospheric Sciences is based on original work carried out by him under my supervision.
    [Show full text]
  • Ecology and Systematics of Foraminifera in Two Thalassia Habitats, Jamaica, West Indies
    Ecology and Systematics of Foraminifera in Two Thalassia Habitats, Jamaica, West Indies MARTIN A. BUZAS, ROBERTA K. SMITH, and KENNETH A. BEEM SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY • NUMBER 31 SERIES PUBLICATIONS OF THE SMITHSONIAN INSTITUTION Emphasis upon publication as a means of "diffusing knowledge" was expressed by the first Secretary of the Smithsonian. In his formal plan for the Institution, Joseph Henry outlined a program that included the following statement: "It is proposed to publish a series of reports, giving an account of the new discoveries in science, and of the changes made from year to year in all branches of knowledge." This theme of basic research has been adhered to through the years by thousands of titles issued in series publications under the Smithsonian imprint, commencing with Smithsonian Contributions to Knowledge in 1848 and continuing with the following active series: Smithsonian Contributions to Anthropology Smithsonian Contributions to Astrophysics Smithsonian Contributions to Botany Smithsonian Contributions to the Earth Sciences Smithsonian Contributions to the Marine Sciences Smithsonian Contributions to Paleobiology Smithsonian Contributions to Zoology Smithsonian Studies in Air and Space Smithsonian Studies in History and Technology In these series, the Institution publishes small papers and full-scale monographs that report the research and collections of its various museums and bureaux or of professional colleagues in the world cf science and scholarship. The publications are distributed by mailing lists to libraries, universities, and similar institutions throughout the world. Papers or monographs submitted for series publication are received by the Smithsonian Institution Press, subject to its own review for format and style, only through departments of the various Smithsonian museums or bureaux, where the manuscripts are given substantive review.
    [Show full text]
  • Studying Found, Together
    STUDIES ON THE FAUNA OF CURAÇAO AND OTHER CARIBBEAN ISLANDS: No. 181 Rare and remarkable Foraminifera of the Caribbean Sea by J. Hofker ('s-Gravenhage) page fig. INTRODUCTION 1 constricta 2 1 Hyperammina n. sp Polysiphotrocha siphonata Seiglie 3 2 Tritaxis siphonifera (Cushman) 5 3 Carterina caribbeana 6 4 n. sp Gaudryinaparvula (Cushman) 8 5 6 Cribrostomoides spiculiferus:n. sp 10 Cornuloculina inconstans (Brady) 11 7 Robertina subcylindrica (Brady) 12 8-9 Patellina altiformis Cushman 14 10 Tretomphalus Moebius 16 Tretomphalus bulloides (d'Orbigny) 20 11 Pseudotretomphaluss n. gen 21 23 12-13 Pseudotretomphalusplanus (Cushman) .... milletti 23 14-15 Pseudotretomphalus (Heron-Allen) ... 24 16 Pseudotretomphalus grandis (Cushman) .... Siphoninellasoluta (Brady) 25 17 Globigerinita glutinata (Egger) 26 18-19 Candeina nitida d'Orbigny 27 20 Globorotalia punctulata (d'Orbigny) 28 21-26 REFERENCES 30 When studying samples from the Virgin Islands area, some new Foraminifera were found, together with other species of which little was known about their internal structure and taxonomic status. 2 Most specimens came from samples W of St. Croix, collected by Th. Mortensen at about 17.5°N 64°W, depths about 200 and 300 These rich in benthonic but m. samples are very Foraminifera, re- latively poor in planktonic species all of which are relatively small and with thin tests. This material contained very many specimens already described in previous publications (e.g. Hofker 1956, 1972 and 1976). The materialstudied has been deposited in the collections of the Netherlands Geological Survey at Haarlem, Netherlands. Family ASTRORHIZIDAE Brady, 1881 Subfamily RHIZAMMININAE Rhumbler, 1895 Genus Hyperammina Brady, 1878 Hyperammina constricta n.
    [Show full text]