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Phenological Responses to Climate in the Alberta Native Flora: Herbarium Specimens Reveal Differential Responsiveness Between Species in Mesic and Xeric Habitats
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2019-03-01 Phenological responses to climate in the Alberta native flora: Herbarium specimens reveal differential responsiveness between species in mesic and xeric habitats Porto, Cassiano Porto, C. (2019). Phenological responses to climate in the Alberta native flora: Herbarium specimens reveal differential responsiveness between species in mesic and xeric habitats (Unpublished master's thesis). University of Calgary, Calgary, AB. http://hdl.handle.net/1880/109929 master thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Phenological responses to climate in the Alberta native flora: Herbarium specimens reveal differential responsiveness between species in mesic and xeric habitats by Cassiano Porto A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE GRADUATE PROGRAM IN BIOLOGICAL SCIENCES CALGARY, ALBERTA MARCH, 2019 © Cassiano Porto 2019 UNIVERSITY OF CALGARY Phenological responses to climate in the Alberta native flora: Herbarium specimens reveal differential responsiveness between species in mesic and xeric habitats by Cassiano Porto A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE GRADUATE PROGRAM IN BIOLOGICAL SCIENCES Research Supervisor: Dr. -
Plant List Bristow Prairie & High Divide Trail
*Non-native Bristow Prairie & High Divide Trail Plant List as of 7/12/2016 compiled by Tanya Harvey T24S.R3E.S33;T25S.R3E.S4 westerncascades.com FERNS & ALLIES Pseudotsuga menziesii Ribes lacustre Athyriaceae Tsuga heterophylla Ribes sanguineum Athyrium filix-femina Tsuga mertensiana Ribes viscosissimum Cystopteridaceae Taxaceae Rhamnaceae Cystopteris fragilis Taxus brevifolia Ceanothus velutinus Dennstaedtiaceae TREES & SHRUBS: DICOTS Rosaceae Pteridium aquilinum Adoxaceae Amelanchier alnifolia Dryopteridaceae Sambucus nigra ssp. caerulea Holodiscus discolor Polystichum imbricans (Sambucus mexicana, S. cerulea) Prunus emarginata (Polystichum munitum var. imbricans) Sambucus racemosa Rosa gymnocarpa Polystichum lonchitis Berberidaceae Rubus lasiococcus Polystichum munitum Berberis aquifolium (Mahonia aquifolium) Rubus leucodermis Equisetaceae Berberis nervosa Rubus nivalis Equisetum arvense (Mahonia nervosa) Rubus parviflorus Ophioglossaceae Betulaceae Botrychium simplex Rubus ursinus Alnus viridis ssp. sinuata Sceptridium multifidum (Alnus sinuata) Sorbus scopulina (Botrychium multifidum) Caprifoliaceae Spiraea douglasii Polypodiaceae Lonicera ciliosa Salicaceae Polypodium hesperium Lonicera conjugialis Populus tremuloides Pteridaceae Symphoricarpos albus Salix geyeriana Aspidotis densa Symphoricarpos mollis Salix scouleriana Cheilanthes gracillima (Symphoricarpos hesperius) Salix sitchensis Cryptogramma acrostichoides Celastraceae Salix sp. (Cryptogramma crispa) Paxistima myrsinites Sapindaceae Selaginellaceae (Pachystima myrsinites) -
WRITTEN FINDINGS of the WASHINGTON STATE NOXIOUS WEED CONTROL BOARD 2018 Noxious Weed List Proposal
DRAFT: WRITTEN FINDINGS OF THE WASHINGTON STATE NOXIOUS WEED CONTROL BOARD 2018 Noxious Weed List Proposal Scientific Name: Tussilago farfara L. Synonyms: Cineraria farfara Bernh., Farfara radiata Gilib., Tussilago alpestris Hegetschw., Tussilago umbertina Borbás Common Name: European coltsfoot, coltsfoot, bullsfoot, coughwort, butterbur, horsehoof, foalswort, fieldhove, English tobacco, hallfoot Family: Asteraceae Legal Status: Proposed as a Class B noxious weed for 2018, to be designated for control throughout Washington, except for in Grant, Lincoln, Adams, Benton, and Franklin counties. Images: left, blooming flowerheads of Tussilago farfara, image by Caleb Slemmons, National Ecological Observatory Network, Bugwood.org; center, leaves of T. farfara growing with ferns, grasses and other groundcover species; right, mature seedheads of T. farfara before seeds have been dispersed, center and right images by Leslie J. Mehrhoff, University of Connecticut, Bugwood.org. Description and Variation: The common name of Tussilago farfara, coltsfoot, refers to the outline of the basal leaf being that of a colt’s footprint. Overall habit: Tussilago farfara is a rhizomatous perennial, growing up to 19.7 inches (50 cm tall), which can form extensive colonies. Plants first send up flowering stems in the spring, each with a single yellow flowerhead. Just before or after flowers have formed seeds, basal leaves on long petioles grow from the rhizomes, with somewhat roundish leaf blades that are more or less white-woolly on the undersides. Roots: Plants have long creeping, white scaly rhizomes (Griffiths 1994, Chen and Nordenstam 2011). Rhizomes are branching and have fibrous roots (Barkley 2006). They are also brittle and can break easily (Pfeiffer et al. -
Buzz-Pollination and Patterns in Sexual Traits in North European Pyrolaceae Author(S): Jette T
Buzz-Pollination and Patterns in Sexual Traits in North European Pyrolaceae Author(s): Jette T. Knudsen and Jens Mogens Olesen Reviewed work(s): Source: American Journal of Botany, Vol. 80, No. 8 (Aug., 1993), pp. 900-913 Published by: Botanical Society of America Stable URL: http://www.jstor.org/stable/2445510 . Accessed: 08/08/2012 10:49 Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at . http://www.jstor.org/page/info/about/policies/terms.jsp . JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. Botanical Society of America is collaborating with JSTOR to digitize, preserve and extend access to American Journal of Botany. http://www.jstor.org American Journalof Botany 80(8): 900-913. 1993. BUZZ-POLLINATION AND PATTERNS IN SEXUAL TRAITS IN NORTH EUROPEAN PYROLACEAE1 JETTE T. KNUDSEN2 AND JENS MOGENS OLESEN Departmentof ChemicalEcology, University of G6teborg, Reutersgatan2C, S-413 20 G6teborg,Sweden; and Departmentof Ecology and Genetics,University of Aarhus, Ny Munkegade, Building550, DK-8000 Aarhus,Denmark Flowerbiology and pollinationof Moneses uniflora, Orthilia secunda, Pyrola minor, P. rotundifolia,P. chlorantha, and Chimaphilaumbellata are describedand discussedin relationto patternsin sexualtraits and possibleevolution of buzz- pollinationwithin the group. The largenumber of pollengrains are packedinto units of monadsin Orthilia,tetrads in Monesesand Pyrola,or polyadsin Chimaphila.Pollen is thesole rewardto visitinginsects except in thenectar-producing 0. -
THE ECOLOGICAL FEATURES of FLOWERS and INFLORESCENCES of TWO SPECIES of the GENUS Petasites Miller (ASTERACEAE)
ACTA AGROBOTANICA Vol. 65 (2): 37–46 2012 THE ECOLOGICAL FEATURES OF FLOWERS AND INFLORESCENCES OF TWO SPECIES OF THE GENUS Petasites Miller (ASTERACEAE) Weronika Haratym, Elżbieta Weryszko-Chmielewska Department of Botany, University of Life Sciences in Lublin, Akademicka 15, 20-950 Lublin, Poland e-mail: [email protected] Received: 06.02.2012 Abstract The above-mentioned species are included The structural features of flowers and inflorescences of in bee plants (Bornus, 1989; Lipiń ski, 2010). Petasites hybridus and P. albus were compared. Only individu- Due to their flowering period (III-V), they can provide als producing flower heads with male flowers and few female a source of early spring pollen and nectar to insects flowers were found in the studied populations. Light microsco- which visit their flowers in large numbers under fa- py (LM) and scanning electron microscopy (SEM) were used vourable conditions. Many papers show that plant for examination. material obtained from their leaves and underground The present study shows that the stems of the above- parts (rhizomes, roots) has been used in herbalism -mentioned species differed in height and number of flower he- and medicine since ancient times (Podhajska and ads, but the number of flowers per head was similar. Larger flo- Rivola, 1992; Schaffner, 1996; Siegenha- wers were found on the stems of P. albus. The following features ler and Neuenschwander, 1996; Wildi et al. has been found to play an important role in pollination ecology: the strongly contrasting colours of the floral parts; on the petals, 1998). However, Smith and Culvenor (1981) the occurrence of several types of cells which can increase the as well as Sadowska (2004) note that P. -
The Effect of Certain Plant Extracts Containing Pyrrolizidine Alkaloids on Lactuca Sativa Radicle Growth
THE EFFECT OF CERTAIN PLANT EXTRACTS CONTAINING PYRROLIZIDINE ALKALOIDS ON LACTUCA SATIVA RADICLE GROWTH OANA-CRISTINA ŞEREMET*, O.T. OLARU**#, D. BĂLĂLĂU***, SIMONA NEGREŞ* * Department of Pharmacology and Clinical Pharmacy, Faculty of Pharmacy, “Carol Davila” University of Medicine and Pharmacy, Bucharest, Romania ** Department of Botany and Cellular Biology, Faculty of Pharmacy, “Carol Davila” University of Medicine and Pharmacy, Bucharest, Romania, #e-mail: [email protected] *** Department of Toxicology, Faculty of Pharmacy, “Carol Davila” University of Medicine and Pharmacy, Bucharest, Romania Abstract: Pyrrolizidine alkaloids (PAs) are toxins synthesized by plants, known to be hepatotoxic, genotoxic and carcinogenic. Tussilago farfara (coltsfoot), Petasites hybridus (common butterbur), Senecio vernalis (eastern groundsel) and Symphytum officinale (comfrey) are species traditionally used in phytotherapy that besides the therapeutic compounds contain PAs. The total PAs and the corresponding N-oxides content of the solid plant extracts were measured using Ehrlich’s method with senecionine as a standard substance. The highest content of PAs was found in Senecio vernalis extract (424.92±9.81 mg%), followed by Symphytum officinale extract (150.24±10.35 mg%) and Petasites hybridus extract (0.021±0.00091 mg%). The lowest concentration was found in Tussilago farfara extract (0.0097±0.00072 mg%). In order to assess the toxicity of the extracts on Lactuca sativa, inhibitory concentrations 50% (IC50) of the root elongation were calculated. All extracts inhibited the elongation of the Lactuca sativa radicles. Senecio vernalis extract exhibited the highest toxicity, whilst Symphytum officinale extract, having the second highest concentration of PAs, has the highest IC50. Further studies should be performed in order to determine whether the inhibitory effect is due to the PAs content and how it is influenced by the other components of the extracts. -
Addendum to the Guide to the Natural Communities of the Delaware Estuary
ADDENDUM TO THE UIDE TO THE ATURAL OMMUNITIES G N C OF THE DELAWARE ESTUARY SEPTEMBER0 2009 Citation: Largay, E. and L. Sneddon. 2009. Addendum to the Guide to the Ecological Systems and Vegetation Communities of the Delaware Estuary. NatureServe. Arlington, Virginia. Partnership for the Delaware Estuary, Report #09-XX. 112 pp. PDE Report No. 09-XX Copyright © 2009 NatureServe COVER PHOTOS Top L: Overwash Dunes, photo from Delaware Natural Heritage Program Top R: Coastal Plain Muck Pondshore, photo by Kathleen Strakosch Walz, New Jersey Natural Heritage Program Bottom L: Dry Oak Hickory Forest, photo by Tony Davis, Pennsylvania Natural Heritage Program Bottom R: Inland Dune and Ridge Forest/Woodland, Kathleen Strakosch Walz, New Jersey Natural Heritage Program ADDENDUM TO THE GUIDE TO THE NATURAL COMMUNITIES OF THE DELAWARE ESTUARY Ery Largay Lesley Sneddon September 2009 Acknowledgements: This work was made possible through funding from the Delaware Estuary Program (EPA 320 Funding). Kristin Snow and Mary Russo from NatureServe provided essential data management services to develop this report and report format. Robert Coxe and Bill McAvoy from the Delaware Natural Heritage Program, Kathleen Strakosch Walz from the New Jersey Natural Heritage Program, Tony Davis from the Pennsylvania Natural Heritage Program, Linda Kelly and Karl Anderson, independent botanists, provided ecological expertise, energy and insight. Mark Anderson and Charles Ferree from The Nature Conservancy developed ecological systems maps to accompany this work. Danielle Kreeger, Laura Whalen, and Martha-Maxwell Doyle from the Partnership for the Delaware Estuary provided support and guidance throughout this project. We thank everyone who helped us with this effort. -
Checklist Flora of the Former Carden Township, City of Kawartha Lakes, on 2016
Hairy Beardtongue (Penstemon hirsutus) Checklist Flora of the Former Carden Township, City of Kawartha Lakes, ON 2016 Compiled by Dale Leadbeater and Anne Barbour © 2016 Leadbeater and Barbour All Rights reserved. No part of this publication may be reproduced, stored in a retrieval system or database, or transmitted in any form or by any means, including photocopying, without written permission of the authors. Produced with financial assistance from The Couchiching Conservancy. The City of Kawartha Lakes Flora Project is sponsored by the Kawartha Field Naturalists based in Fenelon Falls, Ontario. In 2008, information about plants in CKL was scattered and scarce. At the urging of Michael Oldham, Biologist at the Natural Heritage Information Centre at the Ontario Ministry of Natural Resources and Forestry, Dale Leadbeater and Anne Barbour formed a committee with goals to: • Generate a list of species found in CKL and their distribution, vouchered by specimens to be housed at the Royal Ontario Museum in Toronto, making them available for future study by the scientific community; • Improve understanding of natural heritage systems in the CKL; • Provide insight into changes in the local plant communities as a result of pressures from introduced species, climate change and population growth; and, • Publish the findings of the project . Over eight years, more than 200 volunteers and landowners collected almost 2000 voucher specimens, with the permission of landowners. Over 10,000 observations and literature records have been databased. The project has documented 150 new species of which 60 are introduced, 90 are native and one species that had never been reported in Ontario to date. -
Threats to Australia's Grazing Industries by Garden
final report Project Code: NBP.357 Prepared by: Jenny Barker, Rod Randall,Tony Grice Co-operative Research Centre for Australian Weed Management Date published: May 2006 ISBN: 1 74036 781 2 PUBLISHED BY Meat and Livestock Australia Limited Locked Bag 991 NORTH SYDNEY NSW 2059 Weeds of the future? Threats to Australia’s grazing industries by garden plants Meat & Livestock Australia acknowledges the matching funds provided by the Australian Government to support the research and development detailed in this publication. This publication is published by Meat & Livestock Australia Limited ABN 39 081 678 364 (MLA). Care is taken to ensure the accuracy of the information contained in this publication. However MLA cannot accept responsibility for the accuracy or completeness of the information or opinions contained in the publication. You should make your own enquiries before making decisions concerning your interests. Reproduction in whole or in part of this publication is prohibited without prior written consent of MLA. Weeds of the future? Threats to Australia’s grazing industries by garden plants Abstract This report identifies 281 introduced garden plants and 800 lower priority species that present a significant risk to Australia’s grazing industries should they naturalise. Of the 281 species: • Nearly all have been recorded overseas as agricultural or environmental weeds (or both); • More than one tenth (11%) have been recorded as noxious weeds overseas; • At least one third (33%) are toxic and may harm or even kill livestock; • Almost all have been commercially available in Australia in the last 20 years; • Over two thirds (70%) were still available from Australian nurseries in 2004; • Over two thirds (72%) are not currently recognised as weeds under either State or Commonwealth legislation. -
Kenai National Wildlife Refuge Species List - Kenai - U.S
Kenai National Wildlife Refuge Species List - Kenai - U.S. Fish and Wild... http://www.fws.gov/refuge/Kenai/wildlife_and_habitat/species_list.html Kenai National Wildlife Refuge | Alaska Kenai National Wildlife Refuge Species List Below is a checklist of the species recorded on the Kenai National Wildlife Refuge. The list of 1865 species includes 34 mammals, 154 birds, one amphibian, 20 fish, 611 arthropods, 7 molluscs, 11 other animals, 493 vascular plants, 180 bryophytes, 29 fungi, and 325 lichens. Of the total number of species, 1771 are native, 89 are non-native, and five include both native and non-native subspecies. Non-native species are indicated by dagger symbols (†) and species having both native and non-native subspecies are indicated by double dagger symbols (‡). Fifteen species no longer occur on the Refuge, indicated by empty set symbols ( ∅). Data were updated on 15 October 2015. See also the Kenai National Wildlife Refuge checklist on iNaturalist.org ( https://www.inaturalist.org/check_lists/188476-Kenai-National-Wildlife- Refuge-Check-List ). Mammals ( #1 ) Birds ( #2 ) Amphibians ( #3 ) Fish ( #4 ) Arthropods ( #5 ) Molluscs ( #6 ) Other Animals ( #7 ) Vascular Plants ( #8 ) Other Plants ( #9 ) Fungi ( #10 ) Lichens ( #11 ) Change Log ( #changelog ) Mammals () Phylum Chordata Class Mammalia Order Artiodactyla Family Bovidae 1. Oreamnos americanus (Blainville, 1816) (Mountain goat) 2. Ovis dalli Nelson, 1884 (Dall's sheep) Family Cervidae 3. Alces alces (Linnaeus, 1758) (Moose) 4. Rangifer tarandus (Linnaeus, 1758) (Caribou) Order Carnivora Family Canidae 5. Canis latrans Say, 1823 (Coyote) 6. Canis lupus Linnaeus, 1758 (Gray wolf) 7. Vulpes vulpes (Linnaeus, 1758) (Red fox) Family Felidae 8. Lynx lynx (Linnaeus, 1758) (Lynx) 9. -
Vascular Flora and Geoecology of Mont De La Table, Gaspésie, Québec
RHODORA, Vol. 117, No. 969, pp. 1–40, 2015 E Copyright 2015 by the New England Botanical Club doi: 10.3119/14-07; first published on-line March 11, 2015. VASCULAR FLORA AND GEOECOLOGY OF MONT DE LA TABLE, GASPE´ SIE, QUE´ BEC SCOTT W. BAILEY USDA Forest Service, 234 Mirror Lake Road, North Woodstock, NH 03262 e-mail: [email protected] JOANN HOY 21 Steam Mill Road, Auburn, NH 03032 CHARLES V. COGBILL 82 Walker Lane, Plainfield, VT 05667 ABSTRACT. The influence of substrate lithology on the distribution of many vascular and nonvascular plants has long been recognized, especially in alpine, subalpine, and other rocky habitats. In particular, plants have been classified as dependent on high-calcium substrates (i.e., calcicoles) based on common restriction to habitats developed in calcareous rocks, such as limestone and marble. In a classic 1907 paper on the influence of substrate on plants, M. L. Fernald singled out a particular meadow on Mont de la Table in the Chic-Choc Mountains of Que´bec for its unusual co-occurrence of strict calcicole and calcifuge (i.e., acidophile) plant taxa. We re-located this site, investigated substrate factors responsible for its unusual plant diversity, and documented current plant distributions. No calcareous rocks were found on site. However, inclusions of calcareous rocks were found farther up the mountain. The highest pH and dissolved calcium concentrations in surface waters were found in a series of springs that deliver groundwater, presumably influenced by calcareous rocks up the slope. Within the habitat delineated by common occurrences of calcicole species, available soil calcium varied by a factor of five and soil pH varied by almost 1.5 units, depending on microtopography and relative connection with groundwater. -
Plants & Ecology
Population structure and distribution of the declining endangered forest plant Chimaphila umbellata by Anna Lundell Plants & Ecology The Department of Ecology, 2014/1 Environment and Plant Sciences Stockholm University Population structure and distribution of the declining endangared forest plant Chimaphila umbellata by Anna Lundell Supervisors: Ove Eriksson & Sara Cousins Plants & Ecology The Department of Ecology, 2014/1 Environment and Plant Sciences Stockholm University Plants & Ecology The Department of Ecology, Environment and Plant Sciences Stockholm University S-106 91 Stockholm Sweden © The Department of Ecology, Environment and Plant Sciences ISSN 1651-9248 Printed by FMV Printcenter Cover: Flowering Chimaphila umbellata. Photo by Margareta Edqvist. Summary The occurrence of the rare forest plant Chimaphila umbellata has decreased with approximately 80 % in Uppland and its decline is also reported for other regions in Sweden. Suggested causes of decline include the increasingly shaded conditions in understory habitats and increased competition from Vaccinium myrtillus and graminoid species. The aim of the study was to investigate the effects of various biotic and abiotic conditions on C. umbellata populations with regard to population size, flowering frequency, fruit set and seed production. Conditions analyzed include light inflow, coverage of competitive species, soil nitrogen, continuity of forest cover and soil texture, which were investigated in 38 C. umbellata sites in Uppland and Södermanland, Sweden. Results showed that population size was negatively affected by the coverage of competitive species. Population size was not affected by light availability, but increased shading resulted in a decreased flowering frequency. Fruit set decreased with increasing coverage of competitive species and seed production per capsule decreased with increasing soil nitrogen content.