Santana do Cariri, Ceará, Brazil 1 of Palm Swamps in the South of Ceará Maria Daniely Freire Guerra1, Priscila Orlandini2, Rayane de Tasso Moreira Ribeiro3, Maria Iracema Bezerra de Loiola4 & Rubens Teixeira de Queiroz5 1Universidade Regional do Cariri, 2Universidade Estadual de Campinas, 3Universidade Federal Rural de Pernambuco, 4 5 Universidade Federal do Ceará & Universidade Federal da Paraíba Photos by R.T. de Queiroz, M.I.B. Loiola, and M.D.F. Guerra. Produced by M.D.F. Guerra, and Priscila Orlandini. For more information: Guerra et al. (2020). ©Field Museum (2021) CC BY-NC 4.0. Licensed works are free to use/share/remix with attribution, but commercial use of the original work is not permitted. [fieldguides.fieldmuseum.org] [1363] version 1 9/2021 Palm swamps are typical environment of the Cerrado domain. However, they also occur in ecological refugia, in isolated stretches of drainage headwaters in the Araripe sedimentar basin, in the South of Ceará state. These lie inside the polygon of the northeastern Brazil, where the xerophytic Caatinga vegetation predominates. These refugia betray a complex paleoclimatic and paleoecological evolution, being maintained by local particularities, and favored by the perennial water supply from seepage zones fed by the suspended aquifer of Araripe Plateau. This context allows the coexistence and coevolution of the palm swamp within an area of climatic semiaridity. This guide provides information regarding the recognition and identification of typical species of palm swamp, which were cataloged in a refugium area, locally called the Buriti Valley, in the township of Santana do Cariri, state of Ceará.

1 Mendoncia sp. 2 Ruellia inundata 3 Blepharodon pictum 4 Acrocomia intumescens 5 Attalea speciosa ACANTHACEAE ACANTHACEAE APOCYNACEAE ARECACEAE ARECACEAE Santana do Cariri, Ceará, Brazil 2 Plants of Palm Swamps in the South of Ceará Maria Daniely Freire Guerra1, Priscila Orlandini2, Rayane de Tasso Moreira Ribeiro3, Maria Iracema Bezerra de Loiola4 & Rubens Teixeira de Queiroz5 1Universidade Regional do Cariri, 2Universidade Estadual de Campinas, 3Universidade Federal Rural de Pernambuco, 4 5 Universidade Federal do Ceará & Universidade Federal da Paraíba Photos by R.T. de Queiroz, M.I.B. Loiola, and M.D.F. Guerra. Produced by M.D.F. Guerra, and Priscila Orlandini. For more information: Guerra et al. (2020). ©Field Museum (2021) CC BY-NC 4.0. Licensed works are free to use/share/remix with attribution, but commercial use of the original work is not permitted. [fieldguides.fieldmuseum.org] [1363] version 1 9/2021

6 Attalea speciosa 7 Mauritia flexuosa 8 Mauritia flexuosa 9 Aristolochia papillaris 10 Achyrocline saturejoides ARECACEAE ARECACEAE ARECACEAE ARISTOLOCHIACEAE ASTERACEAE

11 Dasyphyllum sprengelianum 12 Dasyphyllum sprengelianum 13 Delilia biflora 14 Dissothrix sp. 15 Pectis gardneri ASTERACEAE ASTERACEAE ASTERACEAE ASTERACEAE ASTERACEAE

16 Garcinia gardneriana 17 Floscopa glabrata 18 Cyathea delgadoi 19 Cyperus odoratus 20 Fuirena lainzii CLUSIACEAE COMMELINACEAE CYATHEACEAE CYPERACEAE CYPERACEAE Santana do Cariri, Ceará, Brazil 3 Plants of Palm Swamps in the South of Ceará Maria Daniely Freire Guerra1, Priscila Orlandini2, Rayane de Tasso Moreira Ribeiro3, Maria Iracema Bezerra de Loiola4 & Rubens Teixeira de Queiroz5 1Universidade Regional do Cariri, 2Universidade Estadual de Campinas, 3Universidade Federal Rural de Pernambuco, 4 5 Universidade Federal do Ceará & Universidade Federal da Paraíba Photos by R.T. de Queiroz, M.I.B. Loiola, and M.D.F. Guerra. Produced by M.D.F. Guerra, and Priscila Orlandini. For more information: Guerra et al. (2020). ©Field Museum (2021) CC BY-NC 4.0. Licensed works are free to use/share/remix with attribution, but commercial use of the original work is not permitted. [fieldguides.fieldmuseum.org] [1363] version 1 9/2021

21 Rhynchospora latifolia 22 Rhynchospora riparia 23 Aeschynomene sensitiva 24 Aeschynomene sensitiva 25 Aeschynomene sensitiva CYPERACEAE CYPERACEAE FABACEAE FABACEAE

26 Bauhinia ungulata 27 Bauhinia ungulata 28 Calopogonium mucunoides 29 Calopogonium velutinum 30 Calopogonium velutinum FABACEAE FABACEAE FABACEAE FABACEAE FABACEAE

31 Calopogonium velutinum 32 Centrosema sagittatum 33 Chamaecrista nictitans 34 Chamaecrista pilosa 35 Clitoria falcata FABACEAE FABACEAE FABACEAE FABACEAE FABACEAE Santana do Cariri, Ceará, Brazil 4 Plants of Palm Swamps in the South of Ceará Maria Daniely Freire Guerra1, Priscila Orlandini2, Rayane de Tasso Moreira Ribeiro3, Maria Iracema Bezerra de Loiola4 & Rubens Teixeira de Queiroz5 1Universidade Regional do Cariri, 2Universidade Estadual de Campinas, 3Universidade Federal Rural de Pernambuco, 4 5 Universidade Federal do Ceará & Universidade Federal da Paraíba Photos by R.T. de Queiroz, M.I.B. Loiola, and M.D.F. Guerra. Produced by M.D.F. Guerra, and Priscila Orlandini. For more information: Guerra et al. (2020). ©Field Museum (2021) CC BY-NC 4.0. Licensed works are free to use/share/remix with attribution, but commercial use of the original work is not permitted. [fieldguides.fieldmuseum.org] [1363] version 1 9/2021

36 Ctenodon histrix 37 Ctenodon paniculatus 38 Ctenodon viscidulus 39 Desmodium barbatum 40 Desmodium distortum FABACEAE FABACEAE FABACEAE FABACEAE FABACEAE

41 Desmodium distortum 42 Desmodium distortum 43 Macropsychanthus sclerocarpus 44 Mimosa ursina 45 latifolia FABACEAE FABACEAE FABACEAE FABACEAE FABACEAE

46 Zornia reticulata 47 Dicranopteris flexuosa 48 Dicranopteris flexuosa 49 Chelonanthus purpurascens 50 Chelonanthus purpurascens FABACEAE GLEICHENIACEAE GLEICHENIACEAE GENTIANACEAE GENTIANACEAE Santana do Cariri, Ceará, Brazil 5 Plants of Palm Swamps in the South of Ceará Maria Daniely Freire Guerra1, Priscila Orlandini2, Rayane de Tasso Moreira Ribeiro3, Maria Iracema Bezerra de Loiola4 & Rubens Teixeira de Queiroz5 1Universidade Regional do Cariri, 2Universidade Estadual de Campinas, 3Universidade Federal Rural de Pernambuco, 4 5 Universidade Federal do Ceará & Universidade Federal da Paraíba Photos by R.T. de Queiroz, M.I.B. Loiola, and M.D.F. Guerra. Produced by M.D.F. Guerra, and Priscila Orlandini. For more information: Guerra et al. (2020). ©Field Museum (2021) CC BY-NC 4.0. Licensed works are free to use/share/remix with attribution, but commercial use of the original work is not permitted. [fieldguides.fieldmuseum.org] [1363] version 1 9/2021

51 Schultesia guianensis 52 Sinningia elatior 53 Sinningia elatior 54 Hyptis lanceolata 55 Vitex rufescens GENTIANACEAE GESNERIACEAE GESNERIACEAE LAMIACEAE LAMIACEAE

56 Ocotea nitida 57 Utricularia sp. 58 Lycopodiella alopecuroides 59 Lygodium venustum 60 Lygodium venustum LAURACEAE LENTIBULARIACEAE LYCOPODIACEAE LYGODIACEAE LYGODIACEAE

61 Herisanthia nemoralis 62 Herisanthia nemoralis 63 Pavonia humifusa 64 Pavonia malacophylla 65 Triumfetta semitriloba MALVACEAE MALVACEAE MALVACEAE MALVACEAE MALVACEAE Santana do Cariri, Ceará, Brazil 6 Plants of Palm Swamps in the South of Ceará Maria Daniely Freire Guerra1, Priscila Orlandini2, Rayane de Tasso Moreira Ribeiro3, Maria Iracema Bezerra de Loiola4 & Rubens Teixeira de Queiroz5 1Universidade Regional do Cariri, 2Universidade Estadual de Campinas, 3Universidade Federal Rural de Pernambuco, 4 5 Universidade Federal do Ceará & Universidade Federal da Paraíba Photos by R.T. de Queiroz, M.I.B. Loiola, and M.D.F. Guerra. Produced by M.D.F. Guerra, and Priscila Orlandini. For more information: Guerra et al. (2020). ©Field Museum (2021) CC BY-NC 4.0. Licensed works are free to use/share/remix with attribution, but commercial use of the original work is not permitted. [fieldguides.fieldmuseum.org] [1363] version 1 9/2021

66 Urena lobata 67 Urena lobata 68 Waltheria viscosissima 69 Clidemia biserrata 70 Pterolepis glomerata MALVACEAE MALVACEAE MALVACEAE MELASTOMATACEAE MELASTOMATACEAE

71 Sauvagesia erecta 72 Ludwigia erecta 73 Ludwigia nervosa 74 Ludwigia nervosa 75 Piper aduncum OCHNACEAE ONAGRACEAE ONAGRACEAE ONAGRACEAE PIPERACEAE

76 Angelonia biflora 77 Angelonia pubescens 78 Anthephora hermaphrodita 79 Melinis minutiflora 80 Bredemeyera floribunda PLANTAGINACEAE PLANTAGINACEAE POACEAE POLYGALACEAE Santana do Cariri, Ceará, Brazil 7 Plants of Palm Swamps in the South of Ceará Maria Daniely Freire Guerra1, Priscila Orlandini2, Rayane de Tasso Moreira Ribeiro3, Maria Iracema Bezerra de Loiola4 & Rubens Teixeira de Queiroz5 1Universidade Regional do Cariri, 2Universidade Estadual de Campinas, 3Universidade Federal Rural de Pernambuco, 4 5 Universidade Federal do Ceará & Universidade Federal da Paraíba Photos by R.T. de Queiroz, M.I.B. Loiola, and M.D.F. Guerra. Produced by M.D.F. Guerra, and Priscila Orlandini. For more information: Guerra et al. (2020). ©Field Museum (2021) CC BY-NC 4.0. Licensed works are free to use/share/remix with attribution, but commercial use of the original work is not permitted. [fieldguides.fieldmuseum.org] [1363] version 1 9/2021

81 Xylosma ciliatifolia 82 Paullinia pinnata 83 Schwenckia mollissima 84 Thelypteris interrupta 85 Thelypteris serrata SALICACEAE SAPINDACEAE SOLANACEAE THELYPTERIDACEAE THELYPTERIDACEAE

86 Thelypteris serrata 87 Piriqueta sidifolia 88 Piriqueta sidifolia 89 Cecropia pachystachya 90 Xyris macrocephala THELYPTERIDACEAE TURNERACEAE TURNERACEAE URTICACEAE XYRIDACEAE Santana do Cariri, Ceará, Brazil 8 Plants of Palm Swamps in the South of Ceará Maria Daniely Freire Guerra1, Priscila Orlandini2, Rayane de Tasso Moreira Ribeiro3, Maria Iracema Bezerra de Loiola4 & Rubens Teixeira de Queiroz5 1Universidade Regional do Cariri, 2Universidade Estadual de Campinas, 3Universidade Federal Rural de Pernambuco, 4 5 Universidade Federal do Ceará & Universidade Federal da Paraíba Photos by R.T. de Queiroz, M.I.B. Loiola, and M.D.F. Guerra. Produced by M.D.F. Guerra, and Priscila Orlandini. For more information: Guerra et al. (2020). ©Field Museum (2021) CC BY-NC 4.0. Licensed works are free to use/share/remix with attribution, but commercial use of the original work is not permitted. [fieldguides.fieldmuseum.org] [1363] version 1 9/2021

Location of isolated palm swamp in selective points of the slope and foothills of the Araripe Plateau

Physiognomy of the palm swamp of the Vale do Buriti, township Santana do Cariri, Ceará, Brazil