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New Generic Concepts in the Triticeae of the Intermountain Region: Key and Comments

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New Generic Concepts in the Triticeae of the Intermountain Region: Key and Comments Great Basin Naturalist Volume 43 Number 4 Article 5 10-31-1983 New generic concepts in the Triticeae of the Intermountain Region: key and comments Mary E. Barkworth Utah State University Douglas R. Dewey Crops Research Laboratory, U.S. Department of Agriculture, Logan, Utah Riley J. Atkins Utah State University Follow this and additional works at: https://scholarsarchive.byu.edu/gbn Recommended Citation Barkworth, Mary E.; Dewey, Douglas R.; and Atkins, Riley J. (1983) "New generic concepts in the Triticeae of the Intermountain Region: key and comments," Great Basin Naturalist: Vol. 43 : No. 4 , Article 5. Available at: https://scholarsarchive.byu.edu/gbn/vol43/iss4/5 This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. NEW GENERIC CONCEPTS IN THE TRITICEAE OF THE INTERMOUNTAIN REGION: KEY AND COMMENTS Mary E. Barkvvorth', R. Riley Atkins' Douglas Dewey% and J. Abstract.— Revision of the perennial genera of North American Triticeae requires recognition of seven new gen- era in the Intermonntain Region: Eltjtrigia, Letjmiis, Psatlnjwstachys, Pseudoroegneria, Thinopijrum, and the hybrid genera X Ehjleijmus and X Pseudelymus. One previously recognized genus, Sitanion, is included in Ehjmiis. Several new combinations are presented to accommodate the taxonomic changes. Ehjmits tmcln/caulus is treated as a wide- spread, polymorphic species with three subspecies in the region: subsp. trachycauhts, subsecundus, and latigliimis. Agropyron dd.systuchyum and A. albicans are treated as conspecific subspecies of Elymus lanceolatus. A key to the genera of the Triticeae occurring in the Intermonntain Region is presented as well as keys and brief descriptions for those genera not included in, or substantially modified from, other regional treatments. Agrostologists have been aware for a long ences between the treatment presented here time that traditional North American treat- and that found in their article. ments of the Triticeae (e.g., A. Hitchcock 1951, Gould 1968, C. Hitchcock 1969, Hol- mgren and Holmgren 1977) do not reflect the Taxonomic Treatment evolutionary relationships within the tribe. Nevertheless, in the absence of any well- The genera that are most affected by the documented revision that included a high revised generic boundaries are Agropyron proportion of the North American species, Gaertn., Elymus L., and Sitanion Raf. Agro- most North American taxonomists have pyron has been restricted to the crested adopted A. Hitchcock's (1951) treatment, wheatgrasses, the remaining species being as- with relatively minor modifications, as the signed to Elymus, Elytrigia, Pseudoroeg- best available. Recently, however, Dewey neria, or Thinopyrum. Several species of (1982, 1983a, 1983b) has published a revision Elymus have been placed in the segregate of the perennial genera that better reflects genus Leymus Hochst., but all species of Sita- the genomic and ecological data available nion are now included in Elymus. The rea- and is consistent with the morphological sons for these and other changes are given in data. Although written in terms of North the discussion of individual genera. To assist American taxa, Dewey's treatment is based those not familiar with the subtribal classifi- on data from the full geographic and tax- cation of the Triticeae, the genera and spe- onomic range of the tribe. cies within genera are treated alphabetically This paper is designed to assist those who after the generic key. wish to use Dewey's generic concepts for The intergeneric hybrids are treated after plants from the Intermonntain Region. It in- the nonhybrid genera. Readers are advised cludes a key to the genera of the tribe in the that such hybrids are relatively rare in na- region and, for those genera not included in, ture. We include them because they do exist or substantially modified from Holmgren and but, in our experience, most plants thought to Holmgren (1977), brief generic descriptions be hybrids are aberrant forms of good spe- and keys to the infrageneric taxa that we rec- cies. Interspecific hybrids are more common, ognize. Readers are referred to the Holm- particularly in disturbed areas. Part of the grens' article for illustrations, detailed de- problem in identifying hybrids in the Tri- scriptions of the species, and the complete ticeae, particularly interspecific hybrids, is synonymy. Table 1 summarizes the differ- that most can backcross to their parents. 'Department of Biology. UMC 45, Utah State University, Logan, Utah 84.322. -Crops Research Laboratory, UMC 63, U.S. Department of .\griculture. Logan, Utah 84322. 561 562 Great Basin Naturalist Vol. 43, No. 4 Table 1. Synopsis of our revised treatment of the Trititeae occurring in the Interniountain Region compared with that presented in the Intermountain Flora (Hohngren and Hohngren 1977). Revised treatment Intermountain Flora Aegilops cylindrica L. Aegilops cylindrica L. Agropyron cristatwn L. Agropyron cristatum L. clesertortim A. Shultes included in A. cristatum L. Agropyron J. Agrpyron fragile Roth included in A. cristatum L. X Elyhordeiiin macounii (Vasey) Barkworth & D. R. X Agrohordeum macounii (Vasey) Lepage Dewey X Ehjleymus aristatits (Merrill) Barkworth & D. R. X Elysitanion aristatum (Merril) Bowden Dewey Elymus canadensis L. Elymus canadensis L. Elymiis elymoides (Raf.) Sweezy Sitanion hystrix (Nutt.) J. G. Smith Elymus glaucus Buckley Elymus glaucus Buckley Elymus X lumsenii Scribner, pro sp. X Elysitanion hansenii (Scribner) Bowden Smith) Gould Elymus lanceolatus (Scribner & J. G. subsp. lanceolatus Agropyron dasystachyum (Hooker) Scribner var. dasystachyum (Scribner Var. riparium & J. G. Smith) Bowden Smith) Barkworth Agropyron albicans Scribner & G. Smith subsp. albicans (Scribner & J. G. J. & D. R. Dewev var. albicans G. A. A. Beetle var. griffithsii (Scribner & J. Smith) Smith) E. Sitanion jubatum G. Smith Elymus multisetiis (J. G. M. Jones J. Agropyron X pseudorepens Scribner & G. Smith Elymus X pseudorepens (Scribner & J. G. Smith) J. Barkworth & D. R. Dewey Elymus X saundersii Vasey X Agrositanion saundersii (Vasey) Bowden Elymus trachycaulus (Link) Gould ex Shinners Agropyron trachycaulum (Link) Malte subsp. trachycaulus var. trachycaulum subsp. subsecundus (Link) Barkworth & D. R. var. glaucum (Pease & Moore) Malte Dewey, pro parte Smith) Barkworth var. latiglumis (Scribner & G. Smith) A. A. Beetle subsp. latiglumis (Scribner & J. G. J. & D. R. Dewey subsp. subsecundus (Link) Barkworth & D. R. var. unilaterale (Cassidy) Malte Dewey, pro parte Elymus virginicus var. submuticus Hooker Elymus virginicus var. submuticus Hooker Elytrigia intermedia (Host) Nevski Agropyron intermedium (Host) Beauv. subsp. intermedia var. intermedium subsp. barbulata (Schur) A. Love var. trichophora (Link) Halac Elytrigia repens (L.) Nevski Agropyron repens (L.) Beau--'. Elytrigia spicata (Pur.sh) D. R. Dewey Agropyron spicatum Pursh Eremopyrum triticeum (Gaertner) Nevski Eremopyrum triticeum (Gaertner) Nevski Hordeum brachyantherum Nevski Hordeum brachyantherum Nevski Hordeum depressum (Scribner & Smith) Rydb. Hordeum depressum (Scribner & Smith) Rydb. Hordeum marinum Hudson subsp. gussonianuni (Pari.) Thell. Hordeum geniculatum All. Hordeum murinum L. subsp. murinum Hordeum murinum L. subsp. glaucum (Steudel) Tsvelev Hordeum glaucum Steudel .subsp. leporinuni (Link) Arcang. Hordeum leporinum Link Hordeum pusillum Nutt. Hordeum pusdlum Nutt. Hordeum vulgare L. Hordeum vulgare L. Leymus cinereus (Scribner & Merrill) A. Love Elymus cinereus (Scribner & Merrill) Leymus flavescens (Scribner & Smith) Pilge Elymus flavescens Scribner & Smith Leymus salinus (M. E. Jones) A. Love subsp. salinus Elymus salina M. E. Jones R. var. salmonis C. L. Hitchc. subsp. stdnionis (C. L. Hitchc.) J. Atkins Elyvtus ambiguus Leymus simplex (Scribner & Williams) D. R. Dewey Elymus simplex Scribner & Williams Leymus triticoides (Buckley) Pilger Elymus triticoides Buckley Pascopyrum smithii (Rydb.) A. Love Agropyron smithii Rydb. October 1983 Barkworth et al.: Intermountain Triticeae 563 Table 1 continued. Revised treatment Intermountain Flora Psetidoroegneria spicata (Pursh) A. Love Agwpyron spicatum Pursh X Psi'udehjnnis saxkolus (Scribner Smith) (Scribner Smith) & J. C X Agrositanion saxicohi & ]. G. Barkwortli & D. R. Dewey Bowden Psdthtjrostaclnjs jtincea (Fischer) Nevski Elymus juncetis Fischer Secale cereale L. Secale cereale L. Taeniatheriim caput-medusae (L.) Nevski Taeniatheriim capitt-mediisae (L.) Nevski Thinopyrum ponticum (Podp.) Barkworth Agropyron elongatum (Host) Beauv. & D. R. Dewey Triticwn aestivttm L. Triticwn aestivitm L. resulting in a morphological continuum as is exacerbated by the relatively small number well as partial restoration of fertility. This of diagnostic characters available for these problem is not unique to the Triticeae but it grasses. Key to the Genera 1. Spikelets three at a node, each with only one floret; only the central floret fertile (exc. in Hordeum vulgare in which all three florets are fertile) Hordeum — Spikelets not three at a node OR with more than one floret per spikelet (n.b watch out for some forms of Elymus elymoides [ = Sitanion hystrix] in which the second floret of the central spikelet is reduced and the lateral spikelets have only a single, sterile, floret) 2 2(1).
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