a

'paUrdllolral se aolDJuqLuluotl.as eqlJo e,\lllluaseJdeJelos aql eq ol leprsuol I tlxtq \'Dsoalt2 Dtuolotr epnlourlou prp srs,(leuelteql ,Jela,lroq,spuodserroc fle8rul eJoqpetdeooe,{uIouoxpt eqlqrrq,r o}snueBeqljo 1ueru1ee4crlsrpeloe pejedold (rg6 J) Trla uolseA\

'suortJas JeqloaqlJo sercods.setels tuolsee eq1 Surst,lar osle sr olleJncooJ€W (/66I)ouornCFquorluBqse,rurrapunsruortcosls€lstrlJ.asuespaoJqoqlutarJir^1121 'lces o1pue 'sdnor8 ,suorljos lpaJl s,.te18ug.ro s,lupqlueg o1 ,{lasrca;dspuodser.roc qcrq,r Jo euou rno; olur snue8eq1 epr,rrp ol posodordetoq sr tI seuese^g olul pepr^rps€,r qctq^\Joeuo ,suotlces eerql otur(9681) relSuE fqpue serresue^es opr (!9g l ) rupqluegfclpepr,rrp su,,n arao.iog snue8 eq1

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uo!lJnporlul

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lJurlsqv

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uoslrld c Inl?d

sJolJuJBqJpaas uo salouqfrAr.arlsrlsnv uJefsea\ rrfolJ (a8or€lnu) otuo,tog snua8oql ut BxEl ,!!,aupuB srruBu ,{toN

(866r)ts r-6 r r:(r)zr 124 NuytsiaVol. 12,No. 1 (1998)

The probability of the being polyphyletic without the inclusion of Bororel/a from New Caledoniawas indicatedby Westonel a/. ( 1984),and an investigationby me ofpetal venationand seed morphology, characterswhich werenot includedin that study,could strengthenthe proposal However, Hartley (1995) doubtsthis closerelationship and supportsthe argumentfor retaining ,oronella as a distinct genus with details from a study of the articulation in the branchletsand of the shapeof the embryonic cotyledons.Ifthe closerelationship ofBororla to Boronellaweretobe accaptedthenfloral and seedcharacters suggest that the latter is more closely relatedto sect.Boloria than that seationis to anyofthe other sectionsin the genus.Logically therefore Boronella shortldeither be madea section of Boronia or the sectionsofBororla shouldbe raisedto genericlevel. In spite ofthese lese ations "Flora Duretto and I have decided for the of Australia" to retain the genus with its traditional circumscriDtion,while BoronellQis assumedto be genedcallydistinct,

As is noted above,the recognitionofthe infragenerictaxa within Boronia,while basedlargely on floral morphology, is strongly suppofiedby seedmorphology The opportunity is thereforetaken to describethe seedtypes that have beenobserved in representativetaxa of Westem Australian of Boronia.

Seedterminology in the Boronieae sclerotesta. The hard, thick, and brittle portion ofthe testa. lt is always black in colour and is either smooth or variably corrugate.

Outer testa. That portion ofthe testaoutside the sclerotesta.It is membranous,papery, coriaceous. or brittle. It may be smooth, or it may follow corrugationsplesent on the sclelotesta,oI it may have conugations that are independentof the sclerotesta.

Micropyle. This is minute and circular or rarely narrowly elliptic. It is alwayspositioned ontheuppel adarial margin ofthe seed.

Hilum. T\e scarformed by the dehiscenceofthefunicle oraril'. Itis alwaysinthe centraladaxial alea ofthe seedand it variesin shapefrom linearto circular. In Boronie sect.Bororra it is sunkenbetween lateral lip-like raisedportions ofthe outer integumentand is herereferred to as labiose.

Raphe. The v asc,,rlarstrand and associatedtissue that passfrom the baseofthe hilum to the chalaza. It varies from being shrivelledand insignificantto large and fleshy

Raphecover. The tissuethat folms a coverto the raphe. It is continuouswith and usually hasthe same texture as the outer testaand tltereforevaries from being membranousto brittle

Chalazalaperture. The circular hole in the sclerotestathroughwhich the raphepasses into the chalaza. It is positionedbeneath the raphe,either at the baseofthe seedor at its lower adaxialmargin.

Hilqr strancls.In somegenerathe seed possesses a cream-coloured ligament-like tissue on its adaxial surface.This tissuehas the appearanceofa strand;it surroundsthe hilum andextends, as a singlethread, to the raDhe.The strandis herecalled the hilar strand. Itis shedon orshofily afterthe dischargeofthe seed.Hilar strandsare found in the seedsofChorilaenq, P hebqliumsect. Gonioclados, Rhadinotharnnus, a

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BruoJog.J3seruoJog.lces uls uruolog .sl

Pruorog.lf,es Lus Eruotog .l

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'pJJS JlltLuo.U (une s;1e3 (11cn:u:re pue .Etset €lsot eqt tuog peletedesseuroceq 1r erntrEru,{l.reou ueq,41 JelnoeqlJo lset eq1o1pesrg .{1e1uu.u1ul sr pulrls eJnlnl eqt lueudole,tep peesJo sa8elsfpua aqt uy unltq eql spunoJ.lns1eq1 (luarunBelur Jolno eql ulo.+ perujoJ)plsel lolno eql uoqrod arorrEu€ ruolJ s3suupuells rulrq eql sruedde 'eearuorog lo 1uq1 lI eqlJo sJequletuJorllo ur lueu_rdola,repjo se8elssnorru,l 1e spees q1r,n rueql ?uuudruoc ,{qpuesxe}eserllurseln^oJoluoudole^epaqturse8e}sJo,{pnlseuorg.altotl

prle'rlsnv uritsor\\ r|orJ Diuarag ,uoslrr^A snuiJ8oqr ur rxrt ,rou pUD s.Lulu ^\oN D lnrd Nuylsiavol. 12,No. 1 (1998)

Boroniqsect.Octqrrlrera F. Muell., Pl. Indig. Col. Victoda 1: I l3 (1862) Type:Boronia PinnatdSm., lectotype here chosen.

Boroniqser.P innqtaeBenth., Fl. Austral. 1: 309,307 (1863). - Boroniqsect.Pinnatae(Benth.) De Wild., Icon.Select. 2:67 (1901).Type; BoroniapinnqtaSm.

Borctniqser.Terminales Benth., Fl. Austral. 1:3 10,323(1863). Type:B. capitataBentl.I.,lectotype llere chosen.

Boroniaser.VariabilesBenth.,F. Austral. 1: 309,320(1863). Type: B. crenulatasm.,lectotypehere chosen.

Boroniaser.Ovatae Paul G. Wilson,Nuytsia1:204 (1971). Type: B. ovqtaL\\\dl.

Seedmorphologt. Seed kidney-shaped,adaxial margin flat or convex. Tetla smooth; outer testa aoriaceous,matt, black. Hllzm sunken,linear to elliptic in centreofadaxial margin, surounded by glossy labiose (lip-like) raised margin which surounds raphe. Raphe fleshy. Chalazal opening basal. Placental endocarpmembmnous, caducous (Figure 1)

Speciesexamined.Ofthe23 Westem Australian species recognized inthis seriesthefollowingninewere examined for seed characters.Boronia clavttal, B. crenulattt, B humifusa, B inornala, B. ovqlq, B. purdieana, B. scabrq, B. spalhulqt.t, andB. tetr.tndrq,

Fig$e L Barcnia crcnuldta s|hsp. r,i ined sceil. A - Iateral vicw, B adaxial surf-ace,C - longitudinal radial section, all \20. Drawn ttont P. lulJ. I Ocr la6i suecseqnddsqns " asolrdstalqouelq :et€rlra-8uol 'eseq 'e1e,roqo ,tlesJuds lu otuproJsaurloruos o1cr1dr11e flpeolq sa^pe-l uaurrur,ldsqns elelorpc,{1e1nutru ro eletltce 'elelncrde '3uo1 'J?ln6ueul ,{ll?rrduqnslou'o1ncz o1 esnlqo rutuZ-E l o1e1e,ro sledeg :g esnlqo dsqns '''''''''''''''''''''olelorlrc'e1a1ncrde,(llecrdeqns 'xede '3uo1 'e1u,r.o {1o1nu[u 1apepunoJ Jo osnlqo urur 7 r ,{lpeorqs1ude5 g etclorlrceJo elulorIrc ,{latnulul 'e1nce ro esnlqosFdes :os FlelnurorJo elulncrdu1ou reqlre sqele4 :7 alslnu?rJ dsqns eleyorlrcflesuap 'unlnctde qlt,lr osnlqo ro eluururncusledes ios ,{ltuouruoldfllensn'alulncrdz fllzcrdtsqns slelod Z snolnreqnd.to srrorqel6 'esEq 'elu,loqo stelqcuejq:elErTrce le pe,rorj€u (lpuorq ol reeurl sa^ee'l I ',,ro1aq papr,rordspeal eqt olur ,{lesrce;d19 tou op s1ueld,(uzru oJoJaJaq:} 'l?qto qcse olul epurSsarcadsqns eqt teql sr uorlecrldruocJeqtlU V ueJDetu€s eql tuo{ suorlcellocur 'elqerl€A ro tuald eul€saqt uo rro^e ,{Je,\sr exel cUrcedsul.+urlueroJ.}rp eqlJo rlruour edDqsJDeleqJ.

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'uesoqc '(86t8 'le4 '€rlur1snV oraqed,{1o1cel S0t rolcalost'.q1.o7cal gaOZSStatd 7

'uesoqc e;eq ed,t1o1cel '(98t8S0l 'lAtr^t 'arl€l1snv 'lfeg iorrape'.>1.opal)Llwo.rgy'[7991 . ;anue1] urolse/[ .{1cn1] 1,{ug :ad,{; ' 'qaeeq 'e8uer 'a8ue1 ,.ratxoU,punos at^xDn eu,{eqJ puE 3u1Fr1g;g 51 s1[q \tepJED s,eBJoeD 'U ']se .1u4snv.lC, 6ur>l:?.1,1.rol&r oc qlnog :uor\o|tc adr(1 (g9g1) 11g : 1 qlueBxa rg U groulqlB sruoJog

(C'CZ ernBrg) eceJI€rx€pu eql uo ra,loc (sso13e fq peleloc enssrlpuolq Dsr eqdul eql ellq,tluorls uruSesr ruqryu4osnfiwr1 gq acejlelrcp€ eqluo EerulezuluqJ,{ssolS eqt oguoute8tlp sp 'Uoqs sessedeqdzl eql :ueluns lou puucrldqle sr urnpq eql rlgt rs 6rul uorlgnlrslpnsn eql ot uorlggeA euos sreJerlJ punorrns,{sso13s1r puu urn1tqeql ol ur6;uruesorqpl aqUo ocuesardeqlsr eeeruo;og eql ur Ex"trerlloJospaesluo{uorlcasstqtJopeeseqlseqsrnBurlsrplEqlJalJE.reqcsnorAqolsoureqJ,ra,?oN

'uos|/A tzl urlDrlsrlv ur:ls.lA rLrcll Dluarag snuei arD ur Expt ^\. pue solusu I\aN D Ine(l t24 NuytsiaVol. 12,No. 1 (1998)

Boroniacrenulata Sm. subsp. crenulata

Distribution. Occurs jn the south-westof Westem Australia, towards the south coast from Augusta eastto Bremer Bay and nofth to the Stirling Rangeand from thereto the eastemmargin ofthe Darling Range.

Nore'.Two varietiesare recognized.

Leavesnarrowly to broadlyobovate...... var. crenulata Leaveslinear " var'angustifolia

Boronia crenulataSm. subsp. crenulata var. crenulata

B. yimineavar, gracilisBenth., Fl. Austral.1: 325 (1863).-B crenulatqvar' gracllls(Benth ) PaulG. Wilson,Nuytsia 1:204 (1971).Type: Western Australia, 1848,J. Drummond 92 (holo:K; i.io;TCD).

nanowly to broadly obovate,7-12 mm long, roundedto obtuse -Flowerspredominantly axillary with I or2 pairsofbracteoles. Sepafu ovate, acuminate,2-3 mm long. Pelal,rglabrous outside, minutelysubapically apiculate.

Dislribriion. Occurs over the entire range ofthe subspecies.

Noles. The variantfrom nearA lbanywith broadlyobovate leaves coffesponds to the typeof B. crenulata while the variant from near Bremer Bay witlt larrowly obovate leavescorresponds to the type of B. \)imined var. gracilis.

The variant found nofih-west ofthe stirling Rangeand towards the Darling Range is sometimes puberulousall over; it hasnarrow leavesand naffowly triangularto triangular . This valialt is somewhatintermediate in morphology betweensubsp crcnulatq ar\ds\bsp. vimineq.

Over most of its distribution the branchesof this vadety are puberulousbut in the Pembedon- Walpole areathey are pilose which suggestsintergmdation with subsp.prir5escens.

Boronia crenulatasubsp. crenulata var. angustifolia Paul G. Wilson,var. nov

Folialinearia, I 5-30 mm longa,acuta. Flores plerumque ad ramulis brevibus axi llaribus terminati. Sepalaanguste triangularia, acuminata, c. 3 mm longa.Petala glabra, manifeste subapicaliter apiculata.

7.Ip1aiNearjunction ofEllen and SouthBlufftracks, Stirling Range,Westenr Australia,26 September 1915.R.Voutier195 (ftolo; PERTH 00962651)

Leqveslinear,l5 -30mm long,acute. F/ou,ers mostly terminal to shortaxillary shoots SeP.l/,tnanow triangular, acuminate,c. 3 mm long. PelaL!glabrous, prominently subapicallyapiculate'

Selectedspecimensexa.ryrned WESTERNAUSTRALIA:Ellen Peak,2lOct.1902,A MotisonQERTH); Ellentrack, Stirling Range,24Oct. 1973,R. Vourierl95 (PERTH). '(gDJ:utrs) saFuDn7'(gga:uts) u spuoLuluu1 | 'urlErlsnv ',{uoloC ,.Iput-l .B uretseld re^rUuD.{\S :ad,t1 $gg1) / t U uE,rS3en qc1e1g oawwt^

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nrlcrtsnv .uosll/d 9Zr urolsarldLuou Diua.tog snue8 aql ur nxEl Mou pue s:rxuu .^,\aN D lnrd

i NuytsiaVol. 12,No. I (1998)

B. yinineavar.latifolla Benth.,Fl. Austral.1: 325 (1863). Type:Canning River, Pefth, Westem Australia, NovemberI 841,r. Preiss2022(lecto;LD; i'roleclo; MEL 1058527),lectotype here chosen.

B. tenuifoliaBartl.in Lehm.,Pl. Preiss l: 168( 1844).Tlpe CanningRiver, Pefih, Western Austalia, November 1841, Z. Preiss2022(lecto:LD isolecto:MEL 1058527),lectotype here chosen.

B. machardiqnaF.Muell., Fragm. g: 115(1875). Type BlackwoodRiver, Western Australia, 1874, M. McI'lard(iso ; PERTH 00997528,fragment)

Typification. Thetype ofB. mqchardiqnais of avariantthat is most similarto materialfoundj ust south ofpertb althoughit is saidto bavecome from theBlackwood River area which is in thefal south-west ofthe State.

Boronia humifusaPaul G. Wilson,s2 irov.

Ramuliacute -angulati,glabri ve I in lineisoppositis scabridi. Folia sessilia, simplicia, oblonga vel oblongo-elliptica,I -2 cm longa,integra, plana, obtusa, infra glabra, supra scabridula. Flores in cymas apeftaslpedunculus gracilis, c. I cm longus;pedicellus gracilis, 5-10 lnm longus,in mediobreviter bibracteolatus.Sepala deltata, c. 1.5 mm longa,glabra. Petala irnbricata, elliptica, 6-7 mm longa,extra glabra,intra sparse pilosa, rubra, ad apicem plana, non apiculata;nelvatumpil]nata, nervo medio unico. Stamina8, omnino fefiilia; filamentadense ciliata, ad apicem verucosa. Discusintegra, glabra. Ovarium glabrum;stylus stigmaque cylindracea, c 0 5 mm longa,glabra

1996,B l Lepschi& T R Lally2611 ryprs: Capelto Donnybrookroad, Westem Australia, l6 June (/rolo.PERTH 04334221;lso; CANB, K)

Low-growing wiryp erennial. stentsandbranches shatply 4-angled, glabrous or sometimesscablid in oppositelines. zeaves sessile, simple, oblongto oblong-elliptic, 1-2 cm long,entile, flat(sometimes recuwed when dry), obtuse,glabrous below, scabridulousab ove. Flowefs in opencymes terminal to main andlateml branches; slender, c. I cm long;pedicelsslender, 5- l0 mm long,medially shottly bibracteolate. sepalsdeltate, c. 1.5mm lol,1g,glabrous, glandular-punctate. imbrrcate,elliptic, 6-7mm long,wrinkledon marginrvhen dry, pilosewithintorvardsbase wjth simplehaim,glabrous outside, red,apex neither inflexed nor apiculate;venationpinnale with a singlelredial vein. Slaneirs8, all similar and fertile; filament compressed-terete,s-curved, dersely ciliate, apexglobular and veffucose;anther affixed subapicallytoadaxial margin, thecae c. I mm long,white apiculumc. 0.5nm long. Drsccushion- shaped,eutire, glabrous. Olary glabrotts;style and stigmatogether cylindrical, c. 0 5 mm long,glabrous' Seedsemicircu lar, c. 1 mm long; outer testasmooth, black; adaxialmargin with glossycover to raphe, without a groove. (Figure 2)

other specinensexan irerl. wESTERN AUSTRALIA: SSEofCapel on Donneybrookroad, R. Da'les 125(PERTFI); 16rd., B.J. Lepschi& T.R Lally 3 l2l (PERTH); ibid..D. Papenluss300 (PERTH); l0 km W ofDornybrook, P.A.Jurjev ich 1831(PERTH)iTutunup'Sep.7945 ' Fare;ter ll/erloz (PERTH)

Distribution. South-westWestern Australia, betweenCapel and Donnybrook

Habitclt. Growirgin Jarrah- Marri open foreston lateritic soil '9ZI sat^Do ./y urc{ u^\Drppaxs:tlgz [llDj y1.p ruo{ rr,,t\nrpsuoruod iurrc^\otjt (C pue '3'C) '(l'B) rya:;dal /g C ruu' I rulu''(V) nri elpos p.osJorrr.tlns -.J.paaslo r\:r^ -:l .lrtsrdpuc _ 'suetrrDls Z:slcq IDIx?pr lrl.lcl osrp !l curFladpuL' curledos - C'lttad - J.1.,(oLl - g.rl.uElq Bu l).{\oLl_V ptntunLl Dtu og.Zerr?tt

(.,\ lt

.uoslrlll EII?ISnV urats:)/A wolJ Dtuotog snu:8 aqt ur !xc1 ^.\oupuE s.urt ^.\.N O Ined 128 Nultsia Vol. 12.No 1 (i9q8)

Conseryation status. Known from one small areaofforest which iS subjectto mining and therefore probably placesthe speciesunder threat. A Priority One classificationmay be appropriate

Etymolog,,. The specific epithet,humiJusa, is aLatin word meaning spread-outover the ground and refersto the low-growing habit ofthe .

NoleJ. The relationshipsofthis speciesare uncertain. It doesnot agreewith othel membels of sect. Boronia sincethepetalslack an apiculumandhave only onemain v€in, while the seedlacks an adaxial groove. In typical membersofsect. Bororunthe petals have several medial nerves and have a subterminal abaxial apiculum while the seedhas a linear groove along the hilum. In inflorescencecharacters it resemblesB. sc4b,,aand B. ovatato which speciesit may possiblybe mostclosely Ielated and fol which reasonit is here included in the seriesBoronla

BoroniapurdieanaDiels,Bot.Jahrb.syst.35:318f.38b,c(1904).Type:NearBullsbrook,Western Australia, July, L D iels 3564 (holo. B n v., ?destroyed).

Distribution. This speciesis fou[d neartl]e west coastof WestemAustralia from Perthnorth to Shark Bay, with one collectionfrom nearLeonora, c. 130km noth ofKalgoorlie.

Notes. Boronia purdieana is vatiable in both vegetativeand floral morphology, however, in anther shapeand lengththereare clearlytwo groups. Thereis a short-antherplant, which is referedto subsp. purclieana,and a long-antherplant, which is refeffedto snbsp.calcicola. The plant that is found near Leonora has shorl anthersbut its habitat is very different flom that of the type locality of subsp. purdieana.

Petalineanthers + square,not curved,.. 0.4 mm long ...... subsp. purdieana Petalineanthers oblong, curved,c. 0.8 mm long . subsp.calcicola

BoroniapurdieanaDiels subsp. purdieana

Distribution. Foundfrom nearGnangara(c.20 kll north ofPerth)north to Coorowwith a 1901collection from near Busselton;recently a sizeablepopulation was discoverednear Leonora

Habitat. Sal(ly seasonallywaterlogged soil in Bair,tsrawoodland; however,the one collection from near Leonora was found growing on a shaleand diorite hillside.

Noles. Diels, in his original description,records that this plant was sold in Perthas an ornamentaldue to its fragrant flowers.

Boronia purdieanasubsp, calcicola Paul G. Wilson,sablp ilov.

A subsp.purdieanae antherispetalinis oblongis, incurvis, ad basim ampliatis, c. 0.8 mm longis recedit.

ryprs: BetweenPotAlley andBluffPoint, nearKalbani,Western Ausfalia, 1Augustl9']2'C Cockman (lolo: PERTH00978752Irso. K). '(StqgE gfUaa :o1ot7) ,0661 ,puerC t t O 1gg1&uo.t1sw7. f reqotrO ,,erluJlsnVuleNeld e.I l4l ..srd,{J

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'aoa dsgzs'uos[16C d slEnuallB dsqnsBlqBJs Bruolo{

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fi11X96) 1qg9nos7u4l) d.tspoc ruo4 ru) I peuJo (II{ lInlg S t:(Hfugd) 0081,?a1al J.}r,ujl?qlp):(HJUEd\ tgg6atuoag.g.y,peelsetuoq€puruJ 3o355[ru1y7e]sellur9t:(HJugd)tlIoaasnoqlxoggt6r,uort€lsesnoHuosrqcrnl:(HJuEd) {qqsy 'JJnlA LZqI t { f peuJo S pror tseol :VI.IV.UJSnV NUEJS3^| paurwDtz suawtcarlsaqtg

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6ZI €rls4snv u.r:tso^\ worJ Djuuog snuag .Lroslry'd eq] ur lr€l ^\au pue solu?u .\\cN I Inrd

\ 130 Nu)tsir Vol. 12, No I (lqq8)

Zeavesnarrowly eiliplic to terete due to tlte recurvedto revolute margins. Flolters congestedat branch apicesor on slenderpeduncles and pedicels, 4- or 5-merous.Sepals narrowly triangular,long- attenuate,c. 6 mm long, strongly hilsute. Pelals c. 8 mm long. Staminal.filamentssmooth except for tuberculateswollen apex.

Selectedspecinensexanfued WESTERNAUSTRALIA: HellflreBay,M. Catter225(PERTH);Cape Le Grand,,J.C. Mqlone 5 (PERTH); SandyHook Island,Recherche Archipelago, 10Nov.1950,J.C. lt/i/lis @ERrrD.

Distibution. Found on the southcoast of WesternAustralia at CapeLe Gralrdand on the neighbouring islands of the RechercheArchipelago.

Hqbitqt. Growing amonggranite rocks.

Conseryation stcttus.Altholrgh this subspeciesis fbund in only a small areait doesoccur in a national park and thereforeis not under threat.A Priority Two classificationwould be appropriate.

Etynologt. The subspecificepithet alludesto the attenuate(Latitl attenuatus)shape ofthe sepals.

Noler. At CapeLe Grandand on someofthe islandsofthe RechercheArchipelago is found the variant with congestedinflorescences while at Thistle Cove,c. 6 krn eastofCape Le Grand,and in CapeArid National Park is found a variant with slenderpeduncles and pedicels.

This subspeciesis of particular interestbecause it liequently has ahernateleaves and 5-merous flowers;for thesereasons it hassomgtimes been assumed to be a speciesof

Boronia scabrasubsp. condensata Paul G. Wilson,r'aD.rp. rov.

Foliaelliptica, plana. Flores iu axillisfoliorum tem inaliumaut bracteis foliaceis breviter pedicellata. Sepalaanguste elliptica vel angustetriangularia et acuminata,4-6 mm longa,glabra vel sparsepilosa. Petalac. 8 mm longa.Filamenta staminalia glanduloso tuberculata.

Iypzs: DinnerHill, WestemAustralia,26 August 1969, K Nevtbey2861(.hoio:PERTH 00986747).

leayes elliptic, flat. F/owersshortly pedicellate in axilsofterminal leavesor -likebracts. Sep.r/,t narrowlyelliptic or narrowlytriangular and acuminate, 4-6 mm long,often open at base,glabrous or sparselypi lose. Pelals c. 8 mm long. Staminalfilaments glandolartubercu late.

Otherspecinens exarrfued. WESTERN AUSTRALIA:Alexander MorisonNationalPark, E.A.Grffin 4725(PDRTI-I);Coomallo Cleek, G.J Ketglery3199(PERTH).

Specimensexamined of snbsp.condensata- subsp.scctbrul intelmediates. WESTERN AUSTRALIA: 15miles [24 knr] E from Mt Peron,C '4. Gardner 9426 (PERTH); I 5 miles [24 km] N of Badgingarra, A.S.George 8621 (PERTH);Mt LesueulReserve,E.A. GrilJin 4169 (PERTH).

Di:ilribution. Foundin the Badgingaraarea ofsouth-west Westem Australia. 9961 .ae lallopu q? tuorJ u,\{nrc ozx lle'uoqres l€rpErl?urpntr8uol - J'o.r.trns l?rxtp! - g',rel^ IErolsl-V peasD tototplp oluara| Eel|o?tl

(€ ern?rC) snocnpDc 'snouurquoru dlocopua y4uaco14 eqdul ,{q paleloc lasuq Suruadop:tzo1ot13 poesJo eseq le ',{ssol8'snoeceuoc sseu,{d1ndu,to:q rludolunene aqdog ur8.rcrulerxe pu 3uole tgeLqwnf H {c€lq 'pedrqs-,(eupry 'tSoloqdntu elset Jelno :qloorus r2l,ral xo^uoc Jo 1zg ur8:uru lerxepe paag paag

'(I/6 t) toz:l PrsfnNruosl/d D d ap{edK1olce1'lpurlDtDryqrods g:adtrJ (t9gl)9Zt:l ls4snv lC. qluegaotofirunpad.tes ouo.tog

aElelnJunpadJes sruoJofl 1c0sBluolog .qI

'e^oq€pelrr eft suorlcalloc al"Ipeuratur oruos sercadsqnsyucrd,{l eqt otorseplxS puu ,{Soloqdrouur elquuuAst uoxq srqJ .saloN

'asuecseJouur eqlJo ernleu (.n?,lrrsuapuocw1e1) ltedatoc eql ol ecualeJeJseq leqlrde cgrcedsqnset11 IBolowtlg

'aier.rdo.rdde eq plno,r uotlcogtssElco,^\J ,&rJorJdg uorlcelord e.llnbar1ou seop eloJoloqt pue ryud luuotluu 'Js^e,rorl 'suorlJelloc u ulqlr,{\ punoJsr lr ,nej e,(1uoruo.rg u,roDl sr uoxpl slllJ .snpls uollol,/asuoJ

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'.jlolJ 'D ?rlDrlsnv urolse/A Diuuog snur8 aqt ur srul Mou pun seuEu.{\.N.uoslrr\\ lritd NuytsirVol 12,No. I (19o8)

Speciesexamined.Ofthe sevenWestern Australian speciesrecognized in this seriesthe following six were examined for seed characters'.Boroniq anceps,B. denticulata, B. dichotomq, B../aiigilta, B. juncea, and B. spqthulata.

Noles. The principalcharacter ofnote isthepulpyrapheat the baseofthe seed.This fomation has not beenobserved in any othersection. The seedm orphologyotlterwise appears to gradeinto that found in typical membersof sect.Boroniq.

Boronia ancepsPaul G. Wilson,sp. nor,.

Herbaperennis, glabra, ad 60 cm alta. Rami applanati,acute 2-angulati. Folia internodiisbreviora, angusteellipticavel lateobovata,2-4 cm longa,multo reductasuperiom,integra, sessilia, adbasin latiora. Floresin cymis longepedunculati dispositi; pedicelli 5- l0 mm longi,graciles, in mediobibracteolati. Sepalaovata, acuta, 3 mm longa,glabm, decidua. Petala late elliptica, c 10mm longa,pallidorosea, ciliata; antheraeminuteapiculatae. Discum integrum, angustum, glabrum Ovariumlanosum;styluspyramidalis vel conicus,c. 0.5mm longus,glaber; minutum.

Zlprs.5kmnorthofBrennansFord(ScottRivercrossing),north-eastofAugusta,WestemAustralia, freshwatermarsh,l6November 1982,,4. .9trid 21427 (holo. PERTH 00946273).

Perennial herb to 60 cm high, glabrous,not foming a lignotuber. Slem'!flattened and ancipitous when young, jntemodesc. 6 cm long. Zower/eares sessile, obovate, c 2 cm long;upperleaves narrowly oblong to narrowly ovate;uppermost narrowly oblongto linear,usually very reduced. Inflorescence a terminal pedunculateayme; peduncleslender, to 6 cm long, bractscaducous; pedicels slender and smooth,5-10 mm long. Sepalsimbricate, broadly oYatetonarowly triangular,2-4mm long, glabrous or woolly ciliate, dark red. Peralr imbricate,broadly ovate,c. 8 mm long, acuminate,glabrous, pink. Stamensall feftile sepalinefilanents very narrowly oblong with a clavateapex, vefiucose abaxially, ciliate below; petalinefilaments similarbut slightly shorter;anthe$ affixed subapicallyon a very shot't stipetoadaxialside olfilament,c. 1.8 mm longwith a smallroundedwhite apiculum Dricc. I mm high, equal in width to ovary, glabrous. Carpels glabrous or shortly woolly pilose; style and stigma continuous,columnar or conical,1-1.5 mm long,glabrous or sparselypilose at base. Coccloblong, c. 3.5mm long. Seedsoblong-ellipsoid, c.2.5 mm long;outer testa smooth, glossyblack (Fjgure4)

Selectedspecimens examined. WESTERN AUSTRALIA: Boggy Lake, 27 Dec. 1957,D Churchill (PERTH);Scott Riv erroad,S. Pat$t265 (PERTH);ScottNationalPark, Cl RoDlnson505 (PERTH).

Distribution and halrltat. Found in the extremesouth-west of WesternAustralia betweenSoott River and Walpole. Growsin seasonallyswampy heaths.

ConserNationstatus. Although this speciesis known from a relativelysmall area some ofthe populations are in a national park. A Priority Thr€e code is thereforeprobably appropriate.

Etynology. The Latin word aircepJmeans two-edged and in this caserefers to the shapeofthe stem

Notes.Baronia qncepsis evidently closelyrelated to B./.rsligiatq andto B. spathalala; it differs from both in having flattened ancipitousupper intemodesand apparentlyir lacking a lignotuber. a

pllts y '.rrou 'S0 .(f ,(C) ,(J) .(C LZI\Z pxrs E ,os rgoy f , uort u^\erpsuori?.Itsnllr Euuo'{\otC H) rulu I rutu f ruru0l pus C'A'g)uru Z'(V) uc S :sleqelscs p.osJoorp.trns lsrr.Ep€ - l,pocsJo,rol^ Irralrl - H lusrupuE 3srp - rJ,suclusls - - - .p ,urpledpu0.urlEdos J'loled g 1a^\oll C'ocue.s.rouur- J.ruols S I- fl lulld.loLl^\- V sdacuottuotog anfug

dlm

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I 13,1 Nu_vtsiaVol. 12,No. I (1998)

BoroniadenticulataSm.,Trans.Linn.soc.London8:284(1807).Type:KingGeorgeSound,Western Australia,1803, A. Menzies(holo.LlNN, photoseen).

"In B. chironiiJbliaBaftl.in Lehrr.,Pl. Preiss. 1:167 ( 1844). Type: uliginosissylvae 17 mill. aburbicula (lecto Albany (Plantagenet)"[Albany, WesternAustralia], 9 November 1840' L. Preiss 202'7. LD' isolec t o ; MEL 1058492),lectotype here chosen.

? B. hypericifoliaRegel,Gartenli. 6: 152(1851), ex ctesc.

Note,. Regardingthe date ofcollection ofthe Menziesspecimen aited above see nole\\nder B. crenulqtd.

Boronia dichotomaLindl., Edwards's Bot. Reg. 27: subtab. 47 ( 1841). Type VasseRiver, Western Australia,Mr Molloy (rolor CGE, photo seen)

B. spathulatavar. elallor Benth.,Fl. Austral. 1: 327 ( I 863). Typecitation; B. dichoforra,Lirdl. Bot. Reg. 1841,under n. 47. - Vasscriver, Mrs. Mol/oy; Swan River, Drummond,Coll l843,n 38 f1'perVasse River, Westem Australia, Mrs Molloy ('ryir''K, photo seen); Swan River Colony, WesternAustralia, J. D r ummo n d 3 8 (syr. PERTFI).

"ln B..flexuos aBartl. inLehm., Pl. Preiss. I : I 66 (1844). Type arenosissylvae ad fluvium Canning(Pefih) (lecto: et in districtuSussex" fCanning River, Perth, Western Austlalia],5 Decembel I 839,L. Preiss 2024 LD; isolecto:MEL1058504, MEL 1058505),lectotypehere chosen.

B. integrifol iaDomin,Vestn. I(ral. CeskeSpolecn. N auk,Tr. Mat -Prjr.2: 53 ( 1923) Typecitalion:W 'A': coll. A.A. Dorrien-Smiih(herb. Kew). Z}pe: Bridgetownto Kojonupand SlabHut Gully, Western Australia, 19lO,A.A. Dorrien-Snith (lecto: K) lectotypehere chosen

Notes.Boronia dicholona is readily recognizedby the glandularaculeate proceSses that are found on the pedicelsand to a lesserextent the pedunclesand terminal portions ofthe stems. The leavesvary from being obovate (c. 2 cm long) towardsthe baseofthe stemto slendel-tefete(c. 4 cm long) belorv the inflorescence.The stems areslender and the internodesvely long,the upperones up to 12 cm long.

The type of B. integrifdia differs from material otherwise included in B dichotoma in having relativelysrnall glandular aculeate processes; it also differs in all theleaves being elliptic and 6- 12 mm longwith the upperinternodes only slightlyexceeding them. The district in whichthe type is statedto havebeen collected is 50- 100km eastofany otherrecord for B. dichotoma.Boron i.t integri-foliqcould be recognizedas a distinct taxon, but it is possiblethat the type is a hybrid betweenB. dicholom.l and ser' Pedunculatae. B.Jastigiatd or betwee.I.B. dichotona and anothermember of Boronia

Boronia exilisPaul G. Wilson,lp. ,lov.

B. junceae subsp.laniflorae (Bar1l.) Paul G. Wilson affinis, a qua imprimis differl pedicellis longioribus,staminibus piJis rigidis manifeste ciliatis

7yl?1j: ScottNational Park,Western Australia, 17 Septembet1990' C.J Robin;on 127(holo PERTH 3054810;lso.CANB). 6t0 tlorutqol fJ tuo{ u'\\prc (A) rlru I'(C) Lurul0'(J'{) LU||rC'(y) urlu 0Z:sl?q ol€.S 'slqsrd - 'suolutts - 'urats fue.srp il lurpl.d pur.urpdis C'lr,\\ou - J'pnq - fl Buu:^\ol+- V sttrcaDiuarag S :,t13t!

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.uoslrlt\ Erlerlsrlv ur3rs.r1ALuorJ Dtuarag snuoi .!n ul €xrl ^\au puD sirusu ^\!N D Ined 136 NuytsiaVol. 12,No. I (1998)

Distribution. Known only from the Scott River area,extreme south-west of WesternAustralia

Habitqt. Growirlgin seasonallywet heath.

Conservdtionstcttus. Classified as DeclaredRare Flora, it is only known from a few sitesover a small area.

Etymolog). The Latin word exilts,meaning slender andweak, refers to the slenderstems ofthis species.

Notes. This speciesis similar to Boroniq juncea, particularly to the subsp.laniflora; it differs most obviously from that speciesinhaving stronglyciliate staminalfilaments (inB.jarceathey areglabrous).

"In Boronia fastigiataBartl. in Lehm.,Pl. Preiss. I : 167( 1844).Zype; locish iemeinundatis districtus Plantagenet" lAlbany area], Western Australia, November 1840, L Preiss 2028 (lecto LD;. isolecto: MEL), lectotypehere chosen.

Distribution. Foundin south-westWestern Australia from Perthto the southcoast as far eastas Albany.

Noles. Two subspeciesare recognized.

Branchletsterete subsp.fastigiata Branchletswith four very naffow glandularundulate wings subsp.tenuior

Boronia fastigiata Bartl. subsp.fastigiata

Brqnchlets tercte. Leavesnarrowly lo broadly elliptic or obovate,1-2(3) cm long, entire or almost so. crlaeJmostly terminal to longbranches.Pedicels oflateralflowers ofcyme with a pairofextremely minutely bracteolesor incipient buds at base.

Distribution. Found in south-westWestern Australia from Perthto Collie and from there south-east to Kojonup and Albany.

Habitctt. Collectors' notesindicate that it occursin latedtic gravel

Noles. This subspeciesis similar in its inflorescenceand in its floral charactersto B. harkeriana from New South Wales.

The typeof B.fqstigiala is ofa variantwith leavescrowded (the intemodesc. 4 mm long), broadly elliptic, 6-9 mm long, andsermte. It wascollected in thePlantagenet county which encompassesthe area aroundAlbany,Denmark, andthe stirling Range,The specimen matches material collectedbetween slab HutCreekand cranbrookby capt. A. Dorrien Smithin 1910and n earKoj onupby w.E. Blackallin 1933. All recent collectionsofsubsp./aslrgrala have come from betweenPerth and collie suggestingthat it no longer occurs fufiher south. Some collections from neal Albany are intermediatebetween this subspeciesand B. denticulata. 'puus ur qnJcs,,!\olur BDt,\l.otD lullgDH

'url?Jlsnv 'qE.lnpqEl{ ulelseld lseM-qlnos pu€ funqung uao^\}oqpuftol uollnqr4sre

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?rlp4snv urolse/A ,uoslrl\ LII worJ rtuo.log snuaB eql ul sxlt .r\ou pue s ueu .,\\eN D lned

I NnytsiaVol. 12,No. I (1998)

No/es.In thestrict sense this subspecies is onlyknown from a few collectionsall from the same locality about2Tkm north ofBunbury. Its distributionis therefore evidently quite separate from that ofthe other subspecies.However, material o f B.juncea frontthePembedonto Donneybrookarea, that is here referred to svbsp.minimq, is somewhatintermediate in morphologybetween the typesofsubsp.jarcea and s}bsp. tninima.

Boronia junceasubsp. lanillora (Bal1l.)Paul G. Wilson,rlal rov.

BoronialanifloraBatl. in Lehm.,Pl. Preiss. l: 165(1844). Trper BehveenMtMelviJleand MtElphinstone, 11 October1840,.L. Preis.r 2030 (synr MEL);Sussex Dish ict, WestemAustl.alia' December 1839, Z P,'e6s 2037 (.lecto:LD),lectolype herechosen.

Boronialaniflctravar.mauanthaBaftl.inlehm., Pl. Preiss l:165(1844). Ilpe: SussexDistrict,Westem Australia,Decembe r 1839,L. P reiss2037 (/eclo. LD), lectotypehere chosen.

Branches (lnd leqvesglabrous or very sparselywoolly. Pedicelswoolly' 2-5 mm long Sepals triangular, acuminate,c. 4 mm long, woolly. Pelcrlsbroadly obovate with rounded apiculateapex, c. 7 mm long, very sparselywoolly alongmidrib outside,sparsely woolly within Ovalt glabrous. (Fjgure6B)

Distribution. Found near the south coastof westem Austlalia betweenAugusta and walpole and also Irear Albany.

Hqbitdt. Growitlgin seasorallyswampy soil

Nomencldturalnote. Bartling did not cite collectionsafter his descriptionof-8. /a niflora,buthe didaller the descriptions of each of his two varieties, micranlhq awl macrctnthd. I have basedthe name B. laniflora onthetype ofvar' fiqcranthq

Boroniaj unceasubsp. micrantha (Bartl )PaulG.Wilson,stal et comb nov

Boronialaniforavar. micrdnLh.tBafil.itllehm.,Pl. Preiss. 1: 165(1844). Zlpe; BehveenMtMelville and MtElphinstone,Westeru Australia, 11October1840,I P/er'ts2030(rso:MEL711185)'

Brenches end leqvesglabrous or very sparselywoolly Pedicelssparsely woolly,2-5 mm long' sepals narrowly triangular, acuminate,5-6 mm long, sparselywoolly. Petals oboyate,acuminate, 5-7(10)mrn long, very sparse ly woolly alongmidrib outside, sparsely woolly within Oval,)'somewhat woolly. (Figure6C)

Distributian. Found near the south coast of westem Austmlia betweenAlbany and walpole.

H ubitat. Growing in seasonallyswampy situatlons

Boroniaj unceasubsp. minima PaulG. Wilson,szbsp llov

Rami et folia glabra.Pedicellisparse lanati, l-2.5mmlongi. sepalaovatavel trjangulafia, acuta, 1.5-3 mm, longa,sparse lanata. Petala obovata, acuminata vel apiculata, c 3 mm longa,glabra vel secus costam sparselanata. 'urur jtq .dsqns Z = rts.S (uorgiC rV) r'rrrl,, dsqns C'6a6( plolxotD f 7\ DLtjuo.l)nu - J 'GgI - uDtupoaA e) onunf dsqtls q'(1n99lDtuaA H) Dtut/iuDt (lsgns, V sr.n\av onun[Dtuua{! 9 etratd

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rrrlnrtsnvurels.rlA rLrarl Diuuot! snuri .ql ur lrxlrt^\au pue s.rulu ^\eN lroslr/A D lnpd I40 Nuylsiavol. 12,No. 1 (1998)

Etymolog,t. TheLatin minimus refersto the small flowers of this subspecies.

Boronia molloyaeJ. Drumm. in Hook.,London J. Bot.2: 169( 1843)as Mo lloyi. TIpe: BetweentheVasse River andAugusta, Westem Australia,l Dntmmond(n.v.).

B. el..ttiorBaftl.ilrLehm.,Pl. Preiss. 1 : 170(1844). Type:"Inhunosa-arenosishumidis interfiutices densos vallisjugi montium Darling's range(Perth)" IDarling Range, Pedh, Westem Australia], September 1841, L. Preiss20l3(lecto:LDl'isolecto;MEL'725044,725045),lectotype herechosen.

B. semi/brlilisF.MtLell., Fragm.2:98 (1860). Iype: FranklinRiver,WesternAustralia,G. Mmwell(n.v.).

Boronia spathulataLindl., SketchVeg. SwanR. 17(1839). Z/per SwanRiverColony,Westem Australia, J. Drummond(holo. CGE photo seen).

B. macrqBarll.inLehm., Pl. Preiss.I : 167(1844). Zype;NearMahogony Creek, Darling Range,Western Australia, l3 September1839, L. Preiss2026 (lecto. LD), lectotypehere choserl.

B. spathulatavar.ramosaBenth., Fl. Austral.l t 32'7(1863). Zlpe. SwanRiverColony, Westem Australia, .J.Drummond s.n. (sy : MEL); EastMt Barren,Western Australia,G. Mmwell (syn;MEL).

Distribution. Found in the south-westof WesternAustralia fron Perth southto Augusta and eastto IsmeliteBay.

Noles. This speciesmay bereadilydistinguished from similarspeciesby the opencymoseinflorescence, the bracteolatepedicels (ofwhich the upper portjon is fleshy and glandularpunctate), and the slender sparselypilose style.

Boroniatetragona Paul G. Wilson,sp. n or,.

Herbaperennis, glabra, ad 70 cm aJta.Rami tetmgoni, angulis acutis, laevibus. Folia intemodiis breviora,sessilia, integra, marginibus dense papillosis, multoreductasuperiora, infedoraremotaelliptica, superioraovata, summa triangularia. F loresin umbellispedunculatis dispositi; pedicelli c. 10mm longi, graciles,ebracteolati, rubri. Bracteaecaducae. Sepala ovata, 3 mm longa,glabra, lanato-ciliata, decidua, Petalalate elliptica, c. 7 mm )onga,pallido Iosea,ad apicerotundata; antherae minutissimae apiculatae. Ovariumglabrum; stylus columnaris vel clavatus,c. I mm longus,glaber; stigma hemisphericum, brunneum.

Tlprsr Westem Australia, Ambergate Reserue,south-west of Busselton,3 November 1993, G.J.Keighery 12938(/rolo: PERTH 04566475).

Perenni.il herb to'70 cm high, glabrous,not forming a lignotuber. Sle,4?stetragonous when young; internodesnostly 4- 12 cmlong. Leavessessile, diminishing in sizetowards apex, with denselypapillate margins; middle and iower leaveselliptic, to 4 cm long; upper ieavesovate; uppermost leaves narrow triangtlar. I nJlorescencea terminalpeduncuJate umbel;peduncle slender, to 2 cm long,bmcts caducous; pedicelss lender and smooth, c. 1 cm long,dark red. Sepals open or stightly imbricate,ovate c. 3 mm long, slightly woo1lyciliate. dark red, deciduous.Pelals imbficate, broadly elliptic, c. 7 mm long, roundedat apex,glablous, pink with redmedial stripe on abaxialsid€. Slazens all fertile;sepaline filaments very 'Jeldua ^q pepnlcurDxel0qlJo ouo ]nq llu sopnlcxslBql 'pazruSooal reuuuure ur pegrdflolcel uoeqsEq ll aDlDAlDj,jtxesur esoql uro.r.1 pedu a.raqsuotlcos aquo qceouo4 sercedspeculd eqqcrq,n olur aa,rDruqtull:ces eqltepun souos a^gpopnlour (g6g 1).te16ug

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Figrrc'7. Boronia cynosa sccd. A - lateral view, B - adaxial surl'ace-C _ lateral view with endocarp and arii attaclred. D - transverse sectioD, all x20 Drawn fto EA. Grif.fin 1610.

3. Boronia sect.Valvatae .t./l7,

Boronia sect.l/qbatae(Benfi.) Engler,Nat.Pflanzenfam.3/4: 135 (1896). - Boroniaser.Valvatae Benth., Fl. Austral. 1: 308, 3 I 1 (1863). - Boroniqsect. ValvoboroniaKurftze,nom. l//eg. in T. Post& Kuntze, Lex.Gen. Phan.74 (1903), based onpreceding. Zype. to bedesignated.

Boronia subg.Robonra Rchb., Iconogr, Bot. Exot. 54 ( 1824). Type:B. ledifolia (Y ent.)J. Gayex DC.

Typirtcqfion.This sectionis beingrevised by Marco Duretto(MEL) who will designatea lectotype. 'pe,unc ,{1lq8r1s'els^elc FI,ro €u Jo urJo]rsnJfl,norjeu o1 eleJel'JepualssaaoaT 8urja,tor ,{3lncs ,{e;6e Surur.rogptru SurlorloJXe uoos elorlnc :sluq pei.lnc esrorluur11r,tr Suno( ueq,,rr re,ro 11e snolnraqnd 'eleiel 'Surpeardg ,(lelnurlu puD ,{lesruds spltlJuD.tg q8rq Lur 0t , g??,/r/,!peqcuuJq flesuep

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'urnsolrd 'ulnsnlqo'uIn8uol 'runl€AEIc 'unnurluoc snsre,tursuq ruruI c le,\runocrJpurl,{c uuFrls la sn1(1s 'ruru8elur 'psocru]ai. :runlnJeqndelmnrtl ulnrla^o u]nrq€I8 runcsrC aululncrduelnur ru eeJeqlue :ol€r [c ruecrdupa 'ef,uolqo'EssDrc Eurleledelue :slErlrf, 'usornrle,\ snsre,\ Luecrde 'ulDcunil'DlrAulc eurledosoluE :srlDulluE]seluetuEllCelelnorduuou'uxaguruouuretrdp'erluacso:qu13'erqnroprlled'elncetiuoltuurg, '€lEAo'eoctsuoc 'plEAlBA 'eFuoJ '€]? 'elecuqull nlEled erluecseguorle rutu [-E Z o eledog snluloelcuJq 'sn8uol 'snsseJc 'u1ul:nc ursuq pe u[u Z-l snllacrped selEuruJelt-l seJolC elnroqndolnurru 'u8uol 'ur ecrporu lrlru0 [ -L LuroJrsnJe]sn8uu 1e,l erla;al olsn8ue ur log rlnraqndotnururle esruds rlnruea

'^o/ dr'uos1116C Irr?4u11,{qdou,{ror eluorog

'drecopue 1e1uece1dlualsrsred 1crq1 'urnlrq ,{lluenbe{ erll puE rpeurlaql'elsolrelno lleLu oql,(q pozua]celeq, sr uorlaes srrluo pees eq.L saloN

'DlpuJal 'op81a 'DtDlD B puu'DlDloacuol g g Druong ;poutruexesB,l\ ,{Soloqdrou peesesoq,,r serJeds 3ur,{olloJ oql sepnlcuruoltcos eql 'peuuep ,{lluesoJdsu're^e,\\oH pouluuotap eq ol lllls eJuuorlces srql3:o uorldrrcsruncrrc eql puesercedsJo Jequnu eql qlog pautwoxasatcadg

'ozx 'o.E]lns '^ror^ 'paes ll! \rorti:rslrlprr luulpnllSuol- J lplxepe- g p-rotcl- V otolDDluoiou 8.rn8t,I

.eqdur,{q (g ern8rg) eurosoreleue seSurlcu tuelsrsrod pue Ic;l:$ AIIensndro)opua puarDld paraloc 'poes Iuspq-qnsJo 1esEqSuuado loztzlp1J Je^oxaluuq fssolE qlr,\\ Jo eseqte pu€ ur8reur lerxupe 'ur8reul ,eso11rded re,&oluo llturs a{dry lrrrc}seurtatuos lurxupr BuolBJee\rll tunltH 1cu1q ,(1e1nurur 'I"ur ,pedEqs-feuprlpaag €tselralno:qloovrsD$aJ o^EcuocJoluU urBJelulErx?pE tBaloqd.tout paas

,uoslr^l Itt erlDqsny urotscr\\ worJ Dlualog snuat c$ ur ?xxl ^\ru pu? sauru r\r3N D InD

\- NuytsiaVol. 12,No. I (1998)

7-10 mm long, c. I mm wide, obtuse,slightly flattenedabove and sometimessulcate, very minutely puberulouswith antrorselycurvedhairs . Flowers l-3 termiral to branches.Pedlcelthick, 1-2mm long, sparselyand minutely puberulous,with a pair of small (c. 1 mm long) ovatebracteoles near the base. Sepalsimbricate, ovate,2.5-3 mm long,two nanowerandslightly longer,coriaceous, glandularpunctate, dark reddish brown, glabrousto very sparselypuberqlous, ciliolate, Pelal,.tvalvate, leathery,ovate, c. 5 mm long, acute,pale red, glabrousto very sparselyand ninutely puberulousoutside, puberulous within, prominently glandular-punctateall over; apex neither inflexed nor sub-apically apiculate. Stamensallferrile sepalinefilaments clavate, c. 2 mm long,obliquely truncate, walty in upperhalf, ciliate; petaline filaments thick, oblong, c. 1.5 mm long, warty at apex,ciliate; antherscordate, pale yellow, c. 0.8mm long,with broadconnective,loculinotrcflexedatdehiscence,minutelywhite-apiculate, affixed by a minute sub-apicalstipe to adaxial side offilaments. Drsc a narrow entire ring, glabrous. Ovary hemispherical,c. I mm high,minutelypuberulous. Si}/eand stigmacontinuous, cylindricalto somewhat clavate,c. I mm long, obtuse,pilose in lower hall F/u il and seednot seen.(Figure 9)

Additional specimenexamined. WESTERN AUSTRALIA: 13 km SW of90 Mile Tank, Frank Hann National Park, 11 Nov. 1979, K. Newbey64?7 (PERTH).

D istribution. This specieshas only beenrecorded fron FrankHann National Park,c. 90 km nofth-east of Lake King township, WesternAustrulia.

Hqbitat. Fol'lmd,in Eucalyptussalmonophloia openwoodland on well-drained clayey sand.

Consemqtionstatus. This speciesis known from only one locality where, accordingto the collector (K.R. Newbey), it is very common. An attemptwas m adein 1996to re-collectthis speciesbut it could not be found, probably due to a fire having passedthrough the areathe previousyear. A Priority Two classificationwith priority for surveywould probably be appropriatefor this species.

Etynologt. The specificepithet is derivedfrom the Greekwordscoryne, a c\tb, andphyllon, aleaf,with referenceto the shapeofthe leaves.

Notes. Boroniq corynophylla is distinctivebecause ofthe shapeofits leavesand the exfoliating cuticle on its brancheswhich gives the plant a glaucousappearance. The valvate aestivationof tl'lepetals is interestingsince the sepalsare imbricate. This sjtuationis also found in B. a/gida, a speciesofsouth- eastemAustralia, and with which -8.corynoplTylla is probablymost closelyrelated (M. Duretto, pers. com.). Seedis requiredto positivelydetemine its sectionalposition.

4. Boronia sect.Cvanothamnus

Boroniasect.Cyqnothannus(Lindl.)F.Muell.,Fragm.9: I 13(1815). - CyanothamntsLindl., SketchVeg. SwanR. Col. I 8 (1839). - Boroniaser.CyaneaeBenth.,Fl.Attstral. 1:309 (1863) basedonCydnothamnus Lindl. - Boroniasect.Cyaneqe (Befih.)DeW ild. nom.illeg.,lcon. Select.2:67 (1901). Type:B. ramosa (Lindl.)Benth., lectotype here chosen.

Typifrcqlion. The two speciesof Cy.tnoth.jmnusthat were describedby Lindley, C ramosus and C. tenuis, are csrrently included in Boronia secl.Cyuathdmnus; the one that is most widespreadis selectedas lectotype. 'LZBLtaq aN tuo4 u,rpro pu! rultl 'G ) G D'E) I puvC'J) ruulZ'(g) urut! Z'(V) ruru :sl?qelDcs pu?.srp- eulplodpue oulledes -0'l?tod - g 'spd:s 0l lqsrd H'suerusls - E're,\\og- C'pnq ra,tou- J lolq.uErqpu? J€el - B'q.urrq Suua^\ou-V DlttqdoutrorDiuorog 6.5mFrC

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Seed morpholog,,. Seed kidney-shaped,adaxial margin flat or convex. Te.tlarugose; outer testa diaphanous,black usually with grey powderyareas between ridges. Hilumlinear alongadaxial margin. Raphe minute at base of seed. Chalqzal opening basal covered by raphe. Placental endocarp membranous,caducous. (Figure 10)

Fi]rnc10. Barcnia coelr/e.rcenr seed. ,{ - lateralvlew, B - adaxialsurface, C - longitudinalradial section. all x20. Drawn tromP.G. Ililson 2/0. species examined. of the nine westem Australian speciesthat are recognizedin this section the following sevenwereexaminedfor seed characlers Boraniabeeckeaceq, B. busselliana,B. coerulescens, B. fabianoides, B- inconspicua,B. penicillata, and B rqmos.t

Nole,r.The seedio this sectionis fairly unjform, exceptfor variationin size. Therugose testa, black with powderygrey betweenthe ridg€s,makes the seedunique in the genusand enables the sectionto be readilyrecognized.

Boroniaacanthoclada PauJG. Wilson,s2. nov.

Fruticuluseffusus c. 30 cm altus;ramulidivaricati, spinescentes, milutissime puberuli. Fol ia alterna, saepein ramisveteribus fasciculata, angustissime obovata,3-6 mm longa,supra concava, crassa' glabra. Floressolitarii, adramulosbreves terminales;pedicellus suprabracteolas breves gracilis,2-3 mm longus. Sepalaanguste triangularia, I .5-2 mm longa,carnosa, glabra. Petala elliptica, c.4 mm longa,glabra, alba.

Typus: Frank Hann National Park, Westem Australia, 29 September1984' J M. Brown 4051555 (/rolo:PERTH 01258494).

Spreadings/rra6 c. 30 cm h igh, divaricatelybranched. Branchlets spinescent, extremely ninutely puberulous. Ledves alterrate,often fasciculateon tbe older wood, very narrowly obovate,rounded below,concave above,3 -6 mm long,thickened, glabrous or almostso. F/ou,erssolitary, telminal to short shoots: (above the smallbracteoles) slender,2-3 mm long SePaAnaffowlytriangular, I.5-2mm 'rd,{toloq LLrorl !r^\?-r( '(C'J) '(E'l) '(A'S) '(y) 'sulultls ultu I ruttl Z tu- t urur g1 :sllrq cp.S Irtsrd puE .srp - D aulplod pu? eullEd.s - C 'Fted 'r5^\ou 'q.u?rq - g - C 1!:lJo o.?.Ins lrrxep! - J \lsurrq Suue.{\o$ - g - y Dpotxol.lluDrDaua./og I I ornirC

{-} I 4-} -{tr d \- ER4K #4Fvn \7

'elurrdorddueq o1srueddu epoc uorle,nrasuoco,rl,{lJJorJd V suorsDrtolero^os uo loJ paqcleesq8noqlle palecolor ueeq lou sEqlr :IJud IBuoll€NuueH {uplC ulpotcalloc sp,r qcrq,r uorlcelloceuoeqluo.g u,,iroul,{lug sntDtsuottt)^-tasuoJ

'IeArxB JeAopues ur 8ur,ro;8 su,r true1deqt teqt etecrpursalou s(rolJelloc aqJ- iDtlqDH

'uIlp]lsnV '8ur; '

'uor1ce11oced,{1 sql uro{,(luo u,rou) pauttur:xasuawnadg

'Suol 'eleJel-Jepuels ( 11 a.rn8tg) ueeslou Trn,ig alnurureu8rts :snolq€18 ruur Z l r el,{ls :snorqelS '3uol 'JIEq ,f,onp snorqelSrsr6 elulncrde-e1rq,u,{pun1qpue (lUoqs tuur g 0 2 sloqlu€:el€rlrc Jeddnur '3uo1qo 'allouptul esocnr:e,t-ru1npue13 ,{1,rorruuslueu€[U lsuture]s a]tq,r punoJBatetcund-rslnpu€18 'elerTrso'snorq€13 'dp poxouul peuo)J!qt e q1r,t'3uo1 ruur y c'tr1dy1asp1a4 snorqu13',{qseg '3uo1

'uoslrrlA elle'qsnv ur:ts:lA wc]] Dtuoroq snuot eq1 ul rxel n\ou puD seurpu /!\cN D InDd I NuytsiaVol. 12,No. I (1998)

Etymotogt. The epithetis derivedfromthe latinizedG teekwofds clcctntho-, spiny, and c/adas, abranch. ly'oles.This speciesis mostclosely relate d.to B. coerulescens.The presenceofalternate leaves, although not unique,is unusualin the genusas are the spinescentbranchlets. Itis similar in appearancetosome yariantsof B. coerulescenssubsp. spinescers but differs in the shapeofthe leavesand in theil alternate phyllotaxy.

Boroniabaeckeacea F. Muell , Fragm.4: 28 ( 1863 )

Distribution. Found in southem WesternAustralia from Pjngrup eastto Mt Ragged

Nole,!. Two subspeciesare recognized.

Leavessimple,erect, suborbjcular, 2-3 mm long ...... subsp baeckeacea patula Leavessimple or tdfoliolate, + spreading,broadly elliptic to obovate'4-7 mm long subsp

Boroniabaeckeacea F. Muel). subsp. baeckeacea

Zeovessimple, suborbicular,concave, + appressedto branch,2-3 mm long'

Distribution. Found in southernWestern Australia from Pingrup eastto Mt Ragged'

heavy soil Habitqt, Recordedas growing in a variety of situations,over granite or limestoneor in

Boronia baeckeaceasubsp. patula Paul G. Wilson,szDsp' rov

Folia+ patula,ad apicemrecurva, simplicia et integravel tri-lobatavel trifoliolata; lamina circularis et c. 3.5mm longa,vel obovataet 5-7mm longa,supra profunde concava'

& Typus: About 45 km north-west of Clyde Hill, Western Australia, 2l May 1993, G F Craig B. Haberley21l3 (holo. PERTH 3218600).

leave.s+ spreading,somewllatrecuryed at apex,sessile, simple and entire or 3-lobed,ortrilbliolate; laminacircular and c. 3.5mm long,to obovateand 5-7 mm long,deeply concave

south- Distribution. Found in the vicinity ofMtNey andMtHeywood, c 90km north-eastofEsperanca' east Westem Australia.

Habitat. Growing on clay-loam in mallee.

Newbey82'79 Selectedspecinrens exqminecl. WESTERN AUSTRALIA: I km W of Mt Heywood' K (PERTH);SWofMtNey,lSGeorgel5g05(PERTH);265kmNEofMtHe)'\tood'lt'RArcher1708912 (PERTH).

attempthas been Consemetion status.The collectionsall comefrom two relatively small areasand no madeto determineits degreeof security A Priority One rating would appearto be appropriate' 'uot1ucgtd,41o1ca1ro; elqelt€A? lou sI euo paloqunuun eql ulq,{q paltc su,'tL1g uotlcelloc ,{1uo ecurslnquor13elloc pareqrunuun eqlJo ruagedButqcuurq eql uro{ leqllde sgIsedsaql pe^lJopuleqlusg 'peJoq[nu 'leqlo ,{lluepr^g soqcup.rqluecseurds,{llueuruord Sutpuards seq lou sl qalq,\\ eI{J soqcuDJq 'dlrJnls tuocsourds,{1ec;ecs Sutpuecse seq 1r lunq € ruo{ ql,tor8 qser3Sutluut sr t€ql luald " Jo sJ '/8 'suollcalloc loqulnu oLpJoeuo (t98 I urtqlueg) ellu.rlsnySoero11 eq1 3ur I u,tl uJq^\ uoull lJds JLI] perpnlspeq oql€ql elecrpurol ruErltueB,{qpasn sD,{\ alnpacold qctq,lt'1ur pel ut pe]u]1epun sua2sawds leqlrdeeql uorpalloc qcpeJoslooqs eql uO () qJoq,^\ou)runuElrolooH lunll€qjeH eql urele,,\\ qclq.{\Jo qloq uoxel srq]Surqucsep ueq,,!\ s.puotuuInJC seruefJo suollcellot o,'r,r1per pnls ,{ltuJpl^ a LucqtuJB

'rl'.Eqlueg 'gL sE' ur ')al fq pelrcfllcelocut su,{\uollselloc puowwntqaql'uottttcl.lidtl

'(8i89f Z1gr{ io.s?:) aloq)L\puowLutuO t' '€rlcnsnv ure1se16'fuo1o3reAIU u?,\\S adtJ Gg80 6le :l lEIsnV Il'tlluog suacsaurdsg

'(1461)g97:1ursltnN'uosp/[ Dln€d( qlueg)suarsaulds dsqns suocseln'taoJ Eluorog

'erpl€SlooJ rEouol lsee pue ur8pelunlrqol qlnos puU sou,{uduIoU ell€Jlsnv ujelse/A luoqlnospuelul ul strxao uollnqlrJSlO

'(l/6l) 002:t €ls nN'uoslUd DlnEdsluctds dsqnssuerslluaoJ sluorog

'urLBllsnVruetsel^[ ureqlnos pue I]llPrtsnVq]noS ur:qtnos qEnorql uuolcr1 uJetsee'sel€/d qlnos ,treN urels?a-qlnosetueJ1xa uI srncco sot.ladsqnsstq1- uotuqlllslO

'(t6t8E0l 'uesoqce;eqed.{lo1cel 'lgt,l iopaps'.96189011AW)raUann !'t98I raqure^oN'IrLrol3lA 'suerduruJo eqJ'oJlol l oJrotrl rad10 ,{eguaqctng spre,trol u€sep 3ql ul pue'suPIdluPlgJql ul sLLrq 'llanl^ A)notuololtt4ll ad,t (tg8I) I I:IEuolcr^ cos lqd sIIaJf cs/aJ.trqndrctsuatsclntaot g

'uosoqcereqed,Qolcel'(t6t8S0I-lat{,triopalnllanq,1'gtr91;eqrualdeg'eqe4snVqlnos',{€l.Inlr'{eql.}o acuPl]ueoqlr€eu qru3s eellEt ,l rzr4l 31ngs.recuedS ol la^lu,{€ljnt{ oql uo qnlts sellBl\ oqlluo4 secBld vaJJBq[l:uo1]DJadlJ (ygg 1) 11 :1euoprl cos ltqd suu.rJ'llenyl gsuacsatqop tc^suaJsalnraulB

sucJsalnraoJdsqns llenl,l C sdeJsalnlaoJsluorog

',{lerl€^ lBcrd,{leql q1r,tcldfioruoq 1reluru o1supagrdftolcel os sl Jell€leql qrlq x1o'suacsa'tqolS J€tpue suacsaqndter, 'seueue^ o^\l eql Jepun,{1uo 1nq eureu setceds eql repunulup uollcellooou e,\€? JelIerV\I uolottJtd'(J

'uesoqc0.10q '(16p9 .lgV\I 'g '€rll?rls '.{PrrnI ed.{1o1cel gg 1 :opal) plptTI,I y yg 1requaldeg nV qlnos Jeq:Uo acuDrtue r'llont{ aql.rceuqnJrs oolp6 :adQ (yggy) I t : l guolcr1 ios IIqd suerJ C suarselnraorBluoJofl

'self,edsqns lecrd,(tarp olul lse,\\ pue qlnos eql ol sepurSll r{Juerqeues eq1uo punoj SurequortBuBr\ e qcnsqll{\ sleu€aleltsses eelql olur popl^lp elP sa,\?alssaquo etrlosFlleuolsPcco Suolruut l ol se,tealole,^oqo Jo lelncJls seqsetcedsqns srql saTortr

'so^€ol eql Jo emlsod eql ol SuuJeJoJ'Surpeetds'sn.zlzd ptortt ulle'I oql uorJ pe^JJepsr teqtrda cgtceds q\s eqJ- ASopwAg

ellxlrsnv llJ)ls)lA wa)].tuotoB snu.13fqt ul pxr],,!\3upue soruru^\aN'uosllA g ln?d 150 NuytsiaVol. 12,No. I (1998)

Boroniafabianoides (Diels) PaulG. Wilson,Nuytsia l: l19 (19'70).-Et'iostemonfabianoidesDiels,Bot. Jahrb.Syst.35: 322, tab.39 K-L (1904). Iype. Norlh ofGrasspatch,Westem Australia, 2 Novemberi 901, l. D/cA528q(isor PERTH 0l600r26).

Distributiotl. Found nearCowcowing and from Lake Ki[g eastto the FraserRange, southem Western Australia.

Noles. Two subspeciesare recognized.

Petalsc. 3 mm long,white with greenmidrib ...... subsp. fabianoides Petals5-7 mm long,white rvith pink midrib ...... subsp, rosea

Boroniafabianoides (Diels) Paul G. Wilson subsp. fabianoides

Low spreadings.4rti6 with divaricatebranches, I0-3 0 cm high. ,Sepalstriangular,c. 1mm long. Pela/s broadlyelliptic,c.3 mm long,2mm wide,glabrous, white with thickenedgreen midrib.

Distribution. Norsemanto Esperancearea of southem Westem Austrajia.

Flabll. In eucalyptwoodland on brown loam, sometimescalcareous.

Notes. This subspeciesgenerally has hispidulous branchlets but a glabrousvariant is foundc 70km eastofNorsenan alongthe Eyre Highway (G.F. Craig3064; R.J.Cranfield9384)

Boronia fabianoidessubsp. rosea PaulG. Wilson, szbsp. nov.

Fruticuluserectus ramis ascendentibus, 30-60 clrraltus. Sepalatriangularia, 2(3 ) mm longa Petala lateelliptica, 5-7 mm longa,glabra, alba praeter costam crassam pallido rubram

Iypzs; Woodline, c. 95 km east-north-eastof Norseman, Western Australia, 7 August 1980, G.J. K ei g hery 2998 (ho I o; PERTH 94229 4).

Erectsrlr, with ascendingbranches, 3 0-60 cm high. Sepalstriangular, 2(3) mrn long Petalsbroadly elliptic,5-7 mm long,4-6mn1 wide, glabrous, white with athickenednarrowpinkmidrib

Selectedspecimens examined. WESTERN AUSTRALIA: I0 km E of Norsema\ D E Albrecht4035 (PERTH);Cowcowing,Septernber 1904, M. Koch1231(MEL);30km S ofNorseman,KNewhey 6869 (PERTH);2.5kmNWoflake Cronin,K Newbe1831I (PERTH)

Distribution. Found betweenLake King and the FraserRange, Westem Australia, with a 1904record from Cowcowing.

flabitat. lo clay-loamoften over greenstoneor granite.

Ery talog. The subspecificepithet rosea, from the Latin,r'ose6, referstothe colourofthe broadmidrib of the petais. '(8€8 t yZZO1.q111g6:o7o4) 9779 ncuadg ,1pplag[uD.tJt U'266t Kl11tI 'erl€llsnvuelsey'A'eJeseJ[Jnense'IlunotrN]Jo,{J€punoqqtnosuopU,,r\er{SJolsEe-qlnoslu{€:.r,?dfJ

'BAecuoc 'e8uo1 'eBuolqo eJdns'BSnlqo ruu 97-9 elsn8ue1e,t etrueutl 'sn8uol 'sqrcur8 '.rFlorloJ-E- ego11o3isqeclnsudns ujru g1 -61 snlorpdie8uol uur 0t-S I gruqeuurderlog

'40, 'dsqns drqr?s'uoslr/d C ln€d Busr nansal esoruu.r uluo,rog

'urlsrlsnVurelssl^d'puerg e1ede3 o11seepue eSue;8uqrr1g eql olqlnos Je^rUuosrqct\V\I uounqulste

'uosorlc '(28n8 .IAhl' oreqod,qorrel S0l I BV8iO|1AVtr;opalosl '.q1:opal) '?'tt8I 'ra,ta ;EOZSs,a'td ,{lnt l Suruue3uo loo.rqseldg;ee51 :adKJ (tgBI) OZt:L 'ls.Ilsnv IC'qtueg( Iwg.)ptlo/i4pup te^DsowD.tg- (tl8l)OLI:t ssrordld' urqo1.J].ptEtsnuo/lqpuo snuutotltotltuots- (E9g1)679:1erqfnN'uoslr^ADInEd(l eB) BlloJlqlausdsqns BsoruBJ eruorog

'€rlu]lsnv 'uorad ulelseld tJ,'{reau osle pue ue8e;epuuqol q ou qUed uo4 e?ueg 3u11.rc9eq1- uolfiqulsr1

'dsqns esorue,r qlueg( 1pur1)esourur uruorog

suulnansal dsqns "" e,toqee,recuoc '8uo1qo ,{1,noleu o1reaurl stegue.l :Z 'elerel BlloJlqleuB dsqns s^oqu pslleuuEqc Jepuelssleuee'l Z alultdeceru8tts iSuol Lutu€-Z sleclped :I esoure.r'dsqns ls3upurt,(o ro uuoJrsnJsul8rls:Auol urur g l -g slecrped I

'pazruSocerore sarcodsqnsoorqJ salop

'(uoos 'gCJ oloqd :uts) saFu?t',[ 1dn3'.(uaes otoqdgDC irL{.s)prouwn.tq f'6tgl'r.rler}snvu.tetseld',(uoloCr.^r1tue,rs:ad,{; (6tgl)gI U ue.r\S '3a,1qcp1g ' Iprn1.rrsowt:u snuwo4louDKT - (g9g1) g7g:1 leasnv lJ' rlluoq ( lpur.I) esoruureruorog

'1.ello, '9 '8 sr 86 leql lseSSnssuorlcelloc Jeqto s[lJo s8zl uo puoulrunro ,(q 6 puu sJequnueq1;o 3ur1r.r,req1 qlt,t uost.tudLuo3g6;equnu eql e,tr?ol seos leoqs oql ot paxg st luql Setraqrunu leut8uo us s€r{'IAJ,\i 'qreq ured,'(1o1celosr eql sEeraq,r98 palaqrunust; qreqle ed,fto1ce1eql Surrueqle eqseqj- uoun![rd,4J

'uesoqc '(S 't 'erlprlsnv ereq edflotcsl t SSSOt 1ifl\I :opapsr lueesoloqd :opal) g6 puoww,?./O1 urelse^\ 'punog e8roeg3ur;,1puuro^r.UuE,\\S uee,t\leg:adt1 (6991)779:t lslsnv lJi qtuag BIBII!JruedE!uoroB

'rasoldsqns jo suorljolloa ,{llolsso ororu 'elncrd€ eql qlr^\ see.l8€esr,,!\reqlo tr ]nq rsqlueelrq,r lueururordpu€ Suol u.rurt J syedas3uolqo-,lroIeu '( 'toutsrp s?q I €ZI tlrox W) tO6I;equeldeg ur epuur Sur,roc,roJ eq1uorj uorlcelloc e18urseq1

'peqsnJc '}uocs ueq,r llelllsIoqlueu u 8ur^€qse roqlouu,{q puD lutrureedse qlr,n rg€ruole fllue?und Sureqs€ rotrello, ouo ^q poqllcsepueeq seq lue1d srql saTo,rg

'uoslr/A nrllrtsnv urolsrrt uJorJolua,log snuat ollt ul l]xvt r\ou puu so(uu ]!\cN D lnt]d Nuytsiavol. 12,No. 1 (1998)

Compact woody perennictlto 30 cm high. Branchesnarrowly 4-winged when young due to the decurent basesof the leaves,becoming tetragonal or terete with age, puberulous between wings otherwiseglabrous; intemodes 8- 15 mm long. Zeale.rpinnately 3 -5-foliolate, in all I 5-40mm long; slender,10- I 5 mm long,channelled above; leaflets linearto narrowly oblong, 5-25 mm long,1.3-2 mm broad, obtuse, concave above.

Other specimensexanlned WESTERNAUSTRALIA: l0km N of JurienBay tum-off , R.J. Cranfield 1467(PERTH);MtLesueur,C.A. Gardner9081 (PERTH);MtLesueur, E.A. Griflitt 1838(PERTH).

Distribution. Recordedfrom nearMt Lesueur,d. 200 km north ofPerth, south-westWestern Australia.

Hal51l.7/.Grows in sandor gravel over laterite.

Conserration stcttus.This subspeciesis known fron tbur populationsofwhich at leastone is in a national park. A Priority Two ranking is recommended.

Etymolog,,. This subspeciesis namedafter the hill where it is to be found.

Notes. Boronia ranosa subsp.Iesueurar.r appears to grade into B. ra,nosa subsp.anethifolia wl'tich is found in the heathlandsurrounding Mt Lesueur;the most obvious differencebetween the two taxa is to be found in the shapeofthe leafletswhich in subsp.anethifolia areslender-terete.

Boronia westringioidesPaul G. Wilsott,sp. rov.

Ramuli stellatopubescentes. Folia confefta, ascendentia, opposita, semiteretia, acuta, 5- 10mm longa. Floressolitariilpedicellus turbinatus, l-2 mm longus,dense stellato pubescens; bracteolae foliaceae, c. I .5 mm longae. Sepalatriangularia ve I ovato acuminata,2-3 mm longa,crassa, sparse pubescentia. Petalaelliptica,5-6 mm 1onga,manifeste rubro glanduloso punctata,tenuia, modice stellato pubescentia.

Zypas. 8 km west of Forrestania-SouthernCross road on Hyden-Norsemanroad, Westem Austmlia, 12October 1995,. F. Craig33l8(holo.PERTH 04202104;1"!oi CANB, K).

Erect shrub to 75 cm high with ascendingbranches. BrQnchlelsstellate pubescant. Ieqres sometimescongested, ascending, opposite (orternate, or the leavessessile and trifoliolate), semiterete or narrowlyelliptic, channelled or concaveabove, acute, 5- 10 mm long,smooth, glabrous or stellate puberulous. F/owers solitary in uppermostleaf-axils; pedicel turbinate, 1-2 mm long, denselystellate pubescentand with shortclavate glandular hairs; bracteoles medially positioned, foliaceous, c. 1.5mm long. Sepalstriangu lar to ovate-acuminate,2-3mm long,thick, with prominentredglandular dots when dry, sparselypubescent. Pelals elliptic, 5-6 mm long, with prominentred glandulardots all over, thin, moderatelystellate pubescent and with clavateglandularhairs, pale pink, nidrib nolprominent.Staminal flanents compressedterete, glandular tuberculate towards apex, ciliate; anthers minutely white- apiculate.Ovr4rglabrous or puberulous;style terete, c. 0.5mm long,sparsely pilose; stigma subcapitate. Seedellipsoid, c. 3 mm long, roundedat apex and base;testa rugose, black with white deposit. (Figure12)

Other specimensexazfued. WESTERN AUSTRALIA: 6 km W ofjunction ofFonestania Rd androad from Hyden,B. & -B. BackhouseHl9 (PERTH); far inland from King GeorgeSound,lSS2, A.Y HQssell (MEL);7km WoflakeCroniu, K ly'ewbey5199(CANB, PERTH);c. 65kn EofHyden,K.Newbey9169 TPER.TTI}. nbbs'uurdfd Crurort u{ero (l '(J) '(g) -c pu?a'C) ruruI ulu.tZ tutug'(V) tuttrS:sreqol?cs poesJo soceJms p}'epE pux ler:lel llsrdpuerslp-f 'su.rrtets 'q.uurq eulplodpuE eulpdes - O'(ar€JrnsplxEqE) IDlod - J'13^{o$ - g - V saP!a!3u!,tls2trDluorog ZI a'n8li

THoo , Cy'iIU' /LU't

'uosllrlA vrlE4sny urrlserld uJou Dluarog snua8 oql u! €r€t ^\eu pu? scrnuu ^{oN C Ined Nuylsiavol. 12,No. I (1998)

Distribution. Known from asmallaread. 70 km nofthoflake King townshipand c. 65 km eastofHyden, in southerninland WesternAustralia.

H..tbitat, kecotded as growing on loamy sandplains.

NoleJ. There is somevariation in tbe leaves.Whereas in most collectionsthey aresimple and decussate in two collectionsthey areternate with someofthe leavessessile and trifoliate.

Conservatianstatus. This specieswarrants a Priority Two ranking sinceit is known from only one area which is not currently under threat.

Etymology. The epithet alludesto the plant's resemblanceto membersofthe genus Westringia.

Affrnities. This speciesdiffers from its closestrelative, B. baeckeacea,principally in its leafshape.

Acknowledgements

The illustrationswere kindly preparedby my daughterAnnemarie. Material wasborrowedfrom the NationalHerbarium ofVictoria(MEL), RoyalBotanicGardens, Kew (K), Trinity College,Dublin (TCD), BotanicGardens ofAdelaide (AD), andth€ Botanical Museum, Lund (LD);I amgateful to tlese herbaria for their assistance.This work was parlly fundedby a grant from the AustualianBiological Resources Studv.

References

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