Formation of Nangwalbibra, Garo Hills, Meghalaya, India

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Formation of Nangwalbibra, Garo Hills, Meghalaya, India Paiaeobotanisl 49 (2000) : 225-237 0031-0174/20001225-237 $2.00 Study ofplant megafossils from the Tura· Formation of Nangwalbibra, Garo Hills, Meghalaya, India R.C. MEHROTRA Sirbal Salmi Institute ofPalaeobotany, 53 University Road, Lucknow 226007, India. (Received 05 April 1999; revised version accepted 07 October 1999) ABSTRACT Mehrotra RC 2000. Study of plant megafossils from the Tura Formation of Nangwalbibra, Garo Hills, Meghalaya, India. Palaeobotanist 49(2) : 225-237. Leaf-impressions collected from the Tura Formation (Upper Palaeocene) ofNangwalbibra. Garo Hills. Meghalaya belong to Grewia ofTiliaceae. Alalalllia of Rutaceae. Schleichera ofSapindaceae. Mallgifera of Anacardiaceae, Termillalia as well as Calycopleris of Combretaceae, Syzygiul11 of Myrtaceae and ChrysophyllulIl of Sapotaceae. In addition, a few palm leaves of uncertain affinities are also described. The modem comparable forms of the fossils indicate a tropical warm and humid climate in the area during the deposition of the beds. Key-words-Leaf-impressions. Angiosperms. Tura Fonnation, Garo Hills, Upper Palaeocene. ~ ~ '1i4IcR'1~<s({1 ~ ~ :ll~HP"l~ ~ ~ ett 1JTU qrfu- % -a j(C\q I~ql~tff qiT -3l&r1'1' m~T%~ mu~ ~ ~ (~qR1~1t111) ~~'1 ~ ~ <t\ miT qcffi~ it 1ijlcR1~0l{1 if @ B quf ~~ Q11<{lI~QW ~ ~ Wt~, vn ~ it;ftfcrqr, ~ it it "ffftim, it ~ ~tm ~. ~ it eWiR741 tzci /lidl'hNRfI, ~ it A7$J{tf:;jlJlf nm it #i1$vi'Jfi!;ITIlf B ~ ~ or~ ~ ~ ~ ~ ~&1fUm rr{~. 01T~ ~ ~ <t\ <t\ c8 A it B ~ ~ ~ ~. 'fffiW it f1"m it ~ IiR if \jOUI iflkOl·tfrl, ~ tzci \3Wj <{iT m ~-quf ~~'1, ~, @ ~, miT qcffi~, ~ qR141W1. INTRODUCTION bearing beds are located on the right bank ofthe Nangal stream within an area of3 km. The fossils are found in greyish-white HE rocks of the Tura Formation are well exposed near to buff coloured clay stones ranging 305-366 m in thickness. T Nangwalbibra (25°28' N, 90° 42' E), a town situated near The locality map (Fig. I) and litholog of the area are already Williamnagar, East Garo Hills District, Meghalaya. The plant gi ven by Bhattacharyya (1983) and Saxena el al. (1996). «:> Birbat Sahni Instilule of Palaeobotany. tndia 226 THE PALAEOBOTANIST Age Group Lithology 2S~ 40' Oligocene Simsang Formation WILLIAHNAGAR (100 m thick) JWEST KHASI Eocene-Palaeocene Jaintia Kopili Formation , HILLS (450 m to 500 m thick) '-~ KHERAPARA 2S' Siju Limestone ) 20' '­ (100 m to 160 m thick) W[ST~ _ ASSAI1 ~' __ , --.,.." 1OCoAL.~.-. '- -).~ v r' 5 ~HARA I ; PI [ G,k ';. LAY> 1:\ Tura Formation ~(;,MC",",.t .""tl&J.\ t "\. .~ ../'-.--. ~J._._. ~~) o 10 (180 m to 250 m thick) r-"·-._. __ ~ ....,. BAHGlAOESH' Upper Cretaceous Coarse grained sandstone Fig. 1- A map showing the fossiliferous locality near Williamnagar (after Saxena el ai., 1996). and conglomerate (60 m thick) however, the general stratigraphic sequence noticed in the area is given below in Fig. 2. As evident from this figure the Tura --- --------- ------Unconform itY---- -- --------------- -- ------- -- --- Formation is considered as Upper Palaeocene in age (Saxena er at., 1996; Mehrotra ef at., 1998). Precambrian Coarse grained granite The average annual rainfall in the Garo Hills varies from ganodiorites, banded gneiss 250 cm to 320 cm. The climate is moderate during winter, but and quartzite gradually becomes warmer from middle of March. The monsoon sets in from mid-May and the area is covered with a Fig. 2-General slraligraphic sequence ofGaro Hills. Meghalaya (after Raja thick forest of mostly deciduous trees having a thick Rao, 1981). undergrowth of thorny bushes. The fossil flora of Nangwalbibra is not well known and SYSTEMATICS only a few plant fossils are described from the area. Lakhanpal (1964) first recorded a palm leaf from this area. Later, DICOTYLEDONS Bhatlacharyya (1983) described several forms, namely NeLumbo nagaLensis (Nymphaeaceae), Lirsea sp., Phoebe Family-TILIACEAE Bentham & Hooker f. subLanceoLara (Lauraceae), Arlocarp us garoensis (Moraceae), Genus-GREWIA Linnaeus Triumfetta rhomboideocarpa (Tiliaceae), Heteropwzax sp. (Araliaceae), Osmanthus eocenicus, Ligustrum turaense, Species-GREWIA SAHNII (Lakhanpal) comb. nov. AnthoLirhes oLeaceaeformis, A. campanuLatus (Oleaceae), PI. 1·1 Nipa sahnii (Palmae) and Poacites sp. A field excursion was undertaken to ascertain the NeoLitsea sahnii palaeofloral composition of the Garo Hills, Meghalaya and a 1954 Lakhanpal, p. 28, PI. I, figs 3, 4. large number of plant megafossils were collected from the Description-Leaf simple, more or less complete, Tura Formation of Nangwalbibra. Most of the fossils are in symmetrical, elliptic, about 9·0 x 4·0 cm in length and width the form of leaf-impressions, a few fruits and seeds were also respectively; apex slightly broken but appearing acute; base collected from the same locality. Some are complete and well asymmetrical, obtuse, normal; margin entire; texture appearing preserved while others are fragmentary in nature. chartaceous; petiole length and width about 1·3 x I'5 cm All the type specimens, negatives etc. ofthe fossils being respectively, normal; venation acrodromous, basal, imperfect; described in the paper are deposited in the repository of the primary veins (10) three in number, mid-rib stout, slightly Birbal Sahni Institute of Palaeobotany, Lucknow. curved near the apex, a pair of lateral primaries coming off at The terminology used in describing leaves is after Hickey base at an angle of25°-30° and running up to the middle of the (1973) and Dilcher (1974). The identification of leaves was leaf; secondary veins (20) arising from the mid-rib at about done by comparing them with the herbarium sheets at Forest one third of its length from the base, about 10 pairs visible, Research Institute (FRI), Dehradun; Central National alternate, 6-11 mm apart, angle ofdivergence moderately acute Herbarium (CNH), Howrah and Birbal Sahni Institute of to narrow acute (60°_40°), decreasing gradually towards the Palaeobotany (BSIP), Lucknow. apex, abruptly curved near the margin to form loops,joining MEHROTRA-STUDY OFPLANT MEGAFOSSILS FROM THETURA FORMATION OF NANGWALBIBRA 227 superadjacent secondaries at right angle to sometimes at acute and running up to the middle ofthe leaf; secondary veins arising angle, moderately thick; intramarginal vein absent; tertiary from the midrib at about one third of its length from the base, veins (30) fine, angle oforigin usually RR, pattern percurrent, alternate, angle of divergence moderately acute 10 narrow simple, obliquely placed, predominantly alternate, close, some acute, decreasing gradually towards the apex, abruptly curved arising directly from mid-rib to join with the lateral veins; near the margin to form loops with the superadjacent higher order venation present; areoles well developed, secondaries; tertiary veins percurrent, simple, obliquely placed, oriented, usually quadrangular, medium to small, vein-lets not predominantly alternate, close, some arisingdirectly from the observed. mid-rib to join with the lateral veins. Holotype-Specimen no. BSIP 38086. Occurrellce-Tura Formation; Nangwalbibra near Species-GREWIA GAROENSIS (Lakhanpal) Williamnagar, East Garo Hills District, Meghalaya; Upper comb. nov. Palaeocene. PI. 1·2 Affillities-The important features of the fossi I leaf are: acrodromous basal imperfect venation, asymmetrical base, Trel1la garoensis entire margin, elliptic shape, secondary veins forming loops 1954 Lakhanpal, p. 27-28, PI. 1'1,2 and percurrent tertiary veins. These characters indicate its Description-Leaf more or less complete with slightly affinities with Grewia Linnaeus of Tiliaceae. Further broken apex, simple, symmetrical to slightly asymmetrical, identification of the fossil leaf up to specific level is very narrow ovate; preserved length (including petiole) ll·5 cm, difficult as there are lot of variations in leaf structure among maximum width 4-4 cm; apex appearing acute; base different species ofGrewia. The base varies from asymmetrical symmetrical; margin serrate, acute, apical side concave to to symmetrical (Grewia laevigala Yahl, Herbarium Sheet Nos straight, basal side straight; sinuses angular, spacing more or FRI 2362, 62203), while the margin varies from serrate to less regular, simple on complete margin; texture seemingly partially serrate to entire (G. calophylla Kurz, Herbarium Sheet chartaceous; attachment with petiole normal; petiole 2·5 cm Nos CNH 80761,1924). in length and 2 mm in thickness; venation acrodromous, Though fossil woods (Bande & Srivastava, 1995) and imperfect; primary veins three, one straight and moderately fruit (Paradkar & Dixit, 1984) of Grewia are of common thick called mid-rib, two laterals coming off at the base at an occurrence in the Deccan lntertrappean beds of India, its only angle of 40° and running half way of the lamina length; leaf, G. foxii Lakhanpal (1954), is known from the Eocene of secondary veins 7 pairs visible, arising slightly above the base, Damalgiri, Tura District, Meghalaya.lt differs from the present alternate, 0·8-1·6 cm apart, angle of divergence narrow to fossil in having serrate-crenate margin, ovate shape, smaller moderately acute (30°-50°), upper pairs more acute than lower, size and twisted petiole. A few years back Srivastava et al. moderately thick, uniformly curved, sometimes joining (1992) also described a fossil leaf of Grewia riliaefolia Yahl superadjacent secondaries; intersecondary veins not present; from the Pleistocene beds of Mahuadanr, Palamu District, intramarginal veins absent; tertiary veins percurrent, angle of Bihar. It also differs from the present
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