County, Utah, U.S.A.; Horizon B Or C, Uinta Formation, Uintan Faunal Age, Late Eocene

Total Page:16

File Type:pdf, Size:1020Kb

County, Utah, U.S.A.; Horizon B Or C, Uinta Formation, Uintan Faunal Age, Late Eocene 1974 AUFFENBERG: FOSSIL TORTOISE CHECKLIST 173 County, Utah, U.S.A.; Horizon B or C, Uinta Formation, Uintan Faunal age, Late Eocene. GEOLOGIC RANGE.-Late Eocene. GEOGRAPHIC RANGE.-Utah, U.S.A. Subgenus fMegalochelys Falconer and Cautley Megalochelys Falconer and Cautley 1837, p. 358. Colossochelys Falconer and Cautley 1844, p. 54. Testudo Lydekker 1880, p. 20 (part). TYPE SPECIES.-Colossochelys atlas Falconer and Cautley (= Geo- chelone [Megalochelys] atlas [Falconer and Cautley]). DEFINITION.-An Old World subdivision of the genus Geochelone in which the included species are large, thick-shelled tortoises with swollen areas on the peripherals, nuchal scute absent, two gular scutes, epiplastron strongly bifurcated anteriorly, and no narrowed pectorals. GEOLOGIC RANCE.-Pleistocene and perhaps Recent. GEOGRAPHIC RANGE.-Southeast Asia, East Indies, Mauritius Island, Mascarene Group, Indian Ocean. REMARKS.-An extinct, but specialized small group of large to giant tortoises. Geochelone (Megalochelys) atlas (Falconer and Cautley) Megalochelys sivalensis Falconer and Cautley 1837, p. 358 (subsequently withdrawn). Colossochelys atlas Falconer and Cautley 1844, p. 54. Colossochelys (Megalochelys) atlas Maack 1869, p. 223. Testudo (Colossochelys) atlas Lydekker 1880, p. 20. Testudo (Megalochelys) atlas Lydekker 1889b, p. 209. Testudo atlas Brown 1931, p. 183. ?Testudo margae Hooijer 1948, p. 1169, fig. 1 (Pleistocene, Celebes). Testudo margae Hooijer 1954, p. 486. TYPE.-British Museum (Natural History); an epiplastron. TYPE LOCALITY AND HORIZON.-Siwalik Hills, Punjab, India; Siwalik Beds, Manchhar Series, Boulder-Conglomerate, Early Pleistocene. GEOLOGIC RANGE.-Early to Late Pleistocene. GEOGRAPHIC RANGE.-Punjab, India, Upper Burma, Java, Celebes, and Timor. REMARKS.-According to Lydekker (1889b) G. atlas is close to G. perpiniana. The 12-foot composite reconstruction of the type and as- sociated specimens by Falconer is incorrect; it should be closer to 6 feet (Lydekker 1889b). For latest discussion, see Hooijer (1971). Geochelone (?Megalochelys) cautleyi (Lydekker) Cautleya annuliger Theobald 1879, p. 186 (nomen nudum)..
Recommended publications
  • §4-71-6.5 LIST of CONDITIONALLY APPROVED ANIMALS November
    §4-71-6.5 LIST OF CONDITIONALLY APPROVED ANIMALS November 28, 2006 SCIENTIFIC NAME COMMON NAME INVERTEBRATES PHYLUM Annelida CLASS Oligochaeta ORDER Plesiopora FAMILY Tubificidae Tubifex (all species in genus) worm, tubifex PHYLUM Arthropoda CLASS Crustacea ORDER Anostraca FAMILY Artemiidae Artemia (all species in genus) shrimp, brine ORDER Cladocera FAMILY Daphnidae Daphnia (all species in genus) flea, water ORDER Decapoda FAMILY Atelecyclidae Erimacrus isenbeckii crab, horsehair FAMILY Cancridae Cancer antennarius crab, California rock Cancer anthonyi crab, yellowstone Cancer borealis crab, Jonah Cancer magister crab, dungeness Cancer productus crab, rock (red) FAMILY Geryonidae Geryon affinis crab, golden FAMILY Lithodidae Paralithodes camtschatica crab, Alaskan king FAMILY Majidae Chionocetes bairdi crab, snow Chionocetes opilio crab, snow 1 CONDITIONAL ANIMAL LIST §4-71-6.5 SCIENTIFIC NAME COMMON NAME Chionocetes tanneri crab, snow FAMILY Nephropidae Homarus (all species in genus) lobster, true FAMILY Palaemonidae Macrobrachium lar shrimp, freshwater Macrobrachium rosenbergi prawn, giant long-legged FAMILY Palinuridae Jasus (all species in genus) crayfish, saltwater; lobster Panulirus argus lobster, Atlantic spiny Panulirus longipes femoristriga crayfish, saltwater Panulirus pencillatus lobster, spiny FAMILY Portunidae Callinectes sapidus crab, blue Scylla serrata crab, Samoan; serrate, swimming FAMILY Raninidae Ranina ranina crab, spanner; red frog, Hawaiian CLASS Insecta ORDER Coleoptera FAMILY Tenebrionidae Tenebrio molitor mealworm,
    [Show full text]
  • Egyptian Tortoise (Testudo Kleinmanni)
    EAZA Reptile Taxon Advisory Group Best Practice Guidelines for the Egyptian tortoise (Testudo kleinmanni) First edition, May 2019 Editors: Mark de Boer, Lotte Jansen & Job Stumpel EAZA Reptile TAG chair: Ivan Rehak, Prague Zoo. EAZA Best Practice Guidelines Egyptian tortoise (Testudo kleinmanni) EAZA Best Practice Guidelines disclaimer Copyright (May 2019) by EAZA Executive Office, Amsterdam. All rights reserved. No part of this publication may be reproduced in hard copy, machine-readable or other forms without advance written permission from the European Association of Zoos and Aquaria (EAZA). Members of the European Association of Zoos and Aquaria (EAZA) may copy this information for their own use as needed. The information contained in these EAZA Best Practice Guidelines has been obtained from numerous sources believed to be reliable. EAZA and the EAZA Reptile TAG make a diligent effort to provide a complete and accurate representation of the data in its reports, publications, and services. However, EAZA does not guarantee the accuracy, adequacy, or completeness of any information. EAZA disclaims all liability for errors or omissions that may exist and shall not be liable for any incidental, consequential, or other damages (whether resulting from negligence or otherwise) including, without limitation, exemplary damages or lost profits arising out of or in connection with the use of this publication. Because the technical information provided in the EAZA Best Practice Guidelines can easily be misread or misinterpreted unless properly analysed, EAZA strongly recommends that users of this information consult with the editors in all matters related to data analysis and interpretation. EAZA Preamble Right from the very beginning it has been the concern of EAZA and the EEPs to encourage and promote the highest possible standards for husbandry of zoo and aquarium animals.
    [Show full text]
  • The Conservation Biology of Tortoises
    The Conservation Biology of Tortoises Edited by Ian R. Swingland and Michael W. Klemens IUCN/SSC Tortoise and Freshwater Turtle Specialist Group and The Durrell Institute of Conservation and Ecology Occasional Papers of the IUCN Species Survival Commission (SSC) No. 5 IUCN—The World Conservation Union IUCN Species Survival Commission Role of the SSC 3. To cooperate with the World Conservation Monitoring Centre (WCMC) The Species Survival Commission (SSC) is IUCN's primary source of the in developing and evaluating a data base on the status of and trade in wild scientific and technical information required for the maintenance of biological flora and fauna, and to provide policy guidance to WCMC. diversity through the conservation of endangered and vulnerable species of 4. To provide advice, information, and expertise to the Secretariat of the fauna and flora, whilst recommending and promoting measures for their con- Convention on International Trade in Endangered Species of Wild Fauna servation, and for the management of other species of conservation concern. and Flora (CITES) and other international agreements affecting conser- Its objective is to mobilize action to prevent the extinction of species, sub- vation of species or biological diversity. species, and discrete populations of fauna and flora, thereby not only maintain- 5. To carry out specific tasks on behalf of the Union, including: ing biological diversity but improving the status of endangered and vulnerable species. • coordination of a programme of activities for the conservation of biological diversity within the framework of the IUCN Conserva- tion Programme. Objectives of the SSC • promotion of the maintenance of biological diversity by monitor- 1.
    [Show full text]
  • Russian Tortoise Care Scientific Taxonomy/Name: Testudo Horsfieldi
    [email protected] http://www.aeacarizona.com Address: 7 E. Palo Verde St., Phone: (480) 706-8478 Suite #1 Fax: (480) 393-3915 Gilbert, AZ 85296 Emergencies: Page (602) 351-1850 Russian Tortoise Care Scientific Taxonomy/name: Testudo horsfieldi Avg. carapace length: Males average around 5.5 inches and 1.3 pounds. Females average around 7.5 inches and 3 pounds. Natural location: Dry/arid, hot areas around the Mediterranean Sea (i.e. parts of Turkey, Afghanistan, Russia, Iran, China, and Pakistan). Russian tortoises live in burrows for nine months of the year. They are the only land tortoise (Testudo spp.) without a hinge on their plastron near the rear legs, and the only land tortoise with only four claws/toes on each foot. Temperature: 70-80 degrees Fahrenheit with basking area of 90 degrees. For tortoises kept inside, keep thermometers at various locations in the enclosure in order to check the temperature in their environment. Humidity: 30-50% (dry environment). Use a hydrometer to check humidity levels in the enclosure. Lighting: Natural sunlight (not blocked by windows) is the best source of light for tortoises (and other reptiles)! If being kept indoors during the winter or due to an illness, use a broad- spectrum heat bulb. The most efficient light source is the Exo Terra Solar Glo. It is a mercury vapor bulb that emits heat, UVB, and UVA. UV light has multiple benefits, including calcium metabolism and improved appetite and activity. Proper calcium metabolism helps protect against metabolic bone disease. Follow manufacture directions on proper installation and use a clamp lamp with a ceramic fixture to prevent melting.
    [Show full text]
  • Environmental and Ecological Factors Affecting the Presence of Giant Land Turtles in the Late Cenozoic Author: Orion Jenkins-Hou
    Environmental and Ecological Factors Affecting the Presence of Giant Land Turtles in the Late Cenozoic Author: Orion Jenkins-Houk GEOL394 Advisor: Dr. Thomas Holtz Due 4/28/2020 1 Abstract: Various species of turtles within Testudinidae (true tortoises) and the recently extinct Meiolaniidae of Australia grew to immense proportions throughout the late Cenozoic, including a significant number of taxa that have persisted into modern times. Although these giant land turtles mostly occur on islands today, there are cases of extinct giant land turtles on every non- Antarctic continent during the Cenozoic. This raises an interesting question: if giant turtles can occur on the continents, presumably in the presence of both predators capable of penetrating their defensive carapace and other herbivores competing for the same food sources, what other factors may be related to the evolution of gigantism in land turtles? This study tests the influence of two ecological factors, presence of durophagous (bone-crushing) predators and competing herbivores, and three environmental factors, mean annual temperature, aridity, and landmass type (insular versus continental) on occurrences of giant land turtles. The results of the Fisher exact tests collected demonstrate that the presence of competing herbivores and insularity have a significant effect on the occurrence of modern giant land turtles. Miocene giant land turtles appear to occur independently of all five factors, while Pliocene giants tend to occur in areas of higher average temperatures. Pleistocene
    [Show full text]
  • Reptilia: Testudines: Testudinidae
    REPTILIA: TESTUDINES: TESTUDINIDAE Catalogue of American Amphibians and Reptiles. Ernst. C.H. andT.E.J. Leuteritz. 1999. Geochelot~edenticrrlnra. Geochelone denticulata (Linnaeus) Yellow-footed Tortoise Te.s/~ldodenticrilat~~ Linnaeus 1766:352. Type locality. errone- ously listed as "Virginia." Syntypes, two specimens (one a stuffed juvenile. the other shown in Schoepff 1792:plate 28, fig. 1) in the Muscum de Geer: one of which was kept in Upsala and the other in Stockholm (Andersson 1900, Schoepff 1792). Anderson (1900) lists the holotype as NRM De Greer 21 (not examined by the author). ~estudotabulatae Walbaum 1782: 122. Nome11 illegitimum, see FIGURE 1. Geochelonedenticulott~. Photograph by Roger W. Barbour. Comment. Testudo ressellata Schneider 1792:262. Type locality, not given. Type, not designated. Testudo tabulara Schoepff 179356. Type locality, "In Africa australi?." Type. no1 designated. See Comment. Testudo terrestris hmsiliensis: Schweigger 18 12:445. See Com- ment. ~ Testudo terrestris arnericnna: Schweigger 18 12:445. See Com- ~ ment. Testudo terrestris cayer~rzensis:Schweigger 18 12:445. See Com- ment. Testudo terrestris .rurinatnen.si.s: Schweigger 18 12445. See Comment. Testudo hercules Spix 1824:20. Type locality, "sylvis de flumen Solimoens" [Rio Solimoes], Brazil. Holotype, an adult, now FIGURE 2. Geochelone denriculata, plastron. Photograph by Roger W. Barbour. I lost (Hoogmoed and Gruber 1983). Testirdo sc[rlpfa Spix 1824:21. Type locality, "sylvis juxta flumen Amazonum" [Rio Amazon], Brazil. Syntypes, originally five specimens in the Zoologische Staatssammlung Miinchen, of tebral and pleural areolae; yellowish or orange pigment also which four juveniles ( 273810, a shell; and 275310 A, B, C), occurs at the lower edge of each marginal.
    [Show full text]
  • Ancient Mitogenomics Clarifies Radiation of Extinct Mascarene Giant Tortoises (Cylindraspis Spp.)
    Title Ancient mitogenomics clarifies radiation of extinct Mascarene giant tortoises (Cylindraspis spp.) Authors Kehlmaier, C; Graciá, E; Campbell, P; Hofmeyr, MD; SCHWEIGER, S; Martínez-Silvestre, A; Joyce, W; Fritz, U Date Submitted 2019-11-26 www.nature.com/scientificreports OPEN Ancient mitogenomics clarifes radiation of extinct Mascarene giant tortoises (Cylindraspis spp.) Christian Kehlmaier1, Eva Graciá2, Patrick D. Campbell3, Margaretha D. Hofmeyr4, Silke Schweiger5, Albert Martínez-Silvestre6, Walter Joyce7 & Uwe Fritz 1* The fve extinct giant tortoises of the genus Cylindraspis belong to the most iconic species of the enigmatic fauna of the Mascarene Islands that went largely extinct after the discovery of the islands. To resolve the phylogeny and biogeography of Cylindraspis, we analysed a data set of 45 mitogenomes that includes all lineages of extant tortoises and eight near-complete sequences of all Mascarene species extracted from historic and subfossil material. Cylindraspis is an ancient lineage that diverged as early as the late Eocene. Diversifcation of Cylindraspis commenced in the mid-Oligocene, long before the formation of the Mascarene Islands. This rejects any notion suggesting that the group either arrived from nearby or distant continents over the course of the last millions of years or had even been translocated to the islands by humans. Instead, Cylindraspis likely originated on now submerged islands of the Réunion Hotspot and utilized these to island hop to reach the Mascarenes. The fnal diversifcation took place both before and after the arrival on the Mascarenes. With Cylindraspis a deeply divergent clade of tortoises became extinct that evolved long before the dodo or the Rodrigues solitaire, two other charismatic species of the lost Mascarene fauna.
    [Show full text]
  • Proposal for Amendment of Appendix I Or II for CITES Cop16
    Original language: English CoP18 Prop. XXX CONVENTION ON INTERNATIONAL TRADE IN ENDANGERED SPECIES OF WILD FAUNA AND FLORA ____________________ Eighteenth meeting of the Conference of the Parties Colombo (Sri Lanka), May 2019 CONSIDERATION OF PROPOSALS FOR AMENDMENT OF APPENDICES I AND II A. Proposal The proponents propose the transfer of Geochelone elegans (Indian Star Tortoise), from CITES Appendix II to Appendix I in accordance with Article II, paragraph 1, of the Convention. Specifically, this species meets the biological criteria found in paragraphs C i) and ii) of Resolution Conf. 9.24 (Rev. CoP16), Annex 1, due to a marked decline in population sizes in the wild observed as ongoing or inferred or projected on the basis of levels or patterns of exploitation, and a high vulnerability to intrinsic (i.e. late maturity, low reproductive output, long generation time and low population recovery potential) and extrinsic (i.e. a decrease in the area and quality of habitat) factors, and a reduction in recruitment due to indiscriminate off take. B. Proponent India and [co-proponent- Sri Lanka] C. Supporting statement 1. Taxonomy 1.1 Class: Reptilia (Laurenti, 1768) 1.2 Order: Testudines (Batsch, 1788) 1.3 Family: Testudinidae (Batsch, 1788) 1.4 Genus, species or subspecies, including author and year: Geochelone elegans (Schoepff, 1795). Taxonomy follows the standard nomenclatural reference for Turtles (Fritz & Havas, 2007). 1.5 Scientific synonyms: Testudo elegans Schoepff, 1795; Testudo stellata (Schweigger, 1812);Testudo actinodes(Bell, 1828);Testudo actinoides(Bell in Gray, 1844);Testudo megalopus(Blyth, 1853);Peltastes stellatus(Gray, 1870); Peltastes stellatus seba(Gray, 1870). 1.6 Common names: English: Indian star tortoise, star tortoise French: Tortue étoilée de l’Inde Spanish: Tortuga estrellada de la India 1.7 Code numbers: ITIS number 551777.
    [Show full text]
  • Gopherus Polyphemus (Daudin) Gopher Tortoise
    215.1 REPTILIA: TESTUDINES: TESTUDINIDAE GOPHERUSPOLYPHEMUS Catalogue of American Amphibians and Reptiles. to Late Pleistocene of Florida (many unpublished localities; see Hay [1930] and Holman [1958, 1959] for a few that have reached AUFFENBERG,WALTERANDRICHARDFRANZ. 1978. Gopherus the literature) and Late Pleistocene of South Carolina (specimens polyphemus. examined by senior author). Closely related (perhaps conspecific) populations occur in the Pleistocene of Kansas (Preston, 1971) and Texas (specimens examined by senior author). The Late Gopherus polyphemus (Daudin) Pleistocene or Recent Gopherus praecedens (Hay, 1916) is a syn• Gopher tortoise onym of G. polyphemus (Auffenberg, 1974). Testudo polyphaemus Bartram, 1791:18. Nomen nudum. • PERTINENTLITERATURE.Douglass (1975, 1977a) provides Testudo polyphemus Daudin, 1802:256. Type-locality, "bords de extensive bibliographies on living Gopherus. A few examples are la riviere Savanna et pres de I'Alatamah"; restricted by listed here. Symbiotic relationships: Young and Goff (1939), Carr Schmidt (1953) to "vicinity of Savannah [Chatham Co.], (1952). Habits: Hallinan (1923), Carr (1952), Brode (1959), Hansen Georgia." No type specimen designated. (1963), Auffenberg and Weaver (1969). Predators: Douglass and Testudo depressa Guerin-Meneville, 1829:5. Type-locality, Weingarner (1977). Growth: Pearse, Lepokovsky, and Hintze "I'Amerique septentrionale." No type specimen designated. (1925), Goin and Goff (1941). Courtship: Auffenberg (1966). Phys• Testudo gopher Gray, 1844:4. Type-locality, "N. America," re• iology: Bogert and Cowles (1947), Cantrell (1964), Sullivan and stricted by Schmidt (1953)to "vicinity of Savannah [Chatham Riggs (1967a-c), Thorson (1968), Spray and May (1972), Ross Co.], Georgia. Two syntypes presumably in the British Mu• (1977). Mental gland secretions: Rose et al. (1969). Parasites: seum (Natural History).
    [Show full text]
  • Psammobates Geometricus, P
    SHORT NOTES 65 SHORT NOTES have more extensive distributions. The genus hw11111ohi11e.1 appears to conform to this general pattern in that the species HERPETOLOGICAL JOURNAL, Vol. 3, pp. 65-67 ( 1993 ) with the smallest range. P. geo111e1ric11s. is a southwestern Cape endemic. Poynton ( 1989: p.2) describes two contrasting faunas in the southern subcontinent. "the one being more A HYPOTHESIS EXPLAINING THE cool-adapted and presently showing re lict patterning in areas ENIGMATIC DISTRIBUTION OF THE of high altitude and latitude, the other being warm-adapted GEOMETRIC TORTOISE, with continuous ranges brought about by recent range expan­ sion". He goes on to point out that this led to a "'dynamic PS AMMOBATES GEOMETRICUS, IN model involving reciprocal spreading and withdrawing of the SOUTH AFRICA interlocking cool- and warm-adapted groups following the cyclic changes in Quaternary climate". Mouton & Oelofsen ' ' E. H. W. BAARD AND P. LE F. N. MOUTON ( 1988) discuss the importance of the Cape Fold Mountains. the coastal lowlands and the cold Benguela sea current in 'Cof'e Na111re Co11ser\'{/fio11. Pril'llle Bag X50/4, 7599 speciation events among cordylid lizards in the southwestern Srelle11bosch, So111h Africa Cape. During changing climates these mountains become ef­ 'John Eller11101111 Museum. Deparrmenr 1fZoology. Uni1·ersitr of fective barriers trapping warm-adapted populations along the Stellenbosch. 7600 Srellenbosch, South Africa western coastal lowlands. Due to a supposedly increased in­ fluence of the Cold Benguela current on coastal climates (Accepted 8.5. 92) during glacial periods, populations along the coastal lowlands become subjected to more adverse climatic conditions than The genus Psammobmes Fitzinger, 1835 comprises three inland sister populations.
    [Show full text]
  • Ecological History and Latent Conservation Potential: Large and Giant Tortoises As a Model for Taxon Substitutions
    Ecography 33: 272Á284, 2010 doi: 10.1111/j.1600-0587.2010.06305.x # 2010 The Authors. Journal compilation # 2010 Ecography Subject Editors: David Nogue´s-Bravo and Carsten Rahbek. Accepted 7 March 2010 Ecological history and latent conservation potential: large and giant tortoises as a model for taxon substitutions Dennis M. Hansen, C. Josh Donlan, Christine J. Griffiths and Karl J. Campbell D. M. Hansen ([email protected]), Dept of Biology, Stanford Univ., 371 Serra Mall, CA 94305, USA. Á C. J. Donlan, Advanced Conservation Strategies, P.O. Box 1201, Midway, UT 84049, USA, and Copeland Fellow in Global Sustainability, Amherst College, Amherst, MA 01002, USA, and Dept of Ecology and Evolutionary Biology, Cornell Univ., Ithaca, NY 14853, USA. Á C. J. Griffiths, School of Biological Sciences, Univ. of Bristol, Woodland Road, Bristol, BS8 1UG, UK, and Inst. of Environmental Sciences, Univ. of Zurich, 190 Winterthurerstrasse, CH-8057 Zurich, Switzerland. Á K. J. Campbell, Island Conservation, LML, 100 Shaffer Road, Santa Cruz, CA 95060, USA, and School of Integrative Systems, Univ. of Queensland, Gatton, Queensland 4343, Australia. Starting in the late 1970s, ecologists began unraveling the role of recently extinct large vertebrates in evolutionary ecology and ecosystem dynamics. Three decades later, practitioners are now considering the role of ecological history in conservation practice, and some have called for restoring missing ecological functions and evolutionary potential using taxon substitutes Á extant, functionally similar taxa Á to replace extinct species. This pro-active approach to biodiversity conservation has proved controversial. Yet, rewilding with taxon substitutes, or ecological analogues, is now being integrated into conservation and restoration programmes around the world.
    [Show full text]
  • Chelonian Advisory Group Regional Collection Plan 4Th Edition December 2015
    Association of Zoos and Aquariums (AZA) Chelonian Advisory Group Regional Collection Plan 4th Edition December 2015 Editor Chelonian TAG Steering Committee 1 TABLE OF CONTENTS Introduction Mission ...................................................................................................................................... 3 Steering Committee Structure ........................................................................................................... 3 Officers, Steering Committee Members, and Advisors ..................................................................... 4 Taxonomic Scope ............................................................................................................................. 6 Space Analysis Space .......................................................................................................................................... 6 Survey ........................................................................................................................................ 6 Current and Potential Holding Table Results ............................................................................. 8 Species Selection Process Process ..................................................................................................................................... 11 Decision Tree ........................................................................................................................... 13 Decision Tree Results .............................................................................................................
    [Show full text]