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Native Orchids in Southeast Alaska
Native Orchids in Southeast Alaska Marlin Bowles & Bob Armstrong 2019 Preface Southeast Alaska's rainforests, peatlands and alpine habitats support a wide variety of plant life. The composition of this vegetation is strongly influenced by patterns of plant distribution and geographical factors. For example, the ranges of some Asian plant species extend into Southeast Alaska by way of the Aleutian Islands; other species extend northward into this region along the Pacific coast or southward from central Alaska. Included in Southeast Alaska's vegetation are at least 27 native orchid species and varieties whose collective ranges extend from Mexico north to beyond the Arctic Circle, and from North America to northern Europe and Asia. These orchids survive in a delicate ecological balance, requiring specific insect pollinators for seed production, and mycorrhizal fungi that provide nutrients essential for seedling growth and survival of adult plants. These complex relationships can lead to vulnerability to human impacts. Orchids also tend to transplant poorly and typically perish without their fungal partners. They are best left to survive as important components of biodiversity as well as resources for our enjoyment. Our goal is to provide a useful description of Southeast Alaska's native orchids for readers who share enthusiasm for the natural environment and desire to learn more about our native orchids. This book addresses each of the native orchids found in the area of Southeast Alaska extending from Yakutat and the Yukon border south to Ketchikan and the British Columbia border. For each species, we include a brief description of its distribution, habitat, size, mode of reproduction, and pollination biology. -
California's Native Ferns
CALIFORNIA’S NATIVE FERNS A survey of our most common ferns and fern relatives Native ferns come in many sizes and live in many habitats • Besides living in shady woodlands and forests, ferns occur in ponds, by streams, in vernal pools, in rock outcrops, and even in desert mountains • Ferns are identified by producing fiddleheads, the new coiled up fronds, in spring, and • Spring from underground stems called rhizomes, and • Produce spores on the backside of fronds in spore sacs, arranged in clusters called sori (singular sorus) Although ferns belong to families just like other plants, the families are often difficult to identify • Families include the brake-fern family (Pteridaceae), the polypody family (Polypodiaceae), the wood fern family (Dryopteridaceae), the blechnum fern family (Blechnaceae), and several others • We’ll study ferns according to their habitat, starting with species that live in shaded places, then moving on to rock ferns, and finally water ferns Ferns from moist shade such as redwood forests are sometimes evergreen, but also often winter dormant. Here you see the evergreen sword fern Polystichum munitum Note that sword fern has once-divided fronds. Other features include swordlike pinnae and round sori Sword fern forms a handsome coarse ground cover under redwoods and other coastal conifers A sword fern relative, Dudley’s shield fern (Polystichum dudleyi) differs by having twice-divided pinnae. Details of the sori are similar to sword fern Deer fern, Blechnum spicant, is a smaller fern than sword fern, living in constantly moist habitats Deer fern is identified by having separate and different looking sterile fronds and fertile fronds as seen in the previous image. -
Outline of Angiosperm Phylogeny
Outline of angiosperm phylogeny: orders, families, and representative genera with emphasis on Oregon native plants Priscilla Spears December 2013 The following listing gives an introduction to the phylogenetic classification of the flowering plants that has emerged in recent decades, and which is based on nucleic acid sequences as well as morphological and developmental data. This listing emphasizes temperate families of the Northern Hemisphere and is meant as an overview with examples of Oregon native plants. It includes many exotic genera that are grown in Oregon as ornamentals plus other plants of interest worldwide. The genera that are Oregon natives are printed in a blue font. Genera that are exotics are shown in black, however genera in blue may also contain non-native species. Names separated by a slash are alternatives or else the nomenclature is in flux. When several genera have the same common name, the names are separated by commas. The order of the family names is from the linear listing of families in the APG III report. For further information, see the references on the last page. Basal Angiosperms (ANITA grade) Amborellales Amborellaceae, sole family, the earliest branch of flowering plants, a shrub native to New Caledonia – Amborella Nymphaeales Hydatellaceae – aquatics from Australasia, previously classified as a grass Cabombaceae (water shield – Brasenia, fanwort – Cabomba) Nymphaeaceae (water lilies – Nymphaea; pond lilies – Nuphar) Austrobaileyales Schisandraceae (wild sarsaparilla, star vine – Schisandra; Japanese -
Biosystematic Studies of Phacelia Capitata
AN ABSTRACT OF THE THESIS OF J. Steph nShelly for the degree of Master of Science in Botanyand PlantPathologypresented on April 11. 1985. Title: Biosystematic Studies of Phacelia capitata (Hydrophyllaceae). a Species Endemic to Serpentine Soils in Southwestern OregonRedacted for privacy Abstract approved: Renton L. Chambers Phacelia capitataRruckeberg is a member of the Phacelia magellanica polyploidcomplex (species group Magellanicae), a group of wide-ranging, polymorphic perennials in westernNorthAmerica related to the South American P. secunda (= P. magellanica). The purpose of thisproject was to determine, to the extent possible, the interrelationships ofP. capitata within the Magellanicae, using studies of its distribution, cytology, morphology, reproductive biology, and population ecology. Phaceliacapitata is endemic to ultramafic (serpentine and peridotite) outcrops, or similar metamorphic substrates, in Coos, Douglas, and JacksonCounties, southwesternOregon. Field surveys revealed22 extant populations. The species occurs on open serpentine slopes, outcrops, and roadbanks or similarlydisturbed sites, generally in areas with a south to southeast exposure. The nativeplant community with which P. capitata is frequently associated is a Pinus ieffreyi/grass savanna. Soils from five P. capitata sites and two E. corvmbosa sites werechemically analyzed. At six of these locations the soils were derived from fully serpentinized parent materials, and were found to possess low levels of phosphorus and very low calcium/magnesium ratios when compared to an eighth sample from a non-serpentine soil. A soil sample fromone P. capitata site, derived from a metamorphic rock similar in appearance to serpentine, had chemical characteristics intermediatebetween those of serpentine and non-serpentine soils. Chromosome counts from 20 P. capitata populations revealed the existence of diploids (n=11) at 19 of them; tetraploid (n=22) counts were obtained from two populations, oneof these populations also containing diploids. -
TO KNOW in Re On
TO KNOW in re on FEDERAL COOPERATIVE EXTENSION SERVICE OREGON STATE COLLEGE,CORVALLIS Cooperative Extension work in Agriculture and Home Economics, F. E. Price, director. Oregon State College and the United States Department of Agriculture cooperating. Printed and distributed in furtherance of Acts of Congress of May 8 and June 30, 1914. Extension Bulletin 785 September 1959 FERN TERMS Evergreen. Said of plants whose leaves re- Pinna. A primary division of a fern leaf. main green at least until new ones are formed, (The plural of pinna is pinnae.) and of leaves that remain green more than a year. Pinnule. A secondary division of a fern leaf. Fertile leaf. A leaf that bears fruit dots or spore cases. Rhizoid. Simple hair-like structures of a Frond. The leaf of a fern. prothallium, functioning as roots. Fruit band. A line of spore cases, instead Rhizone. A somewhat horizontal and us- of fruit dots, appearing on the margin or un- ually elongated creeping subterranean stem. der surface of fertile leaves of some ferns. Rootstock. Rhizome. Stem. Fruit dots. Small groups of spore cases appearing on the underside of fertile leaves. Sorus. A cluster of sporangia. (Plural of Habitat. The typicalsituation under sorus is son.) which a plant grows. Sporangia. (Spore cases.) The vessels Indusium. The shield-like cover of a where spores are formed. sorus. Spore. The small nonsexual fruit of the Leaflet. One of the divisions of a com- fern. A cell that functions as a seed. pound leaf. Midvein. The central and most prominent Stipe. Leafstalk. vein of a pinna or pinnule. -
An Illustrated Key to the Ferns of Oregon
AN ABSTRACT OF THE THESIS OF Helen Patricia O'Donahue Pembrook for the Master of Arts (Name) (Degree) Systematic Botany (Major) Date thesis is presented March 8, 1963 Title AN ILLUSTRATED KEY TO THE FERNS OF OREGON Abstract approved IIIII (Major professor) The purpose of the work is to enable students of botany to identify accurately Oregon ferns, both as living plants and as dried speci- mens. Therefore, it provides vegetative keys to the families, genera and species of the ferns (Class FILICINAE) found in Oregon. Correct names have been determined using the latest available information and in accordance with 1961 edition of the International Code of Botan- ical Nomenclature. The synonomy, a description, and original draw- ings of each species and subspecific taxon are included. An illustrated glossary and a technical glossary have been prepared to explain and clarify the descriptive terms used. There is also a bibliography of the literature used in the preparation of the paper. The class FILICINAE is represented in Oregon by 4 families, 20 genera, 45 or 46 species, 4 of which are represented by more than one subspecies or variety. One species, Botrychium pumicola Coville, is endemic. The taxa are distributed as follows: OPHIO- GLOSSACEAE, 2 genera: Botrychium, 7 species, 1 represented by 2 subspecies, 1 by 2 varieties; Ophioglossum, 1 species. POLYPODI- ACEAE, 15 genera: Woodsia., 2 species; Cystopteris, 1 species; Dryopteris, 6 species; Polystichum, 5 species, 1 represented by 2 distinct varieties; Athyrium, 2 species; Asplenium, 2 species; Stru- thiopteris, 1 species; Woodwardia, 1 species; Pitrogramma, 1 spe- cies; Pellaea, 4 species; Cheilanthes, 3 or 4 species; Cryptogramma, 1 species; Adiantum, 2 species; Pteridium, 1 species; Polypodium, 2 species, 1 represented by 2 varieties. -
Fall 2001 HARDY FERN FOUNDATION QUARTERLY Marlin Rickard to Lecture
THE HARDY FERN FOUNDATION P.O. Box 166 Medina, WA 98039-0166 (206) 870-5363 Web site: www.hardvfems.org The Hardy Fern Foundation was founded in 1989 to establish a comprehen¬ sive collection of the world’s hardy ferns for display, testing, evaluation, public education and introduction to the gardening and horticultural community. Many rare and unusual species, hybrids and varieties are being propagated from spores and tested in selected environments for their different degrees of hardiness and ornamental garden value. The primary fern display and test garden is located at, and in conjunction with, The Rhododendron Species Botanical Garden at the Weyerhaeuser Corpo¬ rate Headquarters, in Federal Way, Washington. Satellite fem gardens are at the Stephen Austin Arboretum, Nacogdoches, Texas, Birmingham Botanical Gardens, Birmingham, Alabama, California State University at Sacramento, Sacramento, California, Coastal Maine Botanical Garden, Boothbay, Maine, Dallas Arboretum, Dallas, Texas, Denver Botanic Gardens. Denver, Colorado, Georgeson Botanical Garden, University of Alaska, Fairbanks, Alaska, Harry P. Leu Garden, Orlando, Florida, Inniswood Metro Gardens, Columbus, Ohio, Lewis Ginter Botanical Garden, Richmond, Virginia, New York Botanical Garden, Bronx, New York, and Strybing Arboretum, San Francisco, California. The fem display gardens are at Bainbridge Island Library, Bainbridge Island, WA, Lakewold, Tacoma, Washington, Les Jardins de Metis, Quebec, Canada, University of Northern Colorado, Greeley, Colorado, and Whitehall Historic Home and Garden, Louisville, KY. Hardy Fem Foundation members participate in a spore exchange, receive a quarterly newsletter and have first access to ferns as they are ready for distribution. Cover Design by Willanna Bradner HARDY FERN FOUNDATION QUARTERLY THE HARDY FERN FOUNDATION Quarterly Volume 11 • No. -
39516 Federal Register / Vol. 50, No. 188 / Friday, September 27, 1985
39516 Federal Register / Vol. 50, No. 188 / Friday, September 27, 1985 / Rules and Regulations reaction irreversibility or by formation polarography or square-wave (3) Irving, H., “The Stability of Metal of two or more complex species in polarography). Complexes and Their Measurement equilibrium with each other. In this last (3) Interpretation and evaluation of Polarographically," Advances ih case it is necessary to apply the method resu lts, (i) Stability constants Polarography Proceedings of the 2nd by De Ford and Hume paragraph (d) (8) determined for a new substance can be International Congress, Ed. I.S. of this section to calculate stepwise compared with literature values for Langmuir (Pergamon Press, 1960). formation constants. standard substances (see Reference (4) Perrin, D.D., Dempsey, B., B u ffe r (2) Test report, (i) The test report substances, above) and used therefore for pH and Metal Ion Controls. should list for each metal ion to evaluate the strength of its (Chapman and Hall: London, 1974). investigated the half-wave potential complexing ability. (5) “Stability Constants of Metal-ion Complexes,” Part B, Organic Ligands, Ei /2 , co-ordination number and overall (ii) The system is physically stability constant. Compiled by D.D. Perrin, IUPAC meaningful if (A) the value of the Publication on Chemical Data Series, stability constant is positive and (B) the (ii) In addition, the following should No. 22 (Pergamon Press, 1979) also be reported: standard error is less than the constant (6) Grabaric, B., Tkalcec, M., Piljac, L, (A) Type of polarisable micro (the t-test should be used as a criterion). -
Biosystematic Studies in the Genus Piperia (Orchidaceae)
BIOSYSTEMATIC STUDIES IN THE GENUS PIPERIA (ORCHIDACEAE) by James D. Ackerman A Thesis Presented to The Faculty of Humboldt State University In Partial Fulfillment of the Requirements for the Degree Master of Arts June, 1976 BIOSYSTEMATIC STUDIES IN THE GENUS PIPERIA (ORCHIDACEAE) by James D. Ackerman Approved by the Master's Thesis Committee Chairman Approved by the Graduate Dean ACKNOWLEDGEMENTS Dennis E. Anderson's patience, guidance and support throughout this project is most gratefully acknowledged. I appreciate members of my committee, Richard L. Hurley, Robert A. Rasmussen, and Farris R. Meredith for lively discussions and reading the manuscript. I also thank William V. Allen for access to laboratory equipment, Jerry A. Powell for identifying the moths, Arlee M. Montalvo for aid through various aspects of this work, and the curators and staffs of those herbaria that provided specimens for study. TABLE OF CONTENTS Page Acknowledgements Introduction 1 Taxonomic History 7 Materials and Methods 12 Results and Discussion 22 Osmophors 22 Chromosomes 22 Chromatography 27 Interfertility 37 Field Studies 46 Morphology 49 Character analysis 49 Taxonomic concepts 60 Summary 66 Distribution 68 Conclusions 74 Taxonomy 79 Keys to Piperia taxa 79 Piperia 81 Piperia elegans 82 Piperia unalascensis 86 Piperia maritima 89 Piperia maritima var. multiflora 92 Piperia transversa 94 Piperia michaeli 96 iii Table of Contents. Continued. Page Appendices 98 Literature Cited 107 INTRODUCTION Piperia Rydberg is a polymorphic genus and presents complex taxonomic and nomenclatural problems. Variability exists in nearly all morphological aspects including the usually conservative features of the column. Many species have been proposed based on plants showing limited ranges of continuous characters; this has resulted in considerable confusion. -
Adiantum Viridimontanum, Aspidotis Densa, Minuartia Marcescens, and Symphyotrichum Rhiannon: Additional Serpentine Endemics from Eastern North America
Soil and Biota of Serpentine: A World View 2009 Northeastern Naturalist 16(Special Issue 5):111–120 Adiantum viridimontanum, Aspidotis densa, Minuartia marcescens, and Symphyotrichum rhiannon: Additional Serpentine Endemics from Eastern North America Tanner Harris1 and Nishanta Rajakaruna2,3,* Abstract - Serpentine outcrops around the world are known to harbor disproportion- ately high rates of plant endemism. Remarkable cases of serpentine endemism occur in New Caledonia and Cuba, with 3178 and 920 endemic taxa, respectively, found solely on serpentine. Despite the patchy occurrence of serpentine in eastern North America from Québec and Newfoundland south to Alabama, only one taxon, Cerastium veluti- num var. villosissimum, has been broadly recognized as a serpentine endemic for the region. Based on reports in the literature, we suggest that Adiantum viridimontanum, Minuartia marcescens, and Symphyotrichum rhiannon be considered endemic to serpentine soils from the east coast of North America. Aspidotis densa, with several disjunct populations on and off serpentine in western North America, is known solely from serpentine soils where it occurs in eastern North America and should be consid- ered endemic to the substrate there. The geobotany of eastern North America in general is poorly understood, and additional taxonomic studies on the region’s unique geologic substrates will likely yield further edaphic endemics. Introduction Narrow endemism can result from any number of biological and en- vironmental interactions. However, within a regional climate, geological discontinuities, both topographic and geochemical, are the most common and striking infl uences of narrow endemism (Kruckeberg 1986, Kruck- eberg and Rabinowitz 1985). Among the endemic species resulting from geological discontinuities are edaphic endemics, those species restricted to chemically and/or physically unique soils (Rajakaruna and Boyd 2008). -
Vascular Plant Species with Documented Or Recorded Occurrence in Placer County
A PPENDIX II Vascular Plant Species with Documented or Reported Occurrence in Placer County APPENDIX II. Vascular Plant Species with Documented or Reported Occurrence in Placer County Family Scientific Name Common Name FERN AND FERN ALLIES Azollaceae Mosquito fern family Azolla filiculoides Pacific mosquito fern Dennstaedtiaceae Bracken family Pteridium aquilinum var.pubescens Bracken fern Dryopteridaceae Wood fern family Athyrium alpestre var. americanum Alpine lady fern Athyrium filix-femina var. cyclosorum Lady fern Cystopteris fragilis Fragile fern Polystichum imbricans ssp. curtum Cliff sword fern Polystichum imbricans ssp. imbricans Imbricate sword fern Polystichum kruckebergii Kruckeberg’s hollyfern Polystichum lonchitis Northern hollyfern Polystichum munitum Sword fern Equisetaceae Horsetail family Equisetum arvense Common horsetail Equisetum hyemale ssp. affine Scouring rush Equisetum laevigatum Smooth horsetail Isoetaceae Quillwort family Isoetes bolanderi Bolander’s quillwort Isoetes howellii Howell’s quillwort Isoetes orcuttii Orcutt’s quillwort Lycopodiaceae Club-moss family Lycopodiella inundata Bog club-moss Marsileaceae Marsilea family Marsilea vestita ssp. vestita Water clover Pilularia americana American pillwort Ophioglossaceae Adder’s-tongue family Botrychium multifidum Leathery grapefern Polypodiaceae Polypody family Polypodium hesperium Western polypody Pteridaceae Brake family Adiantum aleuticum Five-finger maidenhair Adiantum jordanii Common maidenhair fern Aspidotis densa Indian’s dream Cheilanthes cooperae Cooper’s -
Summer 2003 - 61 President’S Message
THE HARDY FERN FOUNDATION P.O. Box 166 Medina, WA 98039-0166 Web site: www.hardvferns.org The Hardy Fern Foundation was founded in 1989 to establish a comprehen¬ sive collection of the world’s hardy ferns for display, testing, evaluation, public education and introduction to the gardening and horticultural community. Many rare and unusual species, hybrids and varieties are being propagated from spores and tested in selected environments for their different degrees of hardiness and ornamental garden value. The primary fern display and test garden is located at, and in conjunction with, The Rhododendron Species Botanical Garden at the Weyerhaeuser Corpo¬ rate Headquarters, in Federal Way, Washington. Satellite fern gardens are at the Stephen Austin Arboretum, Nacogdoches, Texas, Birmingham Botanical Gardens, Birmingham, Alabama, California State University at Sacramento, Sacramento, California, Coastal Maine Botanical Garden, Boothbay, Maine, Dallas Arboretum, Dallas, Texas, Denver Botanic Gardens. Denver, Colorado, Georgeson Botanical Garden, University of Alaska, Fairbanks, Alaska, Harry P. Leu Garden, Orlando, Florida, Inniswood Metro Gardens, Columbus, Ohio, Lewis Ginter Botanical Garden, Richmond, Virginia, New York Botanical Garden, Bronx, New York, and Strybing Arboretum, San Francisco, California. The fern display gardens are at Bainbridge Island Library, Bainbridge Island, WA, Lakewold, Tacoma, Washington, Les Jardins de Metis, Quebec, Canada, University of Northern Colorado, Greeley, Colorado, and Whitehall Historic Home and Garden, Louisville, KY. Hardy Fern Foundation members participate in a spore exchange, receive a quarterly newsletter and have first access to ferns as they are ready for distribution. Cover Design by Willanna Bradner HARDY FERN FOUNDATION QUARTERLY THE HARDY FERN FOUNDATION QUARTERLY Volume 13 No.