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Monograph on Spanish Palaeozoic Crinoidea

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Monograph on Spanish Palaeozoic Crinoidea A Monograph on Spanish Palaeozoic Crinoidea A. Breimer (Pre-issued 9-5-1962) Contents page Abstract 4 Introduction 7 Descriptions 9 Camerata 9 Fam. Reteocrinidae 9 9 Diamenocrinus spec. Fain. Dimerocrinitidae 11 Pterinocrinus decembrachiatus spec, nov 12 Macarocrinus ? 13 spec. Griphocrinus ovetensis spec, nov 15 Fam. Orthocrinidae 18 Orthocrinus planus nomen dubium 19 Orthocrinus robustus spec, nov 20 Orthocrinus 21 elongatus spec, nov Orthocrinus 23 spec. nov. (?) Fam. Periechocrinidae 26 Pradocrinus Baylii De Verneuil 1850 27 1918 Lenneocrinus cirratus Jaekel 30 Lenneocrinus ventanillensis spec, nov 30 Zeiller Pyxidocrinus prumiensis (Wirtgen & 1855) ... 35 Pyxidocrinus collensis spec, nov 35 Pyxidocrinus latus spec, nov 40 nov. comb. 43 Pyxidocrinus San-Migueli (Astre 1925) ... comb. Pyxidocrinus ? bifrons (W. E. Schmidt 1932) nov. 44 Pithocrinus Cooperi Kirk 1945 46 Pithocrinus ovatus spec, nov 47 Pithocrinus spinosus spec, nov 54 Pithocrinus Waliszewskii (Oehlert 1896) 57 59 Stamnocrinus gen. nov Stamnocrinus intrastigmatus (W. E. Schmidt 1932) nov. comb. 59 Stamnocrinus devonicus (Springer 1911) nov. comb. ... 64 Stamnocrinus 1 spec. 65 Stamnocrinus spec. 2 65 Corocrinus ? grandosensis spec, nov 67 Gennaeocrinus cf. G. 69 spec. nyssa (Hall 1862) Aryballocrinus gen. nov 72 72 Aryballocrinus Whitei (Hall 1861) nov. comb comb. 73 Aryballocrinus tenuidiscus (Hall 1861) nov. ... Aryballocrinus awthornsensis (J. Wright 1955) nov. comb. 73 Aryballocrinus Sampsoni (Miller & Gurley 1896) nov. comb. 73 Aryballocrinus parvus (Wachsm. & Spr. 1890) nov. comb. 74 Aryballocrinus spec. (Laudon & Severson 1953) nov. comb. 74 2 page Fam. Paragaricocrinidae 75 Iberocrinus multibrac hiatus Sieverts Doreck 1951 .... 75 Fam. Actinocrinidae 78 Nunnacrinus ? stellaris & Le Hon 78 (Konlnck 1854) ... Fam. Coelocrinidae 78 78 Aorocrinus spec. Fam. Amphoracrinidae 79 Pimlicocrinus latus J. Wright 1943 80 1 Pimlicocrinus spec. 82 83 Pimlicocrinus spec. 2 Fam. Hexacrinitidae 84 Platyhexacrinus Kegeli W. E. Schmidt 1932 84 Fam. Polypeltidae 85 Trybliocrinus Flatheanus Geinitz 1867 86 Fam. Hapalocrimdae 117 Canlharocrinus gen. nov. 117 Cantharocrinus minor spec, nov 117 Cantharocrinus simplex spec, nov 120 nodosus 122 cf. Culicocrinus (Wirtgen & Zeiller 1854) ... Fam. Platycrinidae 124 Oenochoacrinus 124 gen. nov Oenochoacrinus princeps spec, nov 124 Oenochoacrinus 127 pileatus spec, nov Oenochoacrinus scaber spec, nov 129 131 Oenochoacrinus ? spec Platycrinus spec. ex gr. bollandensis Wright 1938 . 131 Pleurocrinus ex 1938 . 133 spec. gr. coplowensis Wright . Inadunata 145 Fam. Iocrinidae 145 Myelodactylus spec. 145 Fam. Synbathocrinidae 145 Oehlert nomen dubium 145 Storthingocrinus Haugi 1896, . 145 Storthingocrinus labiatus W. E. Schmidt 1932, nomen dubium . Fam. Ampheristocrinidae 146 Vasocrinus valens Lyon 1857 147 Vasocrinus turbinatus Kirk 1929 148 Vasocrinus stellaris (Schultze 1867) 149 Vasocrinus cf. V. 1857 spec. sculptus Lyon 151 Fam. Barycrinidae 153 153 Situlacrinus gen. nov 154 Situlacrinus costatus spec, nov Fam. Cyathocrinidae 158 158 Cyalhocrinites spec Fam. Codiacrinidae 159 159 Codiacrinus spec Fam. Cupressocrinitidae 160 Cupressocrinites Sampelayoi (Almela & Revilla 1950) nov. comb. 160 Cupressocrinites inflatus Schultze 1867 163 3 page Cupressocrinites spec. aff. C. Townsendi (König 1825) . 164 cf. C. Schlotheimi 166 Cupressocrinites spec. (Steininger 1831) . 168 Cupressocrinites spec. Fam. Dendrocrinidae 168 168 Bactrocrinites spec Lasiocrinus ? 169 spec Fam. Poteriocrinidae 170 cf. C. Trautschold 1867 . Cromyocrinus spec. simplex 170 Paradelocrinus spec. 172 Conclusions 173 General remarks 173 Stratigraphical and palaeogeographical remarks 174 References 186 Plates Abstract has the aim of this of the It been study to give a comprehensive description important crinoid fauna's of the Palaeozoic core of North Western Spain (provinces Palencia, León and Asturias). This was opportune since fine collections of crinoids had been made during the years 1955—1960 by students of Leiden University (Holland). Moreover, existing collections of Spanish crinoids have not recieved hitherto due attention. Up to crinoids from the present day only ten species of Palaeozoic have been known Spain. Five new genera and sixteen new species of Spanish crinoids are now described. Fourteen described for the first time in either from previously genera are reported to occur Spain, well known allow new species or from species not yet sufficiently to specific arrangement. Six previously described species are recorded for the first time from Spain. So the total been two number of known Spanish crinoid species has now raised to fifty six (thirty five Emsian nine Couvinian six Givetian genera): twenty species, species, species, one Fransnian species, one Visean species, four Namurian species and four Moscovian species. The other species, represented by museum specimens only, are of uncertain Devonian or Carboniferous age. than whereas flexibles Camerates are far more numerous inadunates, are not known with any certainty. Camerates include thirty eight species (twenty two genera); inadunates eightteen species (ten genera). Among camerates only seven species belong to the diplobathrids; stratigraphically they seem to be restricted to the Emsian. Pterinocrinus decembrachiatus, Griphocrinus ovetensis, Orthocrinus robustus and Orthocrinus elongatus are described as new species of The Macarocrinus and diplobathrids. genera Diamenocrinus, Pterinocrinus, Griphocrinus are recorded for the first time from Spain. Orthocrinus was already known. W. E. Schmidt's species Orthocrinus planus is regarded a nomen dubium, since only poor fragments are assignable to the species. which Camerates further include thirty one species of monobathrids, among perie- chocrinids (fourteen species) and hapalocrinids/platycrinids (nine species) are of special importance. Stratigraphical distribution of the Spanish monobathrids is from the Lower Devonian to Upper Carboniferous. Among periechocrinids (restricted to Devonian strata) Pradocrinus is held as an with the only species P. de 1850 as the independent genus Baylii Verneuil, type-species. The genus is only known to occur in Spain. The available generic names Lenneocrinus of Lenneocrinus is and Pyxidocrinus were used for assignment Spanish species. now definitely erected with L. cirratus Jaekel, 1918 as the type-species. A diagnosis has been from the Frasnian given on p. 29. The genus is first reported to occur in Spain new species L. ventanillensis. Pyxidocrinus was proposed as a conditional name but is now erected as with Actinocrinus as and Muller the author. A genus prumiensis type-species J. as collensis and P. latus referred it diagnosis has been given on p. 35. P. are to as new species. P. San-Migueli (Astre, 1925), formerly referred to Periechocrinus and Pithocrinus and P. bifrons (W. E. Schmidt, 1932, formerly referred to Megistocrinus and Pithocrinus been ranked under has German have now Pyxidocrinus. Although Pyxidocrinus a species the it is it is as type, essentially a Spanish genus. Strangely enough only now reported The Pithocrinus with P. for the first time as occurring in Spain. genus Kirk, 1945, Cooperi Kirk, 1945 as the type-species has been emended so as to include forms with globose dorsal cups, anal tube. The a variable number of free arms per ray and a stout subcentral arms are described for the first time. For emended diagnosis see p. 46. Although Pithocrinus has an American type it is essentially a Spanish genus. P. ovatus and P. spinosus are referred Megisto-this P. Waliszewskii Oehlert, 1896, formerly referred to genus as new species, to Sacco-crinus is kept within it, but P. intrastigmatus Schmidt, 1932, formerly referred to the of the Stamnocrinus crinus is excluded and used as type-species new genus diagnosis see p. 59 which is believed to include Dorycrinus devonicus Springer, 1911 and two more definite Spanish species, not yet sufficiently well known so as to allow description. Stamnocrinus is restricted to Devonian strata. The new Emsian species Corocrinus ? characters grandosensis is provisionally referred to Corocrinus because it possesses unknown from Gennaeocrinus is first recorded from from up to now that genus. Spain a species G. very similar to nyssa. 5 of The study the important Devonian periechocrinid fauna has revealed that no Devonian forms can be assigned to the type genus Periechocrinus, which genus must be of Silurian A of Lower Carboniferous and exclusively age. group Mississippian species, hitherto assigned to Periechocrinus, both for morphological as for stratigraphical reasons cannot belong to Periechocrinus nor to any of the Devonian periechocrinid genera. The new genus Aryballocrinus is erected for them with Periechocrinus ? Whitei Hall, 1861 the A for this is 72. The includes as type-species. diagnosis genus given on p. genus six species, four of which were formerly referred with doubt to Periechocrinus: Aryballocrinus Whitei(Hall, 1861), Aryballocrinus tenuidiscus (Hall, 1861), Aryballocrinus awthornsensis and 1 Further (J. Wright, 1955) Aryballocrinus spec. (Laudon & Severson, 1953). are included Aryballocrinus Sampsoni Miller & Gurley, 1896, formerly referred to Corocrinus and Aryballocrinus parvus Wachsmuth & Springer, 1890, formerly referred to Megistocrinus Other monobathrids include specimens of Iberocrinus multibrac hiatus Sieverts Doreck, 1951, which species proved to occur in the Moscovian of Spain. Nunnacrinus ? stellaris first from the Namurian of The Pimlicocrinus is
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