20 © Sahlbergia Vol. 12: 20–32, 2007

Checklist of Finnish flies: superfamilies and (Dip- tera: ) Kaj Winqvist & Jere Kahanpää

Winqvist, K. & Kahanpää, J. 2007: Checklist of Finnish flies: superfamilies Tephritoidea and Sciomyzoidea (Diptera: Brachycera). — Sahlbergia 12:20-32, Helsinki, , ISSN 1237-3273.

Another part of the updated checklist of Finnish flies is presented. part covers the families , , , , , Ulididae, , , Heterocheilidae, Phaeomyii- dae, and . Eight are recorded from Finland for the first time. The following ten species have been erroneously reported from Finland and are here deleted from the Finnish checklist: Chaetolonchaea das- yops (Meigen, 1826), Earomyia crystallophila (Becker, 1895), Lonchaea hirti- ceps Zetterstedt, 1837, Lonchaea laticornis Meigen, 1826, lundbecki (Duda, 1924), achyrophori (Loew, 1869), Campiglossa irrorata (Fallén, 1814), Campiglossa tessellata (Loew, 1844), sororcula (Wie- demann, 1830) and nigricauda (Loew, 1856). The Finnish records of Lonchaeidae: Lonchaea bruggeri Morge, Lonchaea contigua Collin, Lonchaea difficilis Hackman and Piophilidae: Allopiophila dudai (Frey) are considered dubious. The total number of species of Tephritoidea and Sciomyzoidea found from Finland is now 262.

Kaj Winqvist, Zoological Museum, University of Turku, FI-20014 Turku, Finland. Email: [email protected]

Jere Kahanpää, Finnish Environment Institute, P.O. Box 140, FI-00251 Helsinki, Finland. Email: kahanpaa@iki.fi

Introduction new millennium there was no concentrated The last complete checklist of Finnish Dipte- Finnish effort to study just these particular ra was published in Hackman (1980a, 1980b). groups. Consequently, before our work the Recent checklists of Finnish species have level of knowledge on Finnish fauna in these been published for ‘lower Brachycera’ i.e. Xy- families was very heterogeneous particularly lophagidae – by Kahanpää & Win- with regard to Tephritoidea. qvist (2005), by Kahanpää & In Tephritidae the monograph by Merz Grichanov (2004) and Syrphidae by Haarto & (1994) covers not only the Central European Kerppola (2004). This paper presents a new fauna but also the Northern European species. checklist of the 151 Finnish species in the su- This work proved very helpful in updating the perfamily Tephritoidea and 111 species in the list of FinnishTephritidae. North European superfamily Sciomyzoidea. Pallopteridae are adequately covered by An- The superfamilies Tephritoidea and Sci- dersson (1990), Morge (1963) and Rotheray omyzoidea have not exactly been neglected (1999). Both Finnish species of Platystoma- in Finland, but before the beginning of the tidae and most ulidids are included in Rich- Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea 21 ter (1989a, 1989b) keys for European part tion of the Finnish dryomyzids is covered by of Russia. An important paper on the ulidid Ozerov (1987) and Stackelberg (1988). The Homalocephala was published by An- Finnish fauna of superfamily Sciomyzoidea is dersson (1991). well-known and only a few additions can be Yet, Tephritoidea includes two problema- expected. tic families where the taxonomical situation and existing literature are far less comprising. Material and methods In Piophilidae, the most important key to be The material and methods used were docu- used is still the revision of world Piophilidae mented in the previous part of the checklist by McAlpine (1977). This family includes a published by the authors (Kahanpää & Win- number of poorly known and recently descri- qvist 2005). The majority of examined speci- bed arctic and boreal species, some of which mens are stored in the Diptera Fennica collec- may be present in Finland. One such new spe- tion of the Zoological Museum, University of cies was found during the course of this work Helsinki (MZH). and it will be described in a separate article by The material in the collections of Winqvist & Kahanpää (unpublished). the Department of Applied Biology in Viik- The situation was a bit better in Loncha- ki, now part of MZH, Zoological Museum of eidae, where Hackman (1956) had conducted Turku and Natural History and Conservation a good basic study of the Finnish species and Society in Forssa were gone through. Also his work was continued by Nuorteva (1967). several individual collections (coll. Haarto in Also Kovalev (1984) made some important Mietoinen perhaps being the most important) additions to the North European lonchaeids, were checked. Malaise trap and yellow tray but the most helpful contribution in this fami- material caught by Metsähallitus and SYKE ly came from Scotland, where MacGowan has in 2005 were also included in the study. compiled all the existing knowledge of Euro- Species are listed in alphabetic order un- pean lonchaeids in his draft key. Lonchaeidae der the genus name. Additional names used is however a poorly known and taxonomically for the same species are listed under the valid challenging family, so there will certainly be name. Synonyms are listed here only if they future changes in the Finnish list of Loncha- have been used in major Finnish papers, es- eidae. pecially in the previous checklists. All names The second superfamily, Sciomyzoidea, used in the previous checklists are included. has received more attention in Northern Euro- Preoccupied names (preocc.) and emenda- pe during the last decades. Rozkošný’s world tions (emend.) are labelled accordingly. Mi- catalogue of Sciomyzidae and his book on the sidentifications (misid.) are listed only if the Fennoscandian fauna (including Phaeomyii- species was reported as new for Finland under dae) are slightly out of date but still very use- the given name. In this case the author name ful reference works (Rozkošný 1984, 1995). and year refers to the publification of the Fin- Pont (2002) provides keys and descriptions nish record. Species with notes in Results are of all European species of Sepsidae. During marked with an asterisk (*) after the species these two “Fauna Entomologica Scandinavi- name. ca”-projects the Finnish museum material of Sciomyzidae and Sepsidae had already been Results checked. The world sepsid fauna has recently A total of 151 tephritoid and 111 sciomy- been catalogued by Ozerov (2005). Identifica- zoid flies have been found in Finland. Table 22 Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea

1 shows the total number of Finnish species Hackman (1956). Hackman’s specimens for each family in the current checklist and from Karislojo and Utsjoki were reassig- in previously published works (Frey 1941, ned by Morge (1959a) as paratypes of D. Hackman 1980b). Eight species are recorded perpropinquus Morge and D. trichoster- from Finland for the first time. The follo- nalis Morge respectively. A single female wing species have been erroneously reported of D. occultus Collin (sensu Morge) has from Finland - but not necessarily included since been found in Finland: 1 ♀, Ab: in previous checklists - and are here deleted Nauvo (=Nagu), Högsar, 7.-18.vi.1961, from the list of Finnish species: Lonchaeidae: leg. A. Nordman, in MZH. Chaetolonchaea dasyops (Meigen), Earo- Chaetolonchaea dasyops (Meigen): Deleted myia crystallophila (Becker), Lonchaea hirti- from the Finnish checklist. All records ceps Zetterstedt, Lonchaea laticornis Meigen. are based on misidentified specimens of Piophilidae: Prochyliza lundbecki (Duda). C. pallipennis, q. v. Tephritidae: Campiglossa achyrophori (Loew) Chaetolonchaea pallipennis (Zetterstedt): Campiglossa irrorata (Fallén), Campiglossa New to Finland. Misidentified by Fin- tessellata (Loew), (Wie- nish authors (Hackman 1956) and others demann), Tephritis nigricauda (Loew). The as Chaetolonchaea dasyops (Meigen). circumstances leading these deletions are dis- Locally common in coastal forests in cussed in the notes, q. v. Three lonchaeid and the southern provinces Al, Ab, N and Ka. one piophilid species are listed as doubtful. Hackman (1956) mentions a specimen from Sa: Joutseno, but we have not been Notes able to confirm this record. Dasiops latiterebra (Czerny): Morge (1959b) Earomyia crystallophila (Becker): Omitted lists one specimen from Finland: 1 ♀, Ja- from the checklist. Finnish according to kobstad, Mus. Hels. No. 2549, Exemplar- Kovalev and Morge (1984), but never Nr. 1970. There is one female from Pietar- mentioned in Finnish checklists. All spe- saari (= Jakobstad) in MZH (16.vii.1955, cimens in MZH under names E. nigrovio- leg. R. Storå). Unfortunately it lacks the lacea (Frey in Lundström & Frey, 1913) serial number labels normally found on and E. parvicornis Czerny, 1934 nec. Zet- Dasiops specimens examined by Morge terstedt, 1847 are from the Kola Peninsula and it remains unclear whether it was ac- in northwestern Russia; this includes the tually seen by Morge. sole type specimen of nigroviolacea. Dasiops mucronatus Morge: New to Fin- Lonchaea albigena Collin: Not listed in land. There are three specimens in MZH: Hackman (1980b). Two females are kno- a female from Pietarsaari (= Jakobstad), wn from Finland: 1 ♀, Sa: Mikkeli (= St. 16.vii.1955, leg. R. Storå, a female from Michel), leg. Nordqvist; 1 ♀, Ta: Pirkkala Helsinki, leg. R. Frey, and a male from (= Birkkala), leg. R. Frey, ID 3517, labels Pakila, Helsinki, 6.iv.1950 (ex. ), ‘Lonchaea hirticeps Zett. ♀ det. Czerny’/ leg. L. Tiensuu. The female from Helsin- ’albigena Collin det. W. Hackman’. The ki (= Helsingfors) was identified as D. la- female from Otalampi mentioned by titerebra by Hackman (1956) and as D. Hackman (1956) was not found in muse- hennigi by Morge (1959b) um collections. Dasiops occultus Collin: Confirmed as Fin- Lonchaea bruggeri Morge: No males have nish. First reported from the country by been found in Finland. Matti Nuorteva Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea 23

(1967) reared at least two different spe- as stated by Kovalev and Morge (1984)). cies of the ‘watsoni group’ from spruce Two Finnish females have been tentative- stumps. Kovalev (1984) notes that L. ly identified as this species, but their true bruggeri is abundant in Northern Rus- identity could not be confirmed. 1 ♀ Tb: sia and he assumes that at least a part of Saarijärvi, leg. Woldstedt; 1 ♀ Ks: Kuusa- Nuorteva’s material belongs to this spe- mo, 3.vii.1958, leg. M. Nuorteva, ‘L. dif- cies. We failed to locate these females and ficilis Hack. ? Det. J. P. McAlpine 1966’. could not check the identifications. A few Hackman’s (1956) illustration of the ovi- rather worn females in MZH have later positor is based on the first specimen. The been tentatively placed in this species, third female in MZH (1 ♀ Ks: Kuusamo, but as long as no male is known from the leg. R. Frey) is from Paanajärvi National country, we consider all Finnish records Park. Like L. contigua, this species must doubtful. be considered doubtful in Finland until a Lonchaea caledonica MacGowan & Rothe- male is found. ray: Two specimens in MZH, identified Lonchaea freyi Czerny: The male from Kuo- by MacGowan: 1 ♀ N: Hanko, Tvärmin- pio mentioned by Hackman (1956) is still ne, leg. Wegelius; 1 ♂ N: Espoo, leg. P. the only specimen found in Finland; the Nuorteva (MacGowan & Rotheray 2000). type locality (Salmi) is in Russian Care- Hackman’s figures of male genitalia for lia. We have compared the male with the Lonchaea laticornis (Meigen) are based female holotype of L. freyi and consider on Nuorteva’s male and they belong to L. them conspecific. caledonica. Lonchaea hackmani Kovalev: First reported Lonchaea contigua Collin: No males have from Finland in the description of the spe- been found in Finland. The female men- cies (Kovalev 1981). tioned by Hackman (1956) is headless Lonchaea hirticeps Zetterstedt: Deleted from and unidentifiable. Two females collected the Finnish checklist. Hackman (1956) after Hackman’s revision may belong to lists one female specimen under this this species: 1 ♀ Om: Pietarsaari (705:25), name. The specimen was found in MZH 1.vii.1959, leg, R. Storå, in MZH; 1 ♀ (Al: Saltvik, leg. R. Frey, no. 3025-under- Ab: Korppoo, Västankvarn (66796:1957), lined). Unfortunately it is in rather a poor 21.vii.2004, leg. J. Kahanpää, in author’s condition and its head has been glued to collection. The presence of L. contigua in the thorax. Due to the damage it has little Finland must be considered doubtful until faunistic value. If the head and body were a male is found. correctly associated this specimen could Lonchaea defecta McAlpine: First reported not belong to L. hirticeps, but since this by Nuorteva (1967) from Finland on the can not be proved the true identity of the basis of material identified by McAlpine. specimen can not be resolved. We have examined Nuorteva’s specimens Lonchaea ipsiphaga McAlpine: Originally re- and agree with McAlpine’s identificati- ported from Finland by Nuorteva (1967) on. as L. maniola McAlpine on the basis of Lonchaea difficilis Hackman: No males have a single female, identified by McAlpine been found in Finland. The type locality himself. According to Kovalev (1975, is Lake Paanajärvi in Paanajärvi Natio- 1984) this specimen belongs most pro- nal Park, Russian Karelia (not Finland bably to L. ipsiphaga, since the latter spe- 24 Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea

cies is known from northwestern Russia veral localities in province Kb (Polevoi while L. maniola is apparently confined 2001) and province N (Nurmijärvi, Kil- to the Nearctic region. We have examin- javanlähteikkö (6711:3373), 12.vii.2005, ed the specimen in MZH and agree with leg. J. Kahanpää, ID jka-05-04003). Kovalev’s intepretatation. A key to the L. Allopiophila flavipes (Zetterstedt): Not listed corticis-group species with pale calypter in Hackman (1980b). There is one old fringe is provided by McAlpine & Morge Finnish specimen of this species in MZH (1970). (Kb: Eno, leg. Grönvik). Lonchaea laticornis Meigen: Deleted from the Allopiophila dudai (Frey): Doubtful record. Finnish checklist. All records are based Described by Frey (1930) from the Russi- on misidentification of various Lonchaea an Lapland, Kola peninsula. The original and Dasiops species including L. caledo- article did not contain any records from nica MacGowan & Rotheray. Finland and there have not been any sub- Lonchaea scutellaris Rondani: New to Fin- sequent published records. The previous land. 1 ♂ 1 ♀, Kl: Parikkala (6820:3630), Finnish checklists of the family (Frey 30.vi.1990, leg. M. Koponen, in the col- 1941, Hackman 1980b) include A. dudai lection of the Department of Applied Bio- without giving any details. No Finnish logy, Univ. of Helsinki. material was found in Finnish collections, Lonchaea subneatosa Kovalev: First reported but some specimens from MZH are on from Finland by Kovalev (1984). loan abroad and their labels and identifi- Lonchaea tarsata (Fallén): Confirmed as Fin- cation could not be checked. nish. 1 ♂, Al: Föglö, Bänö (6682:3143), Prochyliza lundbecki (Duda): Deleted from 7.vi.2002, leg. J. Kahanpää, in author’s the Finnish checklist. First mentioned as collection. Finnish by Frey (1941). The only Finnish Lonchaea ultima Collin: New to Finland. 1 ♂, specimen is a female of P. varipes (Mei- Al: Lemland, 12.vi.1962, leg. R. Tuomi- gen). There are a few correctly identified koski, in MZH; 1 ♀, Al: Lemland, Nåtö North Russian P. lundbecki specimens in (6680:3109), 7.vi.2002, leg. J. Kahanpää, MZH. in author’s collection. Campiglossa achyrophori (Loew): Deleted anderssoni Rotheray: A recently from the Finnish checklist. All the Finnish described species. Rotheray and McGo- specimens identified as C. achyrophori wan (1999) list a Finnish specimen col- (Loew) belong to C. guttella (Rondani). lected “near Lempola (P. Chandler)” (= C. achyrophori is restricted to elevated Ab: Lohja, Lempola or Ta: Lempäälä?) areas in central Europe (Merz 1994). as a paratype. Three additional specimens Campiglossa irrorata (Fallén): Deleted from were found in MZH: Al: Saltvik, leg. R. the Finnish checklist. Reported from “Ca- Frey, N: Helsinki, leg.L. Tiensuu and Om: relia” by W. Nylander and subsequently Pietarsaari, leg. R. Storå. included in previous Finnish checklists Palloptera marginata (Meigen): A recent (Frey 1941, Hackman 1980b). Nylander’s addition to the checklist, recorded from specimens are from northwestern Russia. one site in the province Ta: (Haarto et al. Campiglossa tessellata Loew: Deleted from 2002). the Finnish checklist. All Finnish mate- Palloptera venusta Loew: Another rather new rial previously identified as C. tessellata addition to the Finnish fauna, found at se- (Loew) belongs to C. difficilis (Hendel). Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea 25

Campiglossa solidaginis (White): A recent Silfverberg 1981). addition to the checklist, recorded from Tephritis fallax (Loew): First listed by Frey two localities in provinces Ta and Sa (Sö- (Frey 1941). Accidentally omitted in derman et al. 2007). Hackman’s list (Hackman 1980b, Silfver- Chetostoma stackelbergi (Rohdendorf): A re- berg 1981). cent addition to the checklist, recorded Tephritis nigricauda (Loew): See Tephritis sp. from five localities in provinces Ab, N and cf. rydeni Hering (q.v.). Ta (Söderman et al. 2007). plagiata (Dahlbom): A recent additi- Dioxyna sororcula (Wiedemann): Deleted on to the checklist, found only in province from the Finnish checklist. All the Fin- Al (Söderman et al. 2007). nish specimens identified as D. sororcu- cardui (Linnaeus): First recorded la (Wiedemann) are in fact D. bidentis from Finland in 1981 (Jansson & Linde- (Robineau-Desvoidy). D. sororcula is berg 1982). a widespread species in the tropical and (Hennig): New subtropical zones. In Europe it is found to Finland. 1 ♂ St: Säkylä (67755:32594), only in southern (Foote 1984, Merz 23.vi.2005, leg. I. Kakko; 2 ♂ 1 ♀, Ta: 1994). Somero, 16.vi.2005, leg. I. Kakko; 1 ♀ Euphranta (Loew): A recent addi- Ab: Karjalohja, Tammisto (66840:33199), tion to the checklist, recorded from three 18.vi.2006, leg. J. Kahanpää, ID jka06- sites in provinces Sa and Ok (Söderman 00690. P. tryptetoptera was erroneously et al. 2007). synonymised with the nearctic P. cortica- nebulosa (Wiedemann): A recent addi- lis (Loew) by Steyskal (1962) (Krivoshei- tion to the checklist based on specimens na & Krivosheina 1997). from provinces Al and Kb in MZH (Sö- omissa (Meigen): New to Finland. derman et al. 2007). Recorded from the coastal provinces Ab, superciliata (Frey): Another re- N, Ka, St, Oa, Om and Oba. This species cent addition to the checklist, recorded was previously known as M. obscuripes from one site in province Ta (Söderman (Loew) in Finland following a misidenti- et al. 2007). Previously misidentified as fication by Krogerus (1932). zoe (Haarto et al. 2002). pectorosa (Hendel): New to Tephritis sp. cf. rydeni Hering: This proble- Finland. Ab: Mietoinen, Perkko 25.- matic taxon is widespread in southern and 28.vii.2002, leg. A. Haarto. Collected central Finland (Söderman et al. 2007). with a light trap. It is close to T. leontodontis (Degeer) in sebezhica Przhiboro: New to Fin- wing pattern (Fig. 1) and colour, but the land. 1 ♂ 2 ♀, Tb: Jyväskylä, Eerolan- ovipositor tip is T. neesii-like. T. rydeni lahti (69055:34338), 22.vii.-10.viii.2004, Hering, 1956 is sometimes considered leg. J. Kahanpää; 2 ♂ 2 ♀, Oba: Liminka, a synonym of T. neesii. Its true status Liminganlahti (719601:342284), 30.vii.- should be reassessed and the Finnish ma- 1.ix.2005, leg. J. Kahanpää. A recently terial compared with the types of T. ryde- described species previously known only ni. In Merz (1994) this form keys out as T. from the type locality, Lake Anninskoe in dioscurea (Loew) or T. tanaceti Hering. Pskov Province, Russia (Przhiboro 2001). Finnish records of Tephritis nigricauda Collected with Malaise traps from chan- (Loew) belong to this taxon (Frey 1943, nels with slowly flowing water and well- 26 Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea

developed shore vegetation (Phragmites Kiado, Budapest & Elsevier, Amsterdam. australis, Potentilla palustris) on a parti- Frey, R. 1941: Diptera Brachycera (excl. Musci- ally floating layer of Sphagnum mosses. dae, ). — In: Anonymous (Ed.), Orygma luctuosum Meigen: In MZH there is Enumeratio Insectorum Fenniae. VI. Diptera: a single specimen of this species allegedly 1-31. Helsingin hyönteisvaihtoyhdistys r.y. Frey, R. 1943: [Föreningsmeddelanden]. — Notu- taken from Rovaniemi by W. Hellén. This lae Entomol. 23: 163. record is highly suspect: O. luctuosum is Haarto, A. & Kerppola, S. 2004: Checklist of Fin- a maritime species and it is unlikely that nish Hoverflies (Diptera, Syrphidae). — Sahl- it could have been recorded so far from bergia 9: 147-164. the seashore. Hellén had collected a long Haarto, A., Mukkala, V. & Koponen, S. 2002: Tut- series of O. luctuosum from the Petsamo kimus Rekijokilaakson hyönteisistä ja hämä- on the arctic coast a few days earlier. It häkkieläimistä. — Lounais-Suomen ympäris- is very probable that he either mislabeled tökeskuksen monistesarja 2002: 1-58. the specimen or inadvertently transferred Hackman, W. 1956: The Lonchaeidae (Dipt.) of it to Rovaniemi, some 400 km from the Eastern Fennoscandia. — Notulae Entomol. arctic coastline. 36: 89-115. Hackman, W. 1980a: A check list of the Finnish Acknowledgements Diptera I. and Brachycera (s. str.). — Notulae Entomol. 60: 17-48. We thank Prof. B. Merz (Muséum d’histoire Hackman, W. 1980b: A check list of the Finnish naturelle, Genève) and Dr. I. Macgowan Diptera II. Cyclorrhapha. — Notulae Ento- (Scottish Natural Heritage) for the time they mol. 60: 117-162. spent studying Finnish specimens. This pro- Jansson, A. & Lindeberg, B. 1982: A spectacular ject would have been impossible without the tephritid fly (Diptera) new to Finland. — No- hospitality of Dr. Pekka Vilkamaa, Dr. Pasi tulae Entomol. 62: 151-152. Sihvonen and Mr. Juha Laiho at the Zoolo- Kahanpää, J. & Grichanov, I. Y. 2004: A check- gical Museum of the University of Helsinki list of Finnish long-legged flies (Diptera: (MZH). The University of Turku provided Dolichopodidae). — Int. J. Dipterol. Res. 15: working facilities and space for a reference 57-62. Kahanpää, J. & Winqvist, K. 2005: Checklist collection for the senior author during three of Finnish flies: families - autumns 2004-2006. We are also grateful for Microphoridae. — Sahlbergia 10: 10-27. the monetary support by Ministry of Envi- Kovalev, V. G. 1975: Palaearctic species of the ronment through the Finnish Expert Group of Lonchaea corticis group (Diptera, Lonchaei- Diptera (Dipteratyöryhmä). dae). — Zool. Zh. 54: 1648-1655. (In Russi- an) References Kovalev, V. G. 1981: On European species of the Andersson, H. 1990: De svenska prickflugorna group Lonchaea peregrina (Diptera, Loncha- (Diptera, Pallopteridae), med typdesignerin- eidae). — Zool. Zh. 60: 221-228. (In Russi- gar och nya synonymer. — Entomol. Tidskr. an) 111: 123-131. Kovalev, V. G. 1984: On Finnish and some other Andersson, H.: Revision of Swedish Homa- species of Lonchaea Fallén (Diptera, Loncha- locephala Zetterstedt (Diptera, Otitidae) Ent. eidae). — Ann. Entomol. Fenn. 50: 17-20. Tidskr. 112: 27-32. Kovalev, V. G. & Morge, G. 1984: Family Lon- Foote, R. H. 1984: Family Tephritidae. — In: chaeidae. — In: Sóos, Á. & Papp, L. (Eds.), Sóos, Á. & Papp, L. (Eds.), Catalogue of Pa- Catalogue of Palaearctic Diptera 9: 247-259. laearctic Diptera Vol. 9: 66-149. Akademiai Akadémiai Kiadó, Budapest. Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea 27

Krivosheina, N. P. & Krivosheina, M. G. 1997: Brill, Leiden, New York, Köln. 221 pp. Review of Palaearctic species of the genus Przhiboro, A. A. 2001: A new species of the snail- Pseudotephritis Johnson (Diptera, Otitidae). killing fly genus Sciomyza (Diptera: Sciomy- — Entomol. Obozr. 76: 671-679. zidae), and a list of Sciomyzidae collected at Krogerus, R. 1932: Über die Ökologie und Ver- Anniskoe and Anisimovo Lakes, Pskov Pro- breitung der Arthropoden der Triebsandge- vince. — Zoosyst. Rossica 10: 183-188. biete an den Küsten Finnlands. — Acta Zoo- Richter, V. A. 1989: Platystomatidae. — In: Bei- logica Fennica 12: 1-308. Bienko, G. Y. & Steyskal, G. C. (Eds.), Keys MacGowan, I. & Rotheray, G. E. R. 2000: New to the of the European part of the species, additions and possible deletions to USSR. Diptera and Siphanoptera. Volume the British Lonchaea Fallen (Diptera, Lon- 5(2): 194-196. Smithsonian Institution Lib- chaeidae). — Dipterists Digest 7: 37-49. raries and the National Science Foundation, McAlpine, J. F. & Morge, G. 1970: The identity, Washington, D. C. distribution, and biology of Lonchaea zetter- Richter, V. A. 1989: Otitidae & . — In: stedti with notes on related species (Diptera: Bei-Bienko, G. Y. & Steyskal, G. C. (Eds.), Lonchaeidae). — The Canadian Entomolo- Keys to the Insects of the European part of gist 102: 1559-1566. the USSR. Diptera and Siphanoptera. Volume Merz, B. 1994: Diptera, Tephritidae. — Insecta 5(2): 197-211. Smithsonian Institution Lib- Helvetica, Fauna 10. Schweizerischen Ento- raries and the National Science Foundation, mologischen Gesellschaft, Genève. 198 pp. Washington, D. C. Morge, G. 1959a: Monographie der palaearkti- Rotheray, G. & McGowan, I. 1999: Palloptera schen Lonchaeidae (Diptera). — Beiträge zur anderssoni sp. n. from Scotland and Finland Entomologie 9: 323-371. (Diptera: Pallopteridae). — Br. J. Ent. Nat. Morge, G. 1959b: Monographie der palaearkti- Hist. 11: 175-179. schen Lonchaeidae (Diptera). — Beiträge zur Rotheray, G. & McGowan, I. 1999: Palloptera Entomologie 9: 1-92. anderssoni sp. n. from Scotland and Finland Morge, G. 1963: Die Lonchaeidae und Pallopte- (Diptera: Pallopteridae). — Br. J. Ent. Nat. ridae Österreichs und der angrenzenden Ge- Hist. 11: 175-179. biete. 1. Teil: Die Lonchaeidae. — Naturk. Rozkošný, R. 1984: The Sciomyzidae (Diptera) Jb. Stadt Linz : 123-312. of Fennoscandia and . — Fauna Ent. Nuorteva, M. 1967: On the habitats of some Lon- Scand. 14. E. J. Brill, Leiden. 224 pp. chaea species in Fennoscandia (Dipt., Lon- Rozkošný, R. 1995: World distribution of Scio- chaeidae). — Ann. Entomol. Fenn. 33: 118- myzidae based on the list of species (Dipte- 121. ra). — Studia dipterologica 2: 221-238. Ozerov, A. L. 1987: Dipterans of the family Dryo- Silfverberg, H. 1981: Additions to the Finnish myzidae in the fauna of the USSR. — Bull. Insect fauna during the years 1976-1980. — Moscow Soc. Nat., Biol. Ser. 92: 36-42. (In Notulae Entomol. 61: 45-61. Russian) Steyskal, G. C. 1962: The American species of the Ozerov, A. L. 2005: World catalogue of the family genera Melieria and Pseudotephritis (Dipte- Sepsidae (Insecta: Diptera). — Zoologiches- ra: Otitidae). — Papers of the Michigan Aca- kie issledovania 8:1-74 demy of Science, Arts, and Letters 47: 247- Polevoi, A. 2001: List of Diptera, collected within 266. Koitajoki area during the study period (1993- Söderman, G., Winqvist, K., Albrecht, A. & Ka- 1996). — Metsähallituksen luonnonsuojelu- hanpää, J. 2007: Suomen hedelmäkärpästen julkaisuja, Sarja A 131: 191-198. (Diptera: Tephritidae) biologia, levinneisyys Pont, A. C. & Meier, R. 2002: The Sepsidae (Dip- ja uhanalaisuus. — Sahlbergia 13. In press. tera) of Europe. — Fauna Ent. Scand. 37. 28 Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea

APPENDIX A: Checklist of Finnish Diptera, = lauta Collin, 1953 superfamilies Tephritoidea and Sciomyzoidea ? = britteni Collin, 1953 defecta McAlpine, 1964 Order DIPTERA deutschi Zetterstedt, 1838 Suborder BRACHYCERA = sarekensis Frey, 1916 Infraorder freyi Czerny, 1934 * Superfamily TEPHRITOIDEA fugax Becker, 1895 = cariecola Czerny, 1934 Family LONCHAEIDAE hackmani Kovalev, 1981 * subfamily Dasiopinae = peregrina auct. nec Becer, 1895 p.p. tribus Dasiopini ipsiphaga McAlpine, 1964 * Dasiops Rondani, 1856 = maniola auct. nec McAlpine, 1964 = Psilolonchaea Czerny, 1934 limatula Collin, 1953 appendiculatus Morge, 1959 = flavidipennis auct. nec Zetterstedt, 1847 = facialis Hackman, 1956 nec Collin, 1953 p.p. nitens (Bigot, 1885) facialis Collin, 1953 = krogerusi Czerny, 1934 mucronatus Morge, 1959 * palposa Zetterstedt, 1847 =hennigi Morge, 1959 misid. patens Collin, 1953 =latiterebra Hackman 1956 nec. Czerny, 1934 * ragnari Hackman, 1956 occultus Collin, 1953 * scutellaris Rondani, 1874 * perpropinquus Morge, 1959 sororcula Hackman, 1956 = occulta Hackman, 1956 nec Collin, 1953 p.p. stackelbergi Czerny, 1934 spatiosus (Becker, 1895) subneatosa Kovalev, 1974 * trichosternalis Morge, 1959 = peregrina auct. nec Becer, 1895 p.p. = occulta Hackman, 1956 nec Collin, 1953 p.p. sylvatica Beling, 1873 = facialis Hackman, 1956 nec Collin, 1953 p.p. = lucidiventris Becker, 1895 subfamily Earomyinae tarsata Fallén, 1820 * Chaetolonchaea Czerny, 1934 ultima Collin, 1953 * pallipennis (Zetterstedt, 1855) * zetterstedti Becker, 1902 = dasyops Hackman, 1956 nec (Meigen, 1826) Doubtful records in Lonchaeidae: Dasiops latiterebra Earomyia Zetterstedt, 1842 Czerny, 1934 *, Lonchaea bruggeri Morge, 1967 *, = Spermatolonchaea Hendel, 1932 Lonchaea contigua Collin, 1953 *, Lonchaea difficilis lonchaeoides Zetterstedt, 1848 Hackman, 1956 * schistopyga Collin, 1953 viridana (Meigen, 1826) Protearomyia McAlpine, 1962 Family PALLOPTERIDAE = Priscoearomyia Morge, 1963 subfamily Eurygnathomyiinae nigra (Meigen, 1826) Eurygnathomyia Czerny, 1904 subfamily Lonchaeinae bicolor (Zetterstedt, 1837) tribus Lonchaeini subfamily Pallopterinae Lonchaea Fallén, 1820 Palloptera Fallén, 1820 = Tricholonchaea Czerny, 1934 ambusta (Meigen, 1826) affinis Malloch, 1920 anderssoni Rotheray & MacGowan, 1999 * = laxa auct. nec. Collin, 1953 ephippium Zetterstedt, 1860 alpigena Collin, 1953 formosa Frey, 1930 albitarsis Zetterstedt, 1838 laetabilis Loew, 1873 caledonica MacGowan & Rotheray, 2000 * marginata (Meigen, 1826) * = laticornis Hackman, 1956 nec Meigen, 1826 = costalis Loew, 1873 p.p. * modesta (Meigen, 1826) chorea (Fabricius, 1781) = parallela Loew, 1858 collini Hackman, 1956 = umbellatarum Hackman, 1980 nec (Fabricius, corusca Czerny, 1934 1775) = alni Ringdahl, 1947 saltuum (Linnaeus, 1758) Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea 29

trimacula (Meigen, 1826) sg. Euphranta Loew, 1862 umbellatarum (Fabricius, 1775) connexa (Fabricius, 1794) = arcuata Hackman, 1980 nec (Fabricius, 1775) sg. Rhacochlaena Loew, 1862 usta (Meigen, 1826) toxoneura (Loew, 1846) * ustulata Fallén, 1820 tribus Carpomyini venusta Loew, 1858 * Rhagoletis Loew, 1862 = atriventris Ringdahl, 1947 alternata (Fallén, 1814) cerasi (Linnaeus, 1758) PIOPHILIDAE tribus Trypetini subfamily Piophilinae Acidia Robineau-Desvoidy, 1830 tribus Piophilini cognata (Wiedemann, 1817) Allopiophila Hendel, 1917 Walker, 1835 = Arctopiophila Duda, 1924 = Phagocarpus Rondani, 1970 = Boreopiophila Frey, 1930 purmunda (Harris, 1780) = Parapiophila McAlpine, 1977 = permundus misspelling calceata (Duda, 1924) Chetostoma Rondani, 1956 flavipes (Zetterstedt, 1847) * stackelbergi (Rohdendorf, 1955) * lonchaeoides (Zetterstedt, 1838) Cornutrypeta Han, Wang & Kim, 1993 luteata (Haliday, 1833) spinifrons (Schroeder, 1913) pectiniventris (Duda, 1924) superciliata (Frey, 1935) * tomentosa Frey, 1930 Euleia Walker, 1835 vulgaris (Fallén, 1820) = Cryptaciura Hendel, 1927 sp. aff. vulgaris heraclei (Linnaeus, 1758) Amphipogon Wahlberg, 1845 rotundiventris (Fallén, 1814) flavum (Zetterstedt, 1838) Rondani, 1856 = spectrum Wahlberg, 1845 lucida (Fallén, 1826) Mycetaulus Loew, 1845 Rondani, 1870 bipunctatus (Fallén, 1823) caesio (Harris, 1776) Fallén, 1810 Trypeta Meigen, 1803 casei (Linnaeus, 1758) artemisiae (Fabricius, 1794) Prochyliza Walker, 1849 immaculata (Macquart, 1835) = Liopiophila Duda, 1924 = hamifera Loew, 1846 nigrimana (Meigen, 1826) zoe Meigen, 1826 varipes (Meigen, 1830) subfamily = lundbecki misid. * tribus Dithrycini Pseudoseps Becker, 1902 Dithryca Rondani, 1856 signata (Fallén, 1820) gutturalis (Meigen, 1826) Stearibia Lioy, 1864 tribus Eutretini nigriceps (Meigen, 1826) Xanthomyia Phillips, 1923 = foveolata (Meigen, 1826) = Paracarphotricha Hendel, 1927 = coerulescens Zetterstedt, 1847 alpestris (Pokorny, 1887) Doubtful records in Piophilidae: Allopiophila dudai = pseudoradiata (Becker, 1900) (Frey, 1930) * tribus Myopitini Eurasimona Korneyev & White, 1991 Family PLATYSTOMATIDAE stigma (Loew, 1840) Platystoma Meigen, 1803 Urophora Robineau-Desvoidy, 1830 seminatione (Fabricius, 1775) = Euribia Meigen, 1800, suppr. Robineau-Desvoidy, 1930 aprica (Fallén, 1820) syngenesiae (Fabricius, 1781) cardui (Linnaeus, 1758) * cuspidata (Meigen, 1826) Family TEPHRITIDAE jaceana (Hering, 1935) subfamily solstitialis (Linnaeus, 1758) tribus Adramini stylata (Fabricius, 1775) Euphranta Loew, 1862 tribus Noeetini 30 Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea

Ensina Robineau-Desvoidy, 1830 Orellia Robineau-Desvoidy, 1830 sonchi (Linnaeus, 1767) falcata (Scopoli, 1763) Noeeta Robineau-Desvoidy, 1830 Terellia Robineau-Desvoidy, 1830 = Oplocheta Rondani, 1856 sg. Cerajocera Rondani, 1856 pupillata (Fallén, 1814) ceratocera (Hendel, 1913) tribus plagiata (Dahlbom, 1850) * Campiglossa Rondani, 1870 tussilaginis (Fabricius, 1775) absinthii (Fabricius, 1805) sg. Terellia Robineau-Desvoidy, 1830 = parvula (Loew, 1862) ruficauda (Fabricius, 1794) argyrocephala (Loew, 1844) serratulae (Linnaeus, 1758) difficilis (Hendel, 1927) winthemi (Meigen, 1830) = tessellata misid. * tribus Xyphosiini grandinata (Rondani, 1870) Xyphosia Robineau-Desvoidy, 1830 = borealis Portschincky, 1875 miliaria (Schrank, 1781) guttella (Rondani, 1870) tribus uncertain = achyrophori misid. * Acinia Robineau-Desvoidy, 1830 loewiana (Hendel, 1927) corniculata (Zetterstedt, 1819) plantaginis (Haliday, 1833) punctella (Fallén, 1814) Family ULIDIIDAE solidaginis (White, 1986) * subfamily Otitinae Dioxyna Frey, 1945 tribus Myennidini bidentis (Robineau-Desvoidy, 1830) Pseudotephritis Johnson, 1802 = sororcula misid. * trypetoptera (Hennig, 1939) * Heringina Aczél, 1940 = corticalis auct. nec. (Loew, 1873) guttata (Fallén, 1814) tribus Otitini Oxyna Robineau-Desvoidy, 1830 Macquart, 1835 flavipennis (Loew, 1844) = Fallén, 1810, preocc. nebulosa (Wiedemann, 1817) * urticae (Linnaeus, 1758) parietina (Linnaeus, 1758) Robineau-Desvoidy, 1830 Robineau-Desvoidy, 1830 frondescentiae (Linnaeus, 1758) marginata (Fallén, 1814) paludum (Fallén, 1820) Tephritis Latreille, 1804 palustris (Meigen, 1826) angustipennis (Loew, 1844) Melieria Robineau-Desvoidy, 1830 bardanae (Schrank, 1803) crassipennis (Fabricius, 1794) cometa (Loew, 1840) omissa (Meigen, 1826) * conura (Loew, 1844) = obscuripes misid. dilacerata (Loew, 1846) Tetanops Fallén, 1820 fallax (Loew, 1844) * sg. Eurycephalomyia Hendel, 1907 hyoscyami (Linnaeus, 1758) sintenisi Becker, 1909 leontodontis (De Geer, 1776) sg. Tetanops Fallén, 1820 neesii (Meigen, 1830) myopina Fallén, 1820 = nesii misspelling subfamily ruralis (Loew, 1844) tribus Seiopterini sp. cf. rydeni Hering, 1956 * Homalocephala Zetterstedt, 1838 = dioscurea misid. albitarsis Zetterstedt, 1838 = nigricauda misid. * = bipunctata (Loew, 1854) Schrank, 1795 angustata (Wahlberg, 1838) stellata (Fuessly, 1775) apicalis (Wahlberg, 1838) tribus Terellini bimaculata (Wahlberg, 1838) Chaetorellia Hendel, 1927 biseta Frey, 1908 jaceae (Robineau-Desvoidy, 1830) biumbrata (Wahlberg, 1838) Chaetostomella Hendel, 1927 = albitarsis auct. nec Zetterstedt, 1838 cylindrica (Robineau-Desvoidy, 1830) Kirby, 1817 = onotrophes (Loew, 1848) vibrans (Linnaeus, 1758) Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea 31

tribus Ulidiini alpina (Frey, 1930) Physiphora Fallén, 1810 argyra Verbeke, 1967 = Chrysomyza Fallén, 1817 brunnipes (Meigen, 1838) alceae (Preyssler, 1791) cinerella (Fallén, 1820) = demandata (Fabricius, 1798) dubia (Fallén, 1820) griseicollis Becker, 1900 Order DIPTERA = lapponica (Ringdahl, 1948) Suborder BRACHYCERA griseola (Fallén, 1820) Infraorder MUSCOMORPHA = dorsata misid. Superfamily SCIOMYZOIDEA hackmani Rozkošný, 1982 nana (Fallén, 1820) Family COELOPIDAE obscura (Ringdahl, 1948) Meigen, 1830 obtusa (Fallén, 1820) sg. Fucomyia Haliday, 1837 pallidiventris (Fallén, 1820) frigida (Fabricius, 1805) rozkosnyi Verbeke, 1967 = scutellaris misid. Family DRYOMYZIDAE schoenherri (Fallén, 1826) subfamily Dryomyzinae sordida (Hendel, 1902) Fallén, 1820 stackelbergi Elberg, 1965 decrepita Zetterstedt, 1838 stylifera Rozkošný, 1982 flaveola (Fabricius, 1794) ventralis (Fallén, 1820) Pseudoneuroctena Ozerov, 1987 Pteromicra Lioy, 1864 senilis (Zetterstedt, 1838) = Dichrochira Hendel, 1901 Neuroctena Rondani, 1868 angustipennis (Staeger, 1845) anilis (Fallén, 1820) = glabricula Frey, 1941 nec (Fallén, 1820) glabricula (Fallén, 1820) Family HETEROCHEILIDAE = nigrimana Meigen, 1830 Rondani, 1857 leucopeza (Meigen, 1838) buccata (Fallén, 1820) pectorosa (Hendel, 1902) * Sciomyza Fallén, 1820 Family PHAEOMYIIDAE = Bischofia Hendel, 1902 Pelidnoptera Rondani, 1856 dryomyzina Zetterstedt, 1846 = Phaeomyia Schiner 1862 sebezhica Przhiboro, 2001 * fuscipennis (Meigen, 1830) simplex Fallén, 1820 = fumipennis (Zetterstedt, 1846) Fallén, 1820 = leptiformis misid. pallidiventris Fallén, 1820 nigripennis (Fabricius, 1794) tribus Tetanocerini Antichaeta Haliday, 1838 Family SCIOMYZIDAE = Heteropteryx Hendel, 1902 subfamily Sciomyzinae analis (Meigen, 1830) tribus Sciomyzini atriseta (Loew, 1849) Colobaea Zetterstedt, 1837 brevipennis (Zetterstedt, 1846) = Ctenulus Rondani, 1856 nigra Karl, 1921 bifasciella (Fallén, 1820) = nigroaenea Frey, 1935 distincta (Meigen, 1830) Rondani, 1856 nigroaristata Rozkošný, 1984 marginata (Fabricius, 1775) pectoralis (Zetterstedt, 1847) = tristis (Harris, 1780) punctata Lundbeck, 1923 Rondani, 1868 Ditaeniella Sack, 1939 finlandica Verbeke, 1964 grisescens (Meigen, 1830) = obliterata auct. nec (Fabricius, 1805) Pherbellia Robineau-Desvoidy, 1830 Meigen, 1803 = Melina Robineau-Desvoidy, 1830 umbranum (Linnaeus, 1758) = Oxytaenia Sack, 1939, preocc. Ectinocera Zetterstedt, 1838 albocostata (Fallén, 1820) borealis Zetterstedt, 1838 32 Winqvist & Kahanpää: Tephritoidea and Sciomyzoidea

Elgiva Meigen, 1838 Family SEPSIDAE cucularia (Linnaeus, 1767) Meroplius Rondani, 1874 divisa (Loew, 1845) minutus (Wiedemann, 1830) solicita (Harris, 1780) = stercorarius (Robineau-Desvoidy, 1830) = rufa Panzer, 1798, preocc. Robineau-Desvoidy, 1830 = sundewalli Kloet & Hincks, 1945 nitidula (Fallén, 1820) Latreille, 1829 = cylindrica (Fabricius, 1794) preocc. = Lunigera Hendel, 1900 pectinulata Loew, 1873 chaerophylli (Fabricius, 1798) Ortalischema Frey, 1925 Hydromya Robineau-Desvoidy, 1830 albitarse (Zetterstedt, 1847) dorsalis (Fabricius, 1775) Pseudonemopoda Duda, 1926 Ilione Haliday in Curtis, 1837 speiseri Duda, 1926 = Knutsonia Verbeke, 1964 Robineau-Desvoidy, 1830 sg. Ilione Haliday in Curtis, 1837 = Pandora Haliday, 1833 albiseta (Scopoli, 1763) sphondylii (Schrank, 1803) sg. Tumidicerus Knutson & Berk, 1967 = scutellaris (Fallén, 1820) lineata (Fallén, 1820) Fallén, 1810 Limnia Robineau-Desvoidy, 1830 = Sepsidimorpha Frey, 1908 paludicola Elberg, 1965 biflexuosa Strobl, 1893 unguicornis (Scopoli, 1763) cynipsea (Linnaeus, 1758) Pherbina Robineau-Desvoidy, 1830 duplicata Haliday, 1838 coryleti (Scopoli, 1763) = pilipes van der Wulp, 1871 Enderlein, 1939 = pilipes Loew, 1873, preocc. zernyi (Mayer, 1953) flavimana Meigen, 1826 = punctata auct. nec Fabricius, 1794 = ruficornis Meigen, 1826 Hendel, 1900 = simplex Goetghebuer & Bastin, 1925 pallida (Fallén, 1820) = borealis Frey, 1925 striata (Meigen, 1830) fulgens Meigen, 1826 stroblii Hendel, 1900 = communis Frey, 1925 = fuscinervis auct. nec Zetterstedt, 1838 luteipes Melander & Spuler, 1917 Sepedon Latreille, 1804 = lamellifera Frey, 1925 sphegea (Fabricius, 1775) nigripes Meigen, 1826 spinipes (Scopoli, 1763) orthocnemis Frey, 1908 Duméril, 1800 punctum (Fabricius, 1794) arrogans Meigen, 1830 thoracica (Robineau-Desvoidy, 1830) elata (Fabricius, 1781) violacea Meigen, 1826 ferruginea Fallén, 1820 Robineau-Desvoidy, 1830 = brunnipennis Frey, 1924 = Enicita Westwood, 1840 freyi Stackelberg, 1963 = Enicomira Duda, 1926 fuscinervis (Zetterstedt, 1838) annulipes (Meigen, 1826) = unicolor Loew, 1874 arctica (Becker, 1915) hyalipennis von Roser, 1840 biloba Andersson, 1975 lapponica Frey, 1924 germanica Duda, 1926 latifibula Frey, 1924 gracilis (Zetterstedt, 1847) montana Day, 1881 leachi (Meigen, 1826) = borealis Frey, 1924 lucida (Staeger, 1844) ornatifrons Frey, 1924 malformans Melander & Spuler, 1917 phyllophora Melander, 1920 minor (Haliday, 1833) = nigricosta auct. nec Rondani, 1868 nigricornis (Meigen, 1826) robusta Loew, 1847 paludosa Elberg, 1963 silvatica Meigen, 1830 pusilla (Zetterstedt, 1847) Trypetoptera Hendel, 1900 putris (Linnaeus, 1758) punctulata (Scopoli, 1763) superba (Haliday, 1833)