FAMILY Gasterosteidae Bonaparte, 1831 – sticklebacks GENUS Apeltes DeKay, 1842 - sticklebacks Species Apeltes quadracus (Mitchill, 1815) - fourspine stickleback [=apeltes, millepunctatus] GENUS Culaea Whitley, 1950 - sticklebacks [=Eucalia] Species Culaea inconstans (Kirtland, 1840) - brook stickleback [=cayuga, gymnetes, micropus, pygmaeus] GENUS Gasterosteus Linnaeus, 1758 - sticklebacks [=Gasteracanthus, Gladiunculus, Leiurus, Obolarius] Species Gasterosteus aculeatus Linnaeus, 1758 - threespined stickleback [=algeriensis, argentatissimus, argyropomus, atkinsii, bailloni, biaculeatus, biarmatus, bispinosus, brachycentrus, cataphractus, cuvieri, dekayi, dimidiatus, elegans, gymnurus, hologymnus, inopinatus, insculptus, intermedius, japonicus, leiurus, loricatus, messinicus, microcephalus, nemausensis, neoboracensis, neustrianus, niger, noveboracensis, obolarius, plebeius, ponticus, pugetti, quadrispinosa, santaennae, semiarmatus, semiloricatus, serratus, spinulosus, suppositus, teraculeatus, tetracanthus, texanus, trachurus, williamsoni] Species Gasterosteus crenobiontus Bacescu & Mayer, 1956 - Techirghiol stickleback Species Gasterosteus islandicus Sauvage, 1874 - Iceland stickleback Species Gasterosteus nipponicus Higuchi et al., 2014 - Higuchi's stickleback Species Gasterosteus wheatlandi Putnam, 1867 - blackspotted stickleback [=gladiunculus] GENUS Pungitius Coste, 1848 - sticklebacks [=Gasterostea, Pygosteus] Species Pungitius bussei (Warpachowski, in Warpachowski & Herzenstein, 1888) – Busse's stickleback Species Pungitius hellenicus Stephanidis, 1971 - ellinopygosteos Species Pungitius kaibarae (Tanaka, 1915) - smoothtail ninespined stickleback Species Pungitius laevis (Cuvier, in Cuvier & Valenciennes, 1829) - laevis stickleback [=breviceps, lotharingus] Species Pungitius platygaster (Kessler, 1877) - Southern ninespined stickleback [=aralensis, caucasicus, danubica, kessleri, niger, nuda] Species Pungitius polyakovi Shedko et al., 2005 - Polyakov's stickleback Species Pungitius pungitius (Linnaeus, 1758) - ninespined stickleback [=blanchardi, brachypoda, burgundianus, carinata, concinnus, dekayi, globiceps, hologymna, mainensis, nebulosus, occidentalis, semiarmata, trachura] Species Pungitius sinensis (Guinchenot, 1869) - Amur stickleback [=brevispinosus, japonicus, steindachneri, wosnesenjenskyi] Species Pingitius stenurus (Kessler, 1876) - Kessler's stickleback Species Pingitius tymensis (Nikolskii, 1889) - Sakhalin stickleback [=undecimalis] Species Pingitius vulgaris (Mauduyt, 1849) - Mauduyt's stickleback GENUS Spinachia Cuvier, 1816 - sticklebacks [=Gastraea, Polycanthus, Spinachia F] Species Spinachia spinachia (Linnaeus, 1758) - sea stickleback [=linnei, marinus, vulagaris]