Atlas of the Bryophytes of Wisconsin Jeff Rose Department of Botany University of Wisconsin-Madison 20 September, 2019

Contents

Diversity of Bryophytes by County 2

History of Bryology in Wisconsin 3

Atlas Methodology 3

Mosses 4 Floristics ...... 4 About the Maps & Nomenclature ...... 4

Hornworts and Liverworts 51 Floristics ...... 51 About the Maps & Nomenclature ...... 51

1 Diversity of Bryophytes by County

331 309 267 175 224 61 58 133 65 48 196 90 137 144 50 189 80 84 37 44 9 54 89 8 204 219 47 78 36 94 0 86 192 61 29 31 81 49 76 118 38 11 45 92 134 number of species 34 45 148 36 68 under 11 92 98 11 − 49 160 126 49 − 68 208 263 35 101139 68 − 109 55 109 − 148 224 157 109 161 172 148 − 175 203 175 − 224 65 81 62 127 102 224 − 267 31 over 267

2 History of Bryology in Wisconsin

The earliest collections of Wisconsin bryophytes date back to those of Increase Lapham from the 1830’s. Other important collectors or students of Wisconsin Bryophytes include C. Barnes (WIS), F. Bowers (UWSP), L. Cheney (WIS), G. Conklin (WIS), J. Christy (MPM), R. Evans (WIS), S. Freckmann (UWSP), H. Iltis (WIS), R. Koch (UWL), and M. Nee (WIS). As seen in chart below, the peak activity in bryophyte collecting was between 1960-1980 with over 10,000 collections in that time frame. 5000 3000 Collections 1000 0

1800 1840 1880 1920 1960 2000

year

Atlas Methodology

Records were downloaded from the Consortium of North American Bryophyte Herbaria website and supplemented by the database at the Milwaukee Public Museum (MPM). Currently, 41,456 Wisconsin collections are databased. Names were then examined to check for synonymy, remove unidentified specimens, and remove specimens lacking locality data. When possible, dubious specimens at WIS have been checked. However, specimens from other herbaria have not been checked unless they are present in duplicate at WIS. In cases where the identifications are highly dubious, these collections have not been mapped and thespecies excluded from the flora for the time being.

3 Mosses

Floristics

Mosses (Bryophyta) can be found in all terrestrial and aquatic habitats throughout Wisconsin. Habitats particularly well-suited for large amounts of moss diversity include cliff faces, wetlands (bogs, fens) and particularly white cedar (Thuja occidentalis) swamps. The distribution and diversity of mosses is far from uniform throughout the states (see map). This is due to a combination of non-uniform distribution of habitats and collection bias towards more “scenic” areas. The problem of collection bias is further hampered by a lack of bryologists actively collecting in Wisconsin in the last several decades. Areas of the highest moss diversity in the state are (1) the Driftless Area, particularly the Baraboo Hills, (2) the Wisconsin River Drainage, (3) the Lake Superior shore, and (4) the Door Peninsula. Conversely, areas of low diversity are (1) West-Central Wisconsin and (2) the Lake Winnebago Area. High diversity areas (1), (2), and (4) are also well-known for their high diversity of vascular plant species. The Baraboo Hills contains species that likely persisted in the Driftless Area during Pleistocene glaciation, but also contains some northern species disjunct from the northern part of the state. Driftless area specialties include Bryoxiphium norvegicum, Dicranodontium denudatum and Jaffueliobryum spp. Another floristically important feature of the Driftless Area is several Ozarkian species that enter the state in Grant County including Forsstroemia trichomitria and Zygodon viridissimus. Associated with the diversity of the Driftless area is a noticeable increase in diversity along the Wisconsin River (and apparent slight decrease in the Adams/Juneau County areas of glacial Lake Wisconsin). This seems to be sometimes associated with species that are more widespread in the northern portion of the state but follow the length of the river to the Mississippi River. Examples of this pattern include Homalia trichomanoides, Isopterygiopsis muelleriana, Mnium lycopodioides, and Saelania glaucescens. Door County is significant in its mixture of northern and southern habitats, but also supports several Great Lakes dune species such as Niphotrichum cansecens. Areas of low diversity are misleading in that the exceptionally low numbers are more than likely due to under collection (for example, Pepin County likely has at least one moss species), although habitat loss due to agriculture and/or relative habitat uniformity is likely to be an additional factor. These areas should specifically be targeted for additional records, especially the west-central part of the state, where speciesof a slightly more western distribution may yet be found.

About the Maps & Nomenclature

The moss flora of Wisconsin presently contains 412 species in 45 families. Nomenclature follows Floraof North America volumes 27 and 28. Grey shading in counties indicates one or more specimens documenting that species, although the identity of the specimen may not be verified.

4 Abietinella abietina Acaulon muticum Amblyodon dealbatus (Hedw.) M. Fleisch. (Schreb.) Mull. Hal. (Sw.) Bruch & Schimp. Thuidiaceae Pottiaceae Meesiaceae

Amblystegium serpens Amphidium lapponicum Anacamptodon splachnoides (Hedw.) Schimp. (Hedw.) Schimp. (Froel.) Brid. Amblystegiaceae Orthotrichaceae Fabroniaceae

Andreaea rupestris Anomobryum concinnatum Anomobryum julaceum Hedw. (Schrad.) Schimp. (Schrad.) Schimp. Andreaeaceae Bryaceae Bryaceae

5 Anomodon attenuatus Anomodon minor Anomodon rostratus (Hedw.) Hub. (Hedw.) Lindb. (Hedw.) Schimp. Thuidiaceae Thuidiaceae Thuidiaceae

Anomodon rugelii Anomodon tristis Anomodon viticulosus (Mull. Hal.) Keissl. (Ces.) Sull. & Lesq. (Hedw.) Hook. & Taylor Thuidiaceae Thuidiaceae Thuidiaceae

Aphanorrhegma serratum Archidium alternifolium Atrichum altecristatum (Hook. & Wilson) Sull. (Dicks.) Mitt. (Ren. & Card.) Smyth & Smyth Funariaceae Archidiaceae Polytrichaceae

6 Atrichum angustatum Atrichum crispum Atrichum oerstedianum (Brid.) Bruch & Schimp. (James) Sull. (Mull. Hal.) Mitt. Polytrichaceae Polytrichaceae Polytrichaceae

Atrichum tenellum Atrichum undulatum Aulacomnium androgynum (Rahl.) Bruch & Schimp. (Hedw.) P. Beauv. (Hedw.) Schwagr. Polytrichaceae Polytrichaceae Aulacomniaceae

Aulacomnium heterostichum Aulacomnium palustre Barbula convoluta (Hedw.) Bruch & Schimp. (Hedw.) Schwagr. Hedw. Aulacomniaceae Aulacomniaceae Pottiaceae

7 Barbula unguiculata Bartramia pomiformis Blindia acuta Hedw. Hedw. (Hedw.) Bruch & Schimp. Pottiaceae Bartramiaceae Seligeriaceae

Brachytheciastrum velutinum Brachythecium acuminatum Brachythecium acutum (Hedw.) Ignatov & Huttunen (Hedw.) Aust. (Mitt.) Sull. Brachytheciaceae Brachytheciaceae Brachytheciaceae

Brachythecium campestre Brachythecium erythrorrhizon Brachythecium laetum (Mull. Hal.) Schimp. Schimp. (Brid.) Schimp. Brachytheciaceae Brachytheciaceae Brachytheciaceae

8 Brachythecium rivulare Brachythecium rotaeanum Brachythecium rutabulum Schimp. De Not. (Hedw.) Schimp. Brachytheciaceae Brachytheciaceae Brachytheciaceae

Brachythecium turgidum Brothera leana Brotherella recurvans (Hartm.) Kindb. (Sull.) Mull. Hal. (Michx.) M. Fleisch. Brachytheciaceae Dicranaceae Sematophyllaceae

Bruchia flexuosa Bryhnia graminicolor Bryhnia novae−angliae (Schwagr.) Mull. Hal. (Brid.) Grout (Sull. & Lesq.) Grout Bruchiaceae Brachytheciaceae Brachytheciaceae

9 Bryoerythrophyllum recurvirostre Bryoxiphium norvegicum Bryum argenteum (Hedw.) Chen (Brid.) Mitt. Hedw. Pottiaceae Bryoxiphiaceae Bryaceae

Bucklandiella heterosticha Buxbaumia aphylla Callicladium haldanianum (Hedw.) Bednarek−Ochyra & Ochyra Hedw. (Grev.) H.A. Crum Grimmiaceae Buxbaumiaceae Hypnaceae

Calliergon cordifolium Calliergon giganteum Calliergon richardsonii (Hedw.) Kindb. (Schimp.) Kindb. (Mitt.) Kindb. Amblystegiaceae Amblystegiaceae Amblystegiaceae

10 Calliergonella cuspidata Campyliadelphus chrysophyllus Campylium stellatum (Hedw.) Loeske (Brid.) R.S. Chopra (Hedw.) C.E.O. Jensen Hypnaceae Amblystegiaceae Amblystegiaceae

Campylophyllum hispidulum Ceratodon purpureus Cirriphyllum piliferum (Brid.) Hedenas (Hedw.) Brid. (Hedw.) Grout Hypnaceae Ditrichaceae Brachytheciaceae

Clasmatodon parvulus Climacium americanum Climacium dendroides (Hampe) Sull. Brid. (Hedw.) F. Weber & D. Mohr Fabroniaceae Climaciaceae Climaciaceae

11 Codriophorus acicularis Conardia compacta Coscinodon calyptratus (Hedw.) P. Beauv. (Mull. Hal.) H. Rob. (Drumm.) C.E.O. Jensen Grimmiaceae Amblystegiaceae Grimmiaceae

Cratoneuron filicinum Ctenidium molluscum Cynodontium strumiferum (Hedw.) Spruce (Hedw.) Mitt. (Hedw.) Lindb. Amblystegiaceae Hypnaceae Dicranaceae

Cynodontium tenellum Cyrtohypnum minutulum Cyrtohypnum pygmaeum (Schimp.) Limpr. (Hedw.) W.R. Buck & H.A. Crum (Schimp.) W.R. Buck & H.A. Crum Dicranaceae Thuidiaceae Thuidiaceae

12 Dichelyma capillaceum Dichelyma falcatum Dichelyma pallescens (With.) Myrin (Hedw.) Myrin Bruch & Schimp. Fontinalaceae Fontinalaceae Fontinalaceae

Dichodontium pellucidum Dicranella cerviculata Dicranella heteromalla (Hedw.) Schimp. (Hedw.) Schimp. (Hedw.) Schimp. Dicranaceae Dicranaceae Dicranaceae

Dicranella rufescens Dicranella schreberiana Dicranella staphylina (With.) Schimp. (Hedw.) Hilf. H. Whitehouse Dicranaceae Dicranaceae Dicranaceae

13 Dicranella subulata Dicranella varia Dicranodontium denudatum (Hedw.) Schimp. (Hedw.) Schimp. (Brid.) E. Britton Dicranaceae Dicranaceae Dicranaceae

Dicranoweisia crispula Dicranum bonjeanii Dicranum condensatum (Hedw.) Milde De Not. Hedw. Dicranaceae Dicranaceae Dicranaceae

Dicranum flagellare Dicranum fulvum Dicranum fuscescens Hedw. Hook. Turner Dicranaceae Dicranaceae Dicranaceae

14 Dicranum montanum Dicranum muehlenbeckii Dicranum ontariense Hedw. Bruch & Schimp. W.L. Peterson Dicranaceae Dicranaceae Dicranaceae

Dicranum polysetum Dicranum scoparium Dicranum undulatum Sw. Hedw. Schrad. Dicranaceae Dicranaceae Dicranaceae

Dicranum viride Didymodon fallax Didymodon rigidulus (Sull. & Lesq.) Lindb. (Hedw.) R.H. Zander Hedw. Dicranaceae Pottiaceae Pottiaceae

15 Didymodon tophaceus Didymodon vinealis Diphyscium foliosum (Brid.) Lisa (Brid.) R.H. Zander (Hedw.) D. Mohr Pottiaceae Pottiaceae Buxbaumiaceae

Distichium capillaceum Distichium inclinatum Ditrichum flexicaule (Hedw.) Bruch & Schimp. (Hedw.) Bruch & Schimp. (Schwagr.) Hampe Ditrichaceae Ditrichaceae Ditrichaceae

Ditrichum lineare Ditrichum pallidum Ditrichum pusillum (Sw.) Lindb. (Hedw.) Hampe (Hedw.) Hampe Ditrichaceae Ditrichaceae Ditrichaceae

16 Drepanocladus aduncus Drepanocladus longifolius Drepanocladus polygamus (Hedw.) Warnst. (Mitt.) Broth. (Schimp.) Hedenas Amblystegiaceae Amblystegiaceae Amblystegiaceae

Drepanocladus sordidus Drummondia prorepens Elodium blandowii (Mull. Hal.) Hedenas (Hedw.) E. Britton (F. Weber & D. Mohr) Eckel Amblystegiaceae Orthotrichaceae Thuidiaceae

Elodium paludosum Encalypta ciliata Encalypta procera (Aust.) Broth. Hedw. Bruch Thuidiaceae Encalyptaceae Encalyptaceae

17 Encalypta rhaptocarpa Entodon brevisetus Entodon challengeri Schwagr. (Hook. & Wilson) Lindb. (Paris) Cardot Encalyptaceae Entodontaceae Entodontaceae

Entodon cladorrhizans Entodon seductrix Ephemerum cohaerens (Hedw.) Mull. Hal. (Hedw.) Mull. Hal. (Hedw.) Hampe Entodontaceae Entodontaceae Ephemeraceae

Ephemerum crassinervium Ephemerum spinulosum Eucladium verticillatum (Schwagr.) Hampe Bruch & Schimp. (Hedw.) Bruch & Schimp. Ephemeraceae Ephemeraceae Pottiaceae

18 Eurhynchiastrum pulchellum Eurhynchium praelongum Fabronia ciliaris (Hedw.) Ignatov & Huttunen (Hedw.) Schimp. (Brid.) Brid. Brachytheciaceae Brachytheciaceae Fabroniaceae

Fissidens adianthoides Fissidens bryoides Fissidens bushii Hedw. Hedw. (Cardot & Ther.) Cardot & Ther. Fissidentaceae Fissidentaceae Fissidentaceae

Fissidens dubius Fissidens elegans Fissidens exilis P. Beauv. Brid. Hedw. Fissidentaceae Fissidentaceae Fissidentaceae

19 Fissidens fontanus Fissidens grandifrons Fissidens minutulus (Bach. Pyl.) Steud. Brid. Sull. Fissidentaceae Fissidentaceae Fissidentaceae

Fissidens obtusifolius Fissidens osmundioides Fissidens subbasilaris Wilson Hedw. Hedw. Fissidentaceae Fissidentaceae Fissidentaceae

Fissidens taxifolius Fontinalis antipyretica Fontinalis dalecarlica Hedw. Hedw. Bruch & Schimp. Fissidentaceae Fontinalaceae Fontinalaceae

20 Fontinalis hypnoides Fontinalis novae−angliae Fontinalis sphagnifolia Hartm. Sull. (Mull. Hal.) Wijk & Margad. Fontinalaceae Fontinalaceae Fontinalaceae

Fontinalis sullivantii Forsstroemia trichomitria Funaria americana Lindb. (Hedw.) Lindb. Lindb. Fontinalaceae Leucodontaceae Funariaceae

Funaria flavicans Funaria hygrometrica Gemmabryum caespiticum Michx. Hedw. (Hedw.) J.R. Spence Funariaceae Funariaceae Bryaceae

21 Gemmabryum dichotomum Grimmia laevigata Grimmia muehlenbeckii (Hedw.) J.R. Spence & H.P. Ramsay (Brid.) Brid. Schimp. Bryaceae Grimmiaceae Grimmiaceae

Grimmia olneyi Grimmia ovalis Grimmia pilifera Sull. (Hedw.) Lindb. P. Beauv. Grimmiaceae Grimmiaceae Grimmiaceae

Grimmia plagiopodia Grimmia teretinervis Gymnostomum aeruginosum Hedw. Limpr. Sm. Grimmiaceae Grimmiaceae Pottiaceae

22 Hamaticaulis vernicosus Haplocladium microphyllum Haplocladium virginianum (Mitt.) Hedenas (Hedw.) Broth. (Brid.) Broth. Amblystegiaceae Thuidiaceae Thuidiaceae

Hedwigia ciliata Herzogiella striatella Herzogiella turfacea (Hedw.) P. Beauv. (Brid.) Z. Iwats. (Lindb.) Z. Iwats. Hedwigiaceae Hypnaceae Hypnaceae

Homalia trichomanoides Homalotheciella subcapillata Homomallium adnatum (Hedw.) Schimp. (Hedw.) Broth. (Hedw.) Broth. Neckeraceae Brachytheciaceae Hypnaceae

23 Hygroamblystegium varium ssp. noterophilum Hygroamblystegium varium ssp. varium Hygroamblystegium varium var. humile (Sull. & Lesq.) Vanderp. & Hedenas (Hedw.) Monk. (P. Beauv.) Vanderp. & Hedenas Amblystegiaceae Amblystegiaceae Amblystegiaceae

Hygrohypnum eugyrium Hygrohypnum luridum Hygrohypnum ochraceum (Schimp.) Loeske (Hedw.) Jenn. (Turner) Loeske Amblystegiaceae Amblystegiaceae Amblystegiaceae

Hylocomiastrum pyrenaicum Hylocomium splendens Hymenostylium recurvirostre (Spruce) M. Fleisch. (Hedw.) Schimp. (Hedw.) Dix. Hylocomiaceae Hylocomiaceae Pottiaceae

24 Hyophila involuta Hypnum cupressiforme Hypnum curvifolium (Hook.) A. Jaeger Hedw. Hedw. Pottiaceae Hypnaceae Hypnaceae

Hypnum fauriei Hypnum imponens Hypnum lindbergii Cardot Hedw. (Mitt.) Hedenas Hypnaceae Hypnaceae Hypnaceae

Hypnum pallescens Hypnum pratense Imbribryum miniatum (Hedw.) P. Beauv. Koch (Lesq.) J.R. Spence Hypnaceae Hypnaceae Bryaceae

25 Isopterygiopsis muelleriana Isopterygiopsis pulchella Jaffueliobryum raui (Schimp.) Z. Iwats. (Hedw.) Z. Iwats. (Aust.) Ther. Plagiotheciaceae Plagiotheciaceae Grimmiaceae

Jaffueliobryum wrightii Leptobryum pyriforme Leptodictyum riparium (Sull.) Ther. (Hedw.) Wilson (Hedw.) Warnst. Grimmiaceae Bryaceae Amblystegiaceae

Leskea gracilescens Leskea obscura Leskea polycarpa Hedw. Hedw. Ehrh. Leskeaceae Leskeaceae Leskeaceae

26 Leskeella nervosa Leucobryum albidum Leucobryum glaucum (Brid.) Loeske (Brid.) Lindb. (Hedw.) Angstr. Leskeaceae Dicranaceae Dicranaceae

Leucodon brachypus Leucodon julaceus Lindbergia brachyptera Brid. (Hedw.) Sull. (Mitt.) Kindb. Leucodontaceae Leucodontaceae Leskeaceae

Meesia triquetra Meesia uliginosa Micromitrium megalosporum (L.) Angstr. Hedw. Aust. Meesiaceae Meesiaceae Ephemeraceae

27 Mnium lycopodioides Mnium marginatum Mnium spinulosum Schwagr. (Dicks.) P. Beauv. Bruch & Schimp. Mniaceae Mniaceae Mniaceae

Mnium stellare Mnium thomsonii Myurella julacea Hedw. Schimp. (Schwagr.) Schimp. Mniaceae Mniaceae Theliaceae

Myurella sibirica Neckera pennata Niphotrichum canescens (Mull. Hal.) Reimers Hedw. (Hedw.) Bednarek−Ochyra & Ochyra Theliaceae Neckeraceae Grimmiaceae

28 Oncophorus virens Oncophorus wahlenbergii Orthotrichum anomalum (Hedw.) Brid. Brid. Hedw. Dicranaceae Dicranaceae Orthotrichaceae

Orthotrichum elegans Orthotrichum obtusifolium Orthotrichum ohioense Schwagr. Brid. Sull. & Lesq. Orthotrichaceae Orthotrichaceae Orthotrichaceae

Orthotrichum pumilum Orthotrichum pusillum Orthotrichum sordidum Sw. Mitt. Sull. & Lesq. Orthotrichaceae Orthotrichaceae Orthotrichaceae

29 Orthotrichum speciosum Orthotrichum stellatum Orthotrichum strangulatum Nees Brid. P. Beauv. Orthotrichaceae Orthotrichaceae Orthotrichaceae

Oxyrrhynchium hians Palustriella falcata Paraleucobryum longifolium (Hedw.) Loeske (Brid.) Hedenas (Ehrh.) Loeske Brachytheciaceae Amblystegiaceae Dicranaceae

Philonotis fontana Philonotis marchica Physcomitrella patens (Hedw.) Brid. (Hedw.) Brid. (Hedw.) Bruch. & Schimp. Bartramiaceae Bartramiaceae Funariaceae

30 Physcomitrium collenchymatum Physcomitrium hookeri Physcomitrium immersum Gier Hampe Sull. Funariaceae Funariaceae Funariaceae

Physcomitrium pyriforme Plagiomnium ciliare Plagiomnium cuspidatum (Hedw.) Hampe (Mull. Hal.) T.J. Kop. (Hedw.) T.J. Kop. Funariaceae Mniaceae Mniaceae

Plagiomnium drummondii Plagiomnium ellipticum Plagiomnium medium (Bruch & Schimp.) T.J. Kop. (Brid.) T.J. Kop. (Bruch & Schimp.) T.J. Kop. Mniaceae Mniaceae Mniaceae

31 Plagiomnium rostratum Plagiopus oederianus Plagiothecium cavifolium (Schrad.) T.J. Kop. (Sw.) H.A. Crum & L.E. Anderson (Brid.) Z. Iwats. Mniaceae Bartramiaceae Plagiotheciaceae

Plagiothecium denticulatum Plagiothecium laetum Plagiothecium latebricola (Hedw.) Schimp. Schimp. Schimp. Plagiotheciaceae Plagiotheciaceae Plagiotheciaceae

Platydictya confervoides Platydictya jungermannioides Platydictya minutissimum (Brid.) H.A. Crum (Brid.) H.A. Crum (Sull. & Lesq.) H.A. Crum Amblystegiaceae Amblystegiaceae Amblystegiaceae

32 Platydictya subtilis Platygyrium repens Pleuridium subulatum (Hedw.) H.A. Crum (Brid.) Schimp. (Hedw.) Rabenh. Amblystegiaceae Hypnaceae Ditrichaceae

Pleurozium schreberi Pogonatum urnigerum Pohlia annotina (Willd.) Mitt. (Hedw.) P. Beauv. (Hedw.) Lindb. Hylocomiaceae Polytrichaceae Bryaceae

Pohlia bulbifera Pohlia cruda Pohlia elongata (Warnst.) Warnst. (Hedw.) Lindb. Hedw. Bryaceae Bryaceae Bryaceae

33 Pohlia filum Pohlia lescuriana Pohlia melanodon (Schimp.) Mertensson (Sull.) Ochi (Brid.) A.J. Shaw Bryaceae Bryaceae Bryaceae

Pohlia nutans Pohlia obtusifolia Pohlia proligera (Hedw.) Lindb. (Vill.) L.F. Koch (Kindb.) S.O. Lindberg Bryaceae Bryaceae Bryaceae

Pohlia wahlenbergii Polytrichastrum alpinum Polytrichastrum formosum (F. Weber & D. Mohr) A.L. Andrews (Hedw.) G.L. Sm. (Hedw.) G.L. Sm. Bryaceae Polytrichaceae Polytrichaceae

34 Polytrichastrum longisetum Polytrichastrum ohioense Polytrichastrum pallidisetum (Sw.) G.L. Sm. (Ren. & Card.) G.L. Sm. (Funck) G.L. Sm. Polytrichaceae Polytrichaceae Polytrichaceae

Polytrichum commune Polytrichum juniperinum Polytrichum piliferum Hedw. Hedw. Hedw. Polytrichaceae Polytrichaceae Polytrichaceae

Polytrichum strictum Pseudobryum cinclidioides Pseudocalliergon trifarium Brid. (Hub.) T.J. Kop. (F. Weber & D. Mohr) Loeske Polytrichaceae Mniaceae Amblystegiaceae

35 Pseudocalliergon turgescens Pseudocampylium radicale Pseudoleskea patens (T. Jensen) Loeske (P. Beauv.) Vanderp. & Hedenas (Lindb.) Kindb. Amblystegiaceae Amblystegiaceae Leskeaceae

Pseudoleskeella catenulata Pseudotaxiphyllum distichaceum Pseudotaxiphyllum elegans (Brid.) Kindb. (Mitt.) Z. Iwats. (Brid.) Z. Iwats. Leskeaceae Hypnaceae Hypnaceae

Pterigynandrum filiforme Ptilium crista−castrensis Ptychostomum cernuum Hedw. (Hedw.) De Not. (Hedw.) Hornsch. Pterigynandraceae Hypnaceae Bryaceae

36 Ptychostomum creberrimum Ptychostomum cyclophyllum Ptychostomum pallens (Taylor) J.R. Spence & H.P. Ramsay (Schwagr.) J.R. Spence (Sw.) J.R. Spence Bryaceae Bryaceae Bryaceae

Ptychostomum pallescens Ptychostomum pendulum Ptychostomum pseudotriquetrum (Schleich.) J.R. Spence Hornsch. (Hedw.) J.R. Spence & H.P. Ramsay Bryaceae Bryaceae Bryaceae

Ptychostomum weigelii Pylaisia intricata Pylaisia polyantha (Spreng.) J.R. Spence (Hedw.) Schimp. (Hedw.) Schimp. Bryaceae Hypnaceae Hypnaceae

37 Pylaisia selwynii Pylaisiadelpha tenuirostris Rauiella scita Kindb. (Bruch & Schimp.) W.R. Buck (P. Beauv.) Reimers Hypnaceae Hypnaceae Thuidiaceae

Rhabdoweisia crispata Rhizomnium appalachianum Rhizomnium magnifolium (Dicks.) Lindb. T.J. Kop. (Horik.) T.J. Kop. Rhabdoweisiaceae Mniaceae Mniaceae

Rhizomnium pseudopunctatum Rhizomnium punctatum Rhodobryum ontariense (Bruch & Schimp.) T.J. Kop. (Hedw.) T.J. Kop. (Kindb.) Paris Mniaceae Mniaceae Bryaceae

38 Rhynchostegium aquaticum Rhynchostegium serrulatum Rhytidiadelphus subpinnatus A. Jaeger (Hedw.) A. Jaeger (Lindb.) T.J. Kop. Brachytheciaceae Brachytheciaceae Hylocomiaceae

Rhytidiadelphus triquetrus Rhytidium rugosum Rosulabryum capillare (Hedw.) Warnst. (Ehrh.) Kindb. (Hedw.) J.R. Spence Hylocomiaceae Hylocomiaceae Bryaceae

Saelania glaucescens Sanionia uncinata Sarmetypnum exannulatum (Hedw.) Broth. (Hedw.) Loeske (Schimp.) Hedenas Ditrichaceae Amblystegiaceae Amblystegiaceae

39 Schistidium agassizii Schistidium apocarpum Schistidium appalachianum Sull. & Lesq. (Hedw.) Bruch & Schimp. H. H. Blom. Grimmiaceae Grimmiaceae Grimmiaceae

Schistidium atrofuscum Schistidium confertum Schistidium crassithecium (Schimp.) Limpr. (Funck) Bruch & Schimp. H.H. Blom Grimmiaceae Grimmiaceae Grimmiaceae

Schistidium dupretii Schistidium pulchrum Schistidium rivulare (Ther.) W.A. Weber H.H. Blom (Brid.) Podp. Grimmiaceae Grimmiaceae Grimmiaceae

40 Schistidium viride Schistostega pennata Schwetschkeopsis fabronia H.H. Blom & C. Darigo (Hedw.) F. Weber & D. Mohr (Schwagr.) Broth. Grimmiaceae Schistostegaceae Fabroniaceae

Sciuro−hypnum curtum Sciuro−hypnum oedipodium Sciuro−hypnum plumosum (Lindb.) Ignatov (Mitt.) Ignatov & Huttunen (Hedw.) Ignatov & Huttunen Brachytheciaceae Brachytheciaceae Brachytheciaceae

Sciuro−hypnum populeum Sciuro−hypnum reflexum Sciuro−hypnum starkei (Hedw.) Ignatov & Huttunen (Starke) Ignatov & Huttunen (Brid.) Ignatov & Huttunen Brachytheciaceae Brachytheciaceae Brachytheciaceae

41 Scorpidium cossonii Scorpidium revolvens Scorpidium scorpioides (Schimp.) Hedenas (Sw.) Rubers (Hedw.) Limpr. Scorpidiaceae Scorpidiaceae Scorpidiaceae

Seligeria campylopoda Seligeria donniana Seligeria pusilla Kindb. (Sm.) Mull. Hal. (Hedw.) Bruch & Schimp. Seligeriaceae Seligeriaceae Seligeriaceae

Seligeria recurvata Sematophyllum demissum Sphagnum andersonianum (Hedw.) Bruch & Schimp. (Wilson) Mitt. R.E. Andrus Seligeriaceae Sematophyllaceae Sphagnaceae

42 Sphagnum angustifolium Sphagnum bartlettianum Sphagnum capillifolium (Warnst.) C.E.O. Jensen Warnst. (Ehrh.) Hedw. Sphagnaceae Sphagnaceae Sphagnaceae

Sphagnum centrale Sphagnum compactum Sphagnum contortum C.E.O. Jensen Lam. & DC Schultz Sphagnaceae Sphagnaceae Sphagnaceae

Sphagnum cuspidatum Sphagnum fallax Sphagnum fimbriatum Ehrh. H. Klinggr. Wilson Sphagnaceae Sphagnaceae Sphagnaceae

43 Sphagnum flexuosum Sphagnum fuscum Sphagnum girgensohnii Dozy & Molk. (Schimp.) H. Klinggr. Russow Sphagnaceae Sphagnaceae Sphagnaceae

Sphagnum henryense Sphagnum imbricatum Sphagnum inundatum Warnst. Hornsch. Russow Sphagnaceae Sphagnaceae Sphagnaceae

Sphagnum lescurii Sphagnum magellanicum Sphagnum majus Sull. Brid. (Russow) C.E.O. Jensen Sphagnaceae Sphagnaceae Sphagnaceae

44 Sphagnum obtusum Sphagnum palustre Sphagnum papillosum Warnst. L. Lindb. Sphagnaceae Sphagnaceae Sphagnaceae

Sphagnum platyphyllum Sphagnum pulchrum Sphagnum recurvum (Lindb.) Warnst. (Lindb.) Warnst. P. Beauv. Sphagnaceae Sphagnaceae Sphagnaceae

Sphagnum riparium Sphagnum rubellum Sphagnum russowii Angstr. Wilson Warnst. Sphagnaceae Sphagnaceae Sphagnaceae

45 Sphagnum squarrosum Sphagnum subfulvum Sphagnum subsecundum Crome Sjors Nees Sphagnaceae Sphagnaceae Sphagnaceae

Sphagnum subtile Sphagnum tenerum Sphagnum teres (Russow) Warnst. Sull. & Lesq. (Schimp.) Angstr. Sphagnaceae Sphagnaceae Sphagnaceae

Sphagnum warnstorfii Sphagnum wulfianum Splachnum ampullaceum Russow Girg. Hedw. Sphagnaceae Sphagnaceae Splachnaceae

46 Splachnum sphaericum Straminergon stramineum Syntrichia papillosa Hedw. (Dicks.) Hedenas (Wilson) Jur. Splachnaceae Amblystegiaceae Pottiaceae

Syntrichia ruralis Taxiphyllum deplanatum Taxiphyllum taxirameum (Hedw.) F. Weber & D. Mohr (Bruch & Schimp.) M. Fleisch. (Mitt.) Fl. Pottiaceae Hypnaceae Hypnaceae

Tetraphis pellucida Thelia asprella Thelia hirtella Hedw. (Schimp.) Sull. & Lesq. (Hedw.) Sull. & Lesq. Tetraphidaceae Theliaceae Theliaceae

47 Thelia lescurii Thuidium delicatulum Thuidium recognitum Sull. (Hedw.) Schimp. (Hedw.) Lindb. Theliaceae Thuidiaceae Thuidiaceae

Timmia megapolitana Tomentypnum nitens Tortella fragilis Hedw. (Hedw.) Loeske (Hook. & Wilson) Limpr. Timmiaceae Brachytheciaceae Pottiaceae

Tortella humilis Tortella inclinata Tortella tortuosa (Hedw.) Jenn. (R. Hedw.) Limpr. (Hedw.) Limpr. Pottiaceae Pottiaceae Pottiaceae

48 Tortula acaulon Tortula cernua Tortula mucronifolia (With.) R.H. Zander (Hub.) Lindb. Schwagr. Pottiaceae Pottiaceae Pottiaceae

Tortula obtusifolia Tortula plinthobia Tortula truncata (Schwagr.) Mathieu (Sull. & Lesq.) Broth. (Hedw.) Mitt. Pottiaceae Pottiaceae Pottiaceae

Trematodon ambiguus Trichostomum spirale Trichostomum tenuirostre (Hedw.) Hornsch. Grout (Hook. & Taylor) Lindb. Bruchiaceae Pottiaceae Pottiaceae

49 Ulota crispa Ulota hutchinsiae Warnstorfia fluitans (Hedw.) Brid. (Sm.) Hammar (Hedw.) Loeske Orthotrichaceae Orthotrichaceae Amblystegiaceae

Warnstorfia pseudostraminea Weissia controversa Weissia muhlenbergiana (Mull. Hal.) Tuom. & T.J. Kop. Hedw. (Sw.) W.D. Reese & B.A.E. Lemmon Amblystegiaceae Pottiaceae Pottiaceae

Zygodon viridissimus (Dicks.) Brid. Orthotrichaceae

50 Hornworts and Liverworts

Floristics

Coming soon!

About the Maps & Nomenclature

Nomenclature follows Schuster (1966-1992, Hepaticae and Anthocerotae of North America East of the Hun- drenth Meridian vols. I-VI), sypplemented by Soderstrom et al. (2016, Phytokeys 59:1-828) The hornwort and liverwort flora presently contains 146 species in 32 families. Grey shading in counties indicates one or more specimens documenting that species, although the identity of the specimen may not be verified.

51 Anastrophyllum hellerianum Anastrophyllum michauxii Anastrophyllum minutum (Nees) R.M. Schust. (F. Weber) H. Buch (Schreb.) R.M. Schust. Anastrophyllaceae Anastrophyllaceae Anastrophyllaceae

Aneura pinguis Anthoceros macounii Anthoceros punctatus (L.) Dum. M. Howe L. Aneuraceae Anthocerotaceae Anthocerotaceae

Asterella tenella Barbilophozia attenuata Barbilophozia barbata (L.) P. Beauv. (Nees) Loeske (Schreb.) Loeske Aytoniaceae Anastrophyllaceae Anastrophyllaceae

52 Barbilophozia kunzeana Bazzania trilobata Blasia pusilla (Hub.) K. Muller (L.) Gray L. Anastrophyllaceae Lepidoziaceae Blasiaceae

Blepharostoma trichophyllum Calypogeia fissa Calypogeia integristipula (L.) Dum. (L.) Raddi Steph. Pseudolepicoleaceae Calypogeiaceae Calypogeiaceae

Calypogeia muelleriana Calypogeia neesiana Calypogeia sphagnicola (Schiffn.) K. Muller (C. Massal. & Carestia) K. Muller (Arnell & J. Perss.) Warnst. & Loeske Calypogeiaceae Calypogeiaceae Calypogeiaceae

53 Calypogeia suecica Cephalozia bicuspidata Cephalozia catenulata (Arnell & J. Perss.) K. Muller (L.) Dum. (Hub.) Lindb. Calypogeiaceae Cephaloziaceae Cephaloziaceae

Cephalozia connivens Cephalozia lacinulata Cephalozia lunulifolia (Dicks.) Lindb. (J.B. Jack) Spruce (Dum.) Dum. Cephaloziaceae Cephaloziaceae Cephaloziaceae

Cephalozia macounii Cephalozia neesiana Cephalozia pleniceps (Aust.) Spruce Steph. (Aust.) Lindb. Cephaloziaceae Cephaloziaceae Cephaloziaceae

54 Cephaloziella byssacea Cephaloziella elachista Cephaloziella hampeana (Roth) Warnst. (J.B. Jack) Schiffn. (Nees) Schiffn. Cephaloziellaceae Cephaloziellaceae Cephaloziellaceae

Cephaloziella rubella Chiloscyphus coadunatus Chiloscyphus pallescens (Nees) Warnst. (Sw.) R.M. Schust. & J.J. Engel (Ehrh.) Dum. Cephaloziellaceae Lophocoleaceae Lophocoleaceae

Chiloscyphus polyanthos Cladopodiella fluitans Cololejeunea biddlecomiae (L.) Corda (Nees) Jarg. (Aust.) A. Evans Lophocoleaceae Cephaloziaceae Lejeuneaceae

55 Conocephalum salebrosum Dichiton integerrimum Diplophyllum apiculatum Szweyk., Buczkowska & Odrzykoski (Lindb.) H. Buch (A. Evans) Steph. Conocephalaceae Cephaloziellaceae Scapaniaceae

Fossombronia foveolata Frullania asagrayana Frullania bolanderi Lindb. Mont. Aust. Fossombroniaceae Jubulaceae Jubulaceae

Frullania brittoniae Frullania eboracensis Frullania inflata A. Evans Gott. Gott. Jubulaceae Jubulaceae Jubulaceae

56 Frullania oakesiana Frullania plana Frullania riparia Aust. Sull. Hampe Jubulaceae Jubulaceae Jubulaceae

Frullania selwyniana Geocalyx graveolens Gymnocolea inflata Pearson (Schrad.) Nees (Huds.) Dum. Jubulaceae Geocalycaceae Jungermanniaceae

Harpanthus drummondii Harpanthus scutatus Jamesoniella autumnalis (Taylor) Grolle (F. Weber & D. Mohr) Spruce (DC) Steph. Geocalycaceae Geocalycaceae Jungermanniaceae

57 Jungermannia lanceolata Jungermannia schraderi Lejeunea cavifolia L. Mart. (Ehrh.) Lindb. Jungermanniaceae Jungermanniaceae Lejeuneaceae

Lepidozia reptans Lophocolea bidentata Lophocolea heterophylla (L.) Dum. (L.) Dum. (Schrad.) Dum. Lepidoziaceae Lophocoleaceae Lophocoleaceae

Lophocolea minor Lophozia alpestris Lophozia badensis Nees (Schleich.) Evans (Gott.) Schiffn. Lophocoleaceae Jungermanniaceae Jungermanniaceae

58 Lophozia bicrenata Lophozia collaris Lophozia excisa (Schmidel) Dum. (Mart.) Dum. (Dicks.) Dum. Jungermanniaceae Jungermanniaceae Jungermanniaceae

Lophozia gillmanii Lophozia heterocolpos Lophozia incisa (Aust.) R.M. Schust. (Thed.) M. Howe (Schrad.) Dum. Jungermanniaceae Jungermanniaceae Jungermanniaceae

Lophozia longidens Lophozia marchica Lophozia muelleri (Lindb.) Macoun (Nees.) Steph. (Nees.) Dym Jungermanniaceae Jungermanniaceae Jungermanniaceae

59 Lophozia porphyroleuca Lophozia quadriloba Lophozia serpens (Nees) Schiffn. (Lindb.) A. Evans (Dum.) Dum. Jungermanniaceae Jungermanniaceae Jungermanniaceae

Lophozia ventricosa Lunularia cruciata Mannia fragrans (Dicks.) Dum. (L.) Dum. (Balb.) Frye & L. Clark Jungermanniaceae Lunulariaceae Aytoniaceae

Mannia pilosa Mannia triandra Marchantia polymorpha (Hornem.) Frye & L. Clark (Scop.) Grolle L. Aytoniaceae Aytoniaceae Marchantiaceae

60 Marsupella emarginata Mesoptychia rutheana Metzgeria conjugata (Ehrh.) Dum. (Limpr.) L. S.derstr. & V._a Lindb. Gymnomitriaceae Mesoptychioideae Metzgeriaceae

Moerckia hibernica Mylia anomala Nardia geoscyphus (Hook.) Gott. (Hook.) A. Gray (De Not.) Lindb. Pallaviciniaceae Jubulaceae Jungermanniaceae

Notothylas orbicularis Nowellia curvifolia Odontoschisma denudatum (Schwein.) Sull. (Dicks.) Mitt. (Nees) Dum. Notothyladaceae Cephaloziaceae Cephaloziaceae

61 Odontoschisma macounii Pallavicinia lyellii Pellia epiphylla (Aust.) Uderw. (Hook.) Gray (L.) Corda Cephaloziaceae Pallaviciniaceae Pelliaceae

Pellia megaspora Pellia neesiana Phaeoceros carolinianus R.M. Schust. (Gott.) Limpr. (Michx.) Prosk. Pelliaceae Pelliaceae Anthocerotaceae

Phaeoceros laevis Plagiochila porelloides Porella pinnata (L.) Prosk. (Torr.) Lindenb. L. Anthocerotaceae Plagiochilaceae Porellaceae

62 Porella platyphylla Preissia quadrata Ptilidium ciliare (L.) Pfeiff. (Scop.) Nees (L.) Hampe Porellaceae Marchantiaceae Ptilidiaceae

Ptilidium pulcherrimum Radula complanata Radula obconica (Weber) Hampe (L.) Dum. Sull. Ptilidiaceae Radulaceae Radulaceae

Reboulia hemisphaerica Riccardia latifrons Riccardia multifida (L.) Raddi (Lindb.) Lindb. (L.) Gray Aytoniaceae Aneuraceae Aneuraceae

63 Riccardia palmata Riccia beyrichiana Riccia bifurca (Hedw.) Carruth. Hampe Hoffm. Aneuraceae Ricciaceae Ricciaceae

Riccia cavernosa Riccia fluitans Riccia frostii Hoffm. L. Aust. Ricciaceae Ricciaceae Ricciaceae

Riccia membranacea Riccia sorocarpa Riccia sullivantii Gott., Lindenb. & Nees Bisch. Aust. Ricciaceae Ricciaceae Ricciaceae

64 Ricciocarpos natans Scapania apiculata Scapania brevicaulis (L.) Corda Spruce Taylor Ricciaceae Scapaniaceae Scapaniaceae

Scapania curta Scapania cuspiduligera Scapania glaucocephala (Mart.) Dum. (Nees) K. Muller (Taylor) Aust. Scapaniaceae Scapaniaceae Scapaniaceae

Scapania gymnostomophila Scapania irrigua Scapania kaurinii Kaal. (Nees) Nees Ryan Scapaniaceae Scapaniaceae Scapaniaceae

65 Scapania mucronata Scapania nemorea Scapania paludicola H. Buch (L.) Grolle Loeske & K. Muller Scapaniaceae Scapaniaceae Scapaniaceae

Scapania paludosa Scapania subalpina Scapania umbrosa (Mull. Frib.) Mull. Frib. (Nees) Dum. (Schrad.) Dum. Scapaniaceae Scapaniaceae Scapaniaceae

Scapania undulata Solenostoma atrovirens Solenostoma cordifolium (L.) Dum. (Dum.) Mull. Frib. (Dum.) Steph. Scapaniaceae Jungermanniaceae Jungermanniaceae

66 Solenostoma crenuliformis Solenostoma fossombronioides Solenostoma gracillimum Steph. (Aust.) R.M. Schust. R.M. Schust. Jungermanniaceae Jungermanniaceae Jungermanniaceae

Solenostoma hyalina Solenostoma pumila Solenostoma sphaerocarpum (Lyell) Mitt. (With.) K. Muller (Hook.) Steph. Jungermanniaceae Jungermanniaceae Jungermanniaceae

Trichocolea tomentella Tritomaria exsecta Tritomaria exsectiformis (Ehrh.) Dum. (Schmidel) Schiffn. (Breidl.) Schiffn. Trichocoleaceae Jungermanniaceae Jungermanniaceae

67 Tritomaria quinquedentata Tritomaria scitula (Huds.) H. Buch (Taylor) Jorg. Jungermanniaceae Jungermanniaceae

68