NORTH-WESTERN JOURNAL OF ZOOLOGY 8 (1): 1-21 ©NwjZ, Oradea, Romania, 2012 Article No.: 111136 http://biozoojournals.3x.ro/nwjz/index.html

Pseudoscorpions of the Czech Republic and Slovakia: An annotated and revised checklist (Arachnida: Pseudoscorpiones)

Jana CHRISTOPHORYOVÁ1*, František ŠŤÁHLAVSKÝ2, Miroslav KRUMPÁL1 and Peter FEDOR3

1. Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynská dolina B–1, SK–842 15 Bratislava, Slovakia. 2. Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ–128 44 Prague 2, Czech Republic. 3. Department of Ecosozology (Conservation Biology), Faculty of Natural Sciences, Comenius University, Mlynská dolina B–1, SK–842 15 Bratislava, Slovakia. * Corresponding author: J. Christophoryová, E-mail: [email protected]

Received: 03. April 2011 / Accepted: 27. August 2011 / Available online: 18. September 2011 / Printed: June 2012

Abstract. An annotated and revised checklist is presented comprising 58 species (8 families) of the arachnid order Pseudoscorpiones from the Czech Republic (38 species) and Slovakia (51 species), formerly Czechoslovakia. According to the latest detailed analyses and revisions Chthonius carinthiacus Beier, 1951, Neobisium validum (L. Koch, 1873), Roncus alpinus L. Koch, 1873 and Pselaphochernes dubius (O. P.-Cambridge, 1892) are excluded from the checklist from Slovakia. Formerly recorded Chthonius diophthalmus Daday, 1888, C. submontanus Beier, 1963, C. kewi Gabbutt, 1966, Neobisium spelaeum (Schiödte, 1847), N. jugorum (L. Koch, 1873), Roncus alpinus and R. transsilvanicus Beier, 1928 have not been included in the list from the Czech Republic. Some generally rare species are discussed.

Keywords: Pseudoscorpiones, checklist, Central Europe, Czechoslovakia, species diversity.

Introduction onymized with Lamprochernes nodosus, see Beier 1932b), “endemic” species in the Czech Republic. Pseudoscorpions are the fourth largest arachnid At the end of the 19th century there were only 14 order (Harvey 2007) with more than 3380 species pseudoscorpion species (belonging to five fami- described in 25 families (Harvey 2009). Europe is a lies) known from this region, but soon the first re- major centre of diversity, and in a review of the vising and summarizing checklist was published major geographical regions of the world (Africa, by Nosek (1901) and later updated (Nosek 1902). Asia, Europe, North America, Australia and Oce- In Slovakia the first records on pseudoscorpions ania, South America), Europe was found to have refer to several Hungarian authors of the late 19th the highest richness with 760 species (Harvey century (e.g. Tömösváry 1882, Daday 1888a, 2007). But even here our knowledge on the distri- 1888b, 1918). Nevertheless, most of our under- bution of many species appears rather sporadic standing of the pseudoscorpion fauna of Slovakia and questionable, especially when older insuffi- dates from the 1950s with the major contributions cient data have been taken into account. More- made by Verner (e.g. Verner 1958a, 1958b, 1960) over, in the Czech Republic and Slovakia, located and later by Krumpál (Krumpál 1979, 1980, in Central Europe, the knowledge on diversity and Krumpál & Kiefer 1981, Krumpál & Cyprich 1987, consequent progress have been limited since the 1988). division of the former Czechoslovakia in 1993. The pseudoscorpions of the Czech Republic The first data on pseudoscorpions from the and Slovakia have been studied more intensively Czech Republic date back to the end of the 18th in the last decades of the 20th century, following century (Preyssler 1790), with the description of the publication of the first Czechoslovak determi- Chthonius tetrachelatus. Almost hundred years nation key (Verner 1971), with a lot of new and later, Stecker (1874) introduced more significant unpublished information, but without distinguish- records on 19 species, but some of them were later ing national diversity. Before the dissolution of synonymized, including the newly described Che- former Czechoslovakia in 1993 many rather gen- lifer serratus and Chernes bohemicus (see Harvey eral faunistic records were published, usually with 2009). Some of the faunistic data published before no detailed information on the exact locality. The the World War I (e.g. Stecker 1875, Simon 1879), world pseudoscorpion catalogue from that period which did not distinguish regions in the Austrian– (Harvey 1991) contains several data on national Hungarian Empire, have thus become confusing, diversity for Czechoslovakia, but difficult to sepa- with the exception of Chernes bohemicus (later syn- rate for the two countries. Just three species (Neo- 2 Christophoryová, J. et al. bisium slovacum, N. beieri and Rhacochelifer quadri- sions on this matter. maculatus) can be specified for Slovakia and five for the Czech Republic (Chthonius tetrachelatus, Materials and Methods Chelifer cancroides and Lamprochernes nodosus, Chthonius diophthalmus and Neobisium sylvaticum The current checklist has been compiled primarily using inaculeatum) (Harvey 1991). all available published records (determination keys, cata- Since the split in 1993, the pseudoscorpion logues, conference proceedings), supplemented by faunis- fauna of the Czech Republic has been studied very tic information from many theses, deposited in university intensively, resulting in plenty of first species re- libraries. Analysing the century old works by Hungarian authors (Tömösváry 1882, Daday 1888a, 1888b, 1918) has cords (Ducháč 1993a, 1995a, 1998a, Schmarda generated many questions, with their specific linguistic 1995, Šťáhlavský & Ducháč 2001, Ducháč & Mle- and geopolitical problems. The present paper operates jnek 2004, Ducháč et al. 2007, Laška et al. 2008, only with exact and clear data. For example, for the town Christophoryová et al. 2011b) and the publication of Slovenské Nové Mesto (Slovakia), established in 1918 of faunistic data from numerous sites (Ducháč after the split of Hungarian Sátoraljaújhely, the previ- 1993b, 1997a, Šťáhlavský 2001, 2006a, 2006b, Šťáh- ously published records were revised to avoid confusion. lavský & Krásný 2007, Šťáhlavský & Tuf 2009). On the other hand, some records omitted by Harvey (1991, 2009) and positively connected with the area of Apart from plenty of faunistic and ecological pa- Slovakia have been taken into account. The data for the pers on pseudoscorpions in Slovakia in the 1990s Czech Republic (CR) are followed by information for Slo- (Krištofík et al. 1993, 1995, 1996, Krumpálová & vakia (SR). All taxa include their relevant references Krumpál 1993, Ducháč 1996a, Fenďa et al. 1998), chronologically listed, accompanied by authors, year and there has been an intensive research recently, ana- faunistic information (relevant pages in literature). The lysing the diversity of pseudoscorpions in caves nomenclature for taxa derived from Harvey (2009). In (Krumpál 2000, Mock et al. 2003, 2004, 2005, Chris- specific cases the faunistic data set may express identifica- tion problems (misidentified or partly misidentified spe- tophoryová & Krumpál 2006c, Kováč et al. 2007, cies). Lapsus calami in names of taxa in certain papers are Papáč et al. 2007a, 2007b), bird nests (Cyprich et al. expressed by [sic]. 2000, Krumpál et al. 2000–2001, Krištofík et al. 2002, 2005, 2007, 2009, Christophoryová et al. 2007a, Christophoryová & Krištofík 2008, Christo- Results phoryová & Krumpál 2009, Christophoryová & Krumpálová 2010) as well as in numerous sites This checklist includes 58 species in eight families throughout the whole country (Krumpál & Krum- of the order Pseudoscorpiones from the Czech Re- pálová 2003, Christophoryová & Krumpál 2005, public (38 species) and Slovakia (51 species), for- 2007b, Christophoryová et al. 2007b, Krumpál & merly Czechoslovakia (Table 1). Chthonius Christophoryová 2007, Christophoryová 2009a, diophthalmus, C. submontanus, C. kewi, Neobisium 2009b, Christophoryová & Mock 2009, Papáč et al. spelaeum, N. jugorum, Roncus alpinus, and R. trans- 2009, Christophoryová & Krumpál 2010). The silvanicus have been recently excluded from the identification key of pseudoscorpions from the fauna of the Czech Republic, and Chthonius carin- Czech Republic and Slovakia was published re- thiacus, Neobisium validum, Roncus alpinus cently (Christophoryová et al. 2011b). and Pselaphochernes dubius from the fauna of Slo- For the Czech Republic, the checklist with 28 vakia. species of seven families was published by Ducháč (1999b), revised soon after by Blick et al. (2004), Order PSEUDOSCORPIONES de Geer, 1778 presenting 33 species, and by Harvey (2009), with Suborder EPIOCHEIRATA Harvey, 1992 5 more species. In Slovakia there is only one Superfamily CHTHONIOIDEA Daday, 1888 checklist available, completed by Harvey (2009) Family CHTHONIIDAE Daday, 1888 and offering information on 37 species (six fami- Subfamily Chthoniinae Daday, 1888 lies). Tribe Chthoniini Daday, 1888 The aim of our study is to revise species diver- sity of pseudoscorpions in Slovakia and the Czech Genus Chthonius C. L. Koch, 1843 Republic, formerly Czechoslovakia, taking the lat- Subgenus Chthonius (Chthonius) C. L. Koch, 1932 est knowledge as well as national fauna specificity into account. The checklist is intended to provide a Chthonius (Chthonius) carinthiacus Beier, 1951 renewed foundation for future studies and discus- CR: Chthonius pygmaeus carinthiacus Beier: Růžička

Checklist of pseudoscorpions from the Czech Republic and Slovakia 3

Table 1. List of pseudoscorpion species of the Czech Republic and Slovakia. Abbreviations: CZ–Czech Republic, SK–Slovakia.

Taxa Country CZ SK Chthoniidae Chthonius (Chthonius) carinthiacus Beier, 1951 + Chthonius (C.) diophthalmus Daday, 1888 + Chthonius (C.) heterodactylus Tömösváry, 1882 + + Chthonius (C.) hungaricus Mahnert, 1980 + Chthonius (C.) ischnocheles (Hermann, 1804) + + Chthonius (C.) orthodactylus (Leach, 1817) + + Chthonius (C.) pygmaeus Beier, 1934 + Chthonius (C.) ressli Beier, 1956 + + Chthonius (C.) subterraneus Beier, 1931 + Chthonius (C.) tenuis L. Koch, 1873 + + Chthonius (Ephippiochthonius) boldorii Beier, 1934 + Chthonius (E.) fuscimanus Simon, 1900 + + Chthonius (E.) tetrachelatus (Preyssler, 1790) + + Mundochthonius carpaticus Rafalski, 1948 + + Mundochthonius styriacus Beier, 1971 + Neobisiidae Microbisium brevifemoratum (Ellingsen, 1903) + Microbisium suecicum Lohmander, 1945 + + Neobisium (Blothrus) slovacum Gulička, 1977 + Neobisium (Neobisium) beieri Verner, 1958 + Neobisium (N.) brevidigitatum (Beier, 1928) + Neobisium (N.) carcinoides (Hermann, 1804) + + Neobisium (N.) carpaticum Beier, 1935 + Neobisium (N.) crassifemoratum (Beier, 1928) + + Neobisium (N.) erythrodactylum (L. Koch, 1873) + + Neobisium (N.) fuscimanum (C. L. Koch, 1843) + + Neobisium (N.) inaequale Chamberlin, 1930 + Neobisium (N.) jugorum (L. Koch, 1873) + Neobisium (N.) macrodactylum (Daday, 1888) + Neobisium (N.) polonicum Rafalski, 1936 + Neobisium (N.) simile (L. Koch, 1873) + Neobisium (N.) sylvaticum (C. L. Koch, 1835) + + Roncus lubricus L. Koch, 1873 + + Roncus transsilvanicus Beier, 1928 + Syarinidae Syarinus strandi (Ellingsen, 1901) + Larcidae Larca lata (Hansen, 1884) + + Cheiridiidae Apocheiridium (Apocheiridium) ferum (Simon, 1879) + Cheiridium museorum (Leach, 1817) + + Atemnidae Atemnus politus (Simon, 1878) + Diplotemnus insolitus Chamberlin, 1933 + Cheliferidae Chelifer cancroides (Linnaeus, 1758) + + Dactylochelifer latreillii (Leach, 1817) + + Mesochelifer ressli Mahnert, 1981 + + Rhacochelifer peculiaris (L. Koch, 1873) + Rhacochelifer quadrimaculatus (Tömösváry, 1882) + Chernetidae Chernes cimicoides (Fabricius, 1793) + + Chernes hahnii (C. L. Koch, 1839) + + 4 Christophoryová, J. et al.

Table 1. (continued)

Taxa Country CZ SK Chernes nigrimanus Ellingsen, 1897 + Chernes similis (Beier, 1932) + + Chernes vicinus (Beier, 1932) + + Dendrochernes cyrneus (L. Koch, 1873) + + Dinocheirus panzeri (C. L. Koch, 1837) + + Allochernes peregrinus Lohmander, 1939 + + Allochernes wideri (C. L. Koch, 1843) + + Anthrenochernes stellae Lohmander, 1939 + Lamprochernes chyzeri (Tömösváry, 1882) + + Lamprochernes nodosus (Schrank, 1803) + + Lasiochernes pilosus (Ellingsen, 1910) + Pselaphochernes scorpioides (Hermann, 1804) + + Total number of species 38 51 et al., 1995: 47; Ducháč, 2002c: 68. Chthonius (Chthonius) heterodactylus Tömös- Chthonius (Chthonius) pygmaeus carinthiacus váry: Ducháč et al., 2007: 31–33; Harvey, 2009: –; Beier: Šťáhlavský, 1996: 4. Christophoryová et al., 2011b: 36. Chthonius (Chthonius) carinthiacus Beier: Har- SR: Chthonius heterodactylus Tömösváry: vey, 2009: –; Šťáhlavský & Tuf, 2009: 98; Christo- Tömösváry, 1882: 241–242; Krumpál & Krum- phoryová et al., 2011b: 36. pálová, 2003: 118–119; Ducháč & Mlejnek, 2004: 37; Chthonius cf. pygmaeus carinthiacus Beier: Christophoryová, 2006b: 7, 8; Kárpáthegyi, 2006: Ducháč, 1997b: –; Ducháč, 2002c: 124, 139–141; 116; Mikula, 2006: 38; Ducháč et al., 2007: 32; Christophoryová, 2006b: 9. Christophoryová & Mock, 2009: 17. Chthonius cf. submontanus Beier: Ducháč, Chthonius (Chthonius) heterodactylus Tömös- 2002c: 102 (misidentification). váry: Szent-Ivány, 1941: 86; Rafalski, 1967: 6; Chthonius (Chthonius) tenuis L. Koch, 1873: Krumpál & Kiefer, 1981: 128; Ducháč, 1989: 180; Šťáhlavský, 2000: 17; Šťáhlavský, 2001: 80; Šťáh- Ducháč, 2002c: 31; Harvey, 2009: –; Christophory- lavský & Král, 2004: 50; Šťáhlavský, 2006a: 169; ová et al., 2011b: 36.

Šťáhlavský, 2006c: 37, 79 (all misidentification). Chthonius (Chthonius) hungaricus Mahnert, 1980 SR (note 1): Chthonius (Chthonius) pygmaeus SR: Chthonius (Chthonius) hungaricus Mahnert: carinthiacus Beier: Šťáhlavský, 1996: 4. Christophoryová, 2009a: 256; Papáč et al., 2009: 37;

Chthonius (Chthonius) diophthalmus Daday, 1888 Christophoryová et al., 2011a: 2–3; Christophory- CR (note 2): Chthonius (Chthonius) diophthalmus ová et al., 2011b: 36. Daday: Harvey, 2009: –. Chthonius hungaricus Mahnert: Christophory- SR: Chthonius diophthalmus Daday: Christo- ová, 2009b: 128; Christophoryová & Krumpál, phoryová, 2006b: 8; Christophoryová, 2008a: 132, 2009: 14.

Christophoryová & Mock, 2009: 17; Mock et al., Chthonius (Chthonius) ischnocheles (Hermann, 2009: 268; Papáč et al., 2009: 37. 1804) Chthonius (Chthonius) diophthalmus Daday: CR: Chthonius Rayi L. Koch, 1873: Stecker, 1874: Krumpál & Kiefer, 1981: 127–128; Christophory- 238; Simon, 1879: 74–75; Nosek, 1901: 21; Nosek, ová, 2009a: 256; Christophoryová et al., 2011b: 36. 1902: 60; Caporiacco, 1926: 75; Ducháč, 2002c: 11– Chthonius ksenemanni Hadži, 1939: Krumpál & 12, 13 (synonymized by Chamberlin, 1929: 72). Krumpálová, 2003: 119; Christophoryová, 2006b: 8 Chthonius ischnocheles (Hermann): Ducháč, (synonymized by Schawaller, 1989: 6). 1999b: 14; Ducháč, 2002c: 20, 68, 82, 136, 138; Blick Chthonius (Chthonius) ksenemanni Hadži: et al., 2004: –. Krumpál & Kiefer, 1981: 128. Chthonius (Chthonius) ischnocheles (Hermann):

Chthonius (Chthonius) heterodactylus Šťáhlavský, 1996: 4; Jagoš, 2001: 33–34; Harvey, Tömösváry, 1882 2009: –; Christophoryová et al., 2011b: 36. CR: Chthonius heterodactylus Tömösváry: Ducháč & SR: Chthonius ischnocheles (Hermann): Krum- Mlejnek, 2004: 37–38; Mikula, 2006: 18; Laška et al., pál, 2000: 96; Krumpál & Krumpálová, 2003: 119; 2008: 171. Christophoryová, 2009a: 254.

Checklist of pseudoscorpions from the Czech Republic and Slovakia 5

Chthonius (Chthonius) ischnocheles (Hermann): Chthonius (Chthonius) submontanus Beier, 1963 Krumpál, 1980: 23; Krumpál & Kiefer, 1981: 128; CR (note 4): not Chthonius cf. submontanus Beier: Šťáhlavský, 1996: 4; Jagoš, 2001: 33–34; Harvey, Ducháč, 2002c: 102 (misidentification, see Chtho- 2009: –; Christophoryová et al., 2011b: 36. nius carinthiacus).

Chthonius (Chthonius) ischnocheles ischnocheles Chthonius (Chthonius) subterraneus Beier, 1931 (Hermann, 1804) SR (note 5): Chthonius (Chthonius) subterraneus CR: Chthonius (Chthonius) ischnocheles ischnocheles Beier: Christophoryová et al., 2011b: 36. (Hermann): Harvey, 2009: –. Chthonius subterraneus Beier: Krumpál & SR: Chthonius (Chthonius) ischnocheles Krumpálová, 2003: 119. ischnocheles (Hermann): Harvey, 2009: –. Chthonius (Chthonius) tenuis L. Koch, 1873

Chthonius (Chthonius) orthodactylus CR (note 6): Chthonius tenuis L. Koch: Stecker, (Leach, 1817) 1874: 238–239; Simon, 1879: 72–73; Nosek, 1901: 21; CR: Chthonius orthodactylus (Leach): Krásný, 1993: Nosek, 1902: 60; Caporiacco, 1926: 76; Krásný, 37; Ducháč, 1998b: –; Ducháč, 1999b: 14; Ducháč, 1993: 38; Ducháč, 1998b: –; Ducháč, 1999b: 14; 2002c: 17, 18, 19, 68, 82, 138; Blick et al., 2004: –; Ducháč, 2002c: 12, 13, 19, 68, 82, 141; Blick et al., Christophoryová, 2006b: 85; Mikula, 2006: 18. 2004: –. Chthonius (Chthonius) orthodactylus (Leach): Chthonius (Chthonius) tenuis L. Koch: Šťáh- Studničková, 1968: 25; Krumpál & Kiefer, 1981: lavský, 1996: 6; Šťáhlavský & Krásný, 2007: 430; 128; Ducháč, 1989: 180; Ducháč, 1994a: 140; Šťáh- Harvey, 2009: –; Christophoryová et al., 2011b: 36. lavský, 1996: 5–6; Šťáhlavský, 2000: 17; Šťáh- not Chthonius (Chthonius) tenuis L. Koch: Šťáh- lavský, 2001: 80; Ducháč, 2002c: 31, 43; Šťáhlavský lavský, 2000: 17; Šťáhlavský, 2001: 80; Šťáhlavský & Král, 2004: 50; Šťáhlavský, 2006c: 37, 79; Šťáh- & Král, 2004: 50; Šťáhlavský, 2006a: 169; Šťáh- lavský & Krásný, 2007: 430; Harvey, 2009: –; Chris- lavský, 2006c: 37, 79 (all misidentification, see tophoryová et al., 2011b: 36. Chthonius carinthiacus). SR: Chthonius (Chthonius) orthodactylus SR: Chthonius tenuis L. Koch: Krumpál & (Leach): Krumpál, 1980: 23; Krumpál & Kiefer, Krumpálová, 2003: 119; Mock et al., 2004: 38; Mock 1981: 128; Šťáhlavský, 1996: 5; Harvey, 2009: –; et al., 2005: 100; Christophoryová, 2009a: 254. Christophoryová et al., 2011b: 36. Chthonius (Chthonius) tenuis L. Koch: Šťáh-

Chthonius (Chthonius) pygmaeus Beier, 1934 lavský, 1996: 6; Christophoryová et al., 2011b: 36.

SR: Chthonius (Neochthonius) pygmaeus Beier: Subgenus Chthonius (Ephippiochthonius) Krumpál & Kiefer, 1981: 127; Kárpáthegyi, 2007b: Beier, 1930 92. Chthonius (Ephippiochthonius) boldorii Chthonius pygmaeus Beier: Ducháč, 2002c: 139. Beier, 1934 Chthonius (Neochthonius) pygmaeus pygmaeus SR: Chthonius (Ephippiochthonius) boldorii Beier: Beier: Beier, 1963: 23. Christophoryová & Krumpál, 2005: 79; Christo- Chthonius (Chthonius) pygmaeus Beier: Kár- phoryová, 2006a: 63; Christophoryová, 2006b: 24; páthegyi, 2007a: 82–83; Christophoryová et al., Christophoryová & Krumpál, 2006a: 43; Christo- 2011b: 36. phoryová & Krumpál, 2007a: 14; Christophoryová Chthonius aff. pygmaeus Beier: Krumpál & & Krumpál, 2007b: 29; Christophoryová et al., Krumpálová, 2003: 120. 2007b: 20; Christophoryová, 2008b: 22; Christo- Chthonius (Chthonius) ressli Beier, 1956 phoryová & Krištofík, 2008: 22; Christophoryová CR (note 3): Chthonius (Chthonius) ressli Beier: et al., 2011b: 36. Šťáhlavský & Ducháč, 2005: 83; Šťáhlavský, 2006a: Chthonius boldorii Beier: Christophoryová & 169; Harvey, 2009: –; Christophoryová et al., Krumpál, 2006b: 4; Christophoryová, 2008a: 133; 2011b: 36. Christophoryová & Krumpál, 2009: 15. SR (note 3): Chthonius (Chthonius) ressli Beier: Chthonius (Ephippiochthonius) fuscimanus Christophoryová et al., 2011b: 36. Simon, 1900 Chthonius ressli Beier: Christophoryová, 2008a: CR: Chthonius austriacus Beier, 1931: Ducháč, 132. 1998b: –; Ducháč, 1999b: 14; Ducháč, 2002c: 16, 82, Chthonius aff. ressli Beier: Mock et al., 2004: 38. 136 (synonymized by Gardini, 1980: 261). Chthonius cf. ressli Beier: Mock et al., 2005: 100; Chthonius (Ephippiochthonius) austriacus austria- Christophoryová, 2009a: 254.

6 Christophoryová, J. et al. cus Beier, 1931: Rafalski, 1967: 7. Ducháč, 1994a: 140–141; Šťáhlavský, 1996: 7–8; Chthonius (Ephippiochthonius) fuscimanus Šťáhlavský, 2000: 18; Jagoš, 2001: 34–35, 48; Šťáh- Simon: Šťáhlavský, 1996: 9; Šťáhlavský, 2001: 80– lavský, 2001: 81–82; Ducháč, 2002c: 26, 30–31, 43– 81; Muster et al., 2004: 306; Šťáhlavský & Král, 44; Šťáhlavský & Král, 2004: 50; Šťáhlavský, 2006a: 2004: 50; Šťáhlavský, 2006a: 169; Šťáhlavský, 169–170; Šťáhlavský, 2006c: 37, 80; Christophory- 2006c: 37, 80; Harvey, 2009: –; Šťáhlavský & Tuf, ová et al., 2007b: 19; Šťáhlavský & Krásný, 2007: 2009: 98; Christophoryová et al., 2011b: 36. 430; Harvey, 2009: –; Šťáhlavský & Tuf, 2009: 98; Chthonius fuscimanus Simon: Ducháč, 1995c: –; Christophoryová, 2010: 4; Christophoryová et al., Ducháč, 1997b: –; Ducháč, 2002c: 67, 68, 124; Blick 2011b: 36. et al., 2004: –; Christophoryová, 2006b: 9, 85; SR: Chthonius tetrachelatus (Preyssler): Daday, Laška, 2006: 19. 1888a: 134; Daday, 1888b: 190–191; Daday, 1918: 2; SR: Chthonius (Ephippiochthonius) austriacus Krumpál & Cyprich, 1988: 42; Ducháč, 1994a: 146; austriacus Beier: Krumpál & Kiefer, 1981: 127 (syn- Fenďa et al., 1998: 27; Ducháč, 2002c: 49; Krumpál onymized by Gardini, 1980: 261). & Krumpálová, 2003: 119–120; Christophoryová, Chthonius (Ephippiochthonius) fuscimanus Si- 2006b: 8, 84; Christophoryová & Krumpál, 2006b: mon: Šťáhlavský, 1996: 9; Christophoryová & 4; Christophoryová, 2008a: 133; Christophory- Krumpál, 2005: 79; Christophoryová, 2006a: 63; ová & Mock, 2009: 16. Christophoryová, 2006b: 24; Christophoryová & Chthonius (Ephippiochthonius) tetrachelatus Krumpál, 2006a: 43; Christophoryová & Krumpál, (Preyssler): Szent-Ivány, 1941: 86–87 (as Chthonius 2007a: 14; Christophoryová & Krumpál, 2007b: 29; (Epiphiochthonius) [sic] tetrachelatus); Studničková, Christophoryová et al., 2007b: 20. 1968: 24–25; Krumpál & Kiefer, 1981: 127; Ducháč, Chthonius fuscimanus Simon: Christophoryová, 1989: 179–180; Šťáhlavský, 1996: 7; Jagoš, 2001: 34– 2006b: 8, 84; Christophoryová & Krumpál, 2006b: 35; Ducháč, 2002c: 30–31; Christophoryová, 2006a: 4; Christophoryová, 2008a: 133; Christophoryová 63; Christophoryová, 2006b: 24; Christophoryová et al., 2011b: 36. & Krumpál, 2007a: 14; Christophoryová & Krum-

Chthonius (Ephippiochthonius) kewi pál, 2007b: 29; Harvey, 2009: –; Christophoryová, Gabbutt, 1966 2010: 4; Christophoryová et al., 2011b: 36. CR (note 7): Chthonius (Ephippiochthonius) kewi Chthonius (Ephippiochthonius) cf. tetrachelatus Gabbutt: Harvey, 2009: –. (Preyssler): Christophoryová & Krumpál, 2006a: Chthonius cf. kewi Gabbutt: Ducháč, 1996c: 81– 43.

84; Ducháč, 1997b: –; Ducháč, 2002c: 69–72, 127, Genus Mundochthonius Chamberlin, 1929 138; Christophoryová, 2006b: 9. Mundochthonius carpaticus Rafalski, 1948 Chthonius (Ephippiochthonius) tetrachelatus CR: Mundochthonius carpaticus Rafalski: Christo- (Preyssler, 1790) phoryová et al., 2011b: 36, 38. CR: Scorpio tetrachelatus Preyssler: Preyssler, 1790: SR: Mundochthonius carpaticus Rafalski: Krum- 56; Ducháč, 2002c: 7. pál & Krumpálová, 2003: 118; Christophoryová, Chthonius trombidioides (Latreille, 1804): 2008a: 133; Christophoryová, 2009a: 256; Christo- Stecker, 1874: 238; Ducháč, 2002c: 12 (misidentifi- phoryová, 2009b: 129; Christophoryová & Mock, cation). 2009: 16; Mock et al., 2009: 268; Papáč et al., 2009: Chthonius tetrachelatus (Preyssler): Simon, 1879: 37; Christophoryová, 2010: 4; Christophoryová et 70–71; Nosek, 1901: 21; Nosek, 1902: 60–61; al., 2011b: 36. Ducháč, 1994b: –; Ducháč, 1995a: 52; Ducháč, 1995d: 130; Růžička et al., 1995: 47; Ducháč, 1996c: Mundochthonius styriacus Beier, 1971 82; Šťáhlavský, 1998: –; Ducháč, 1999b: 14; Ducháč, CR: Mundochthonius styriacus Beier: Ducháč, 1998b: 2002c: 13, 19, 65, 68, 82, 102, 110–111, 141–143; –; Šťáhlavský, 1998: –; Ducháč, 1999b: 14; Šťáh- Blick et al., 2004: –; Christophoryová, 2006b: 9, 85; lavský, 2000: 19; Šťáhlavský, 2001: 83; Šťáhlavský Laška, 2006: 19; Mikula, 2006: 18; Šťáhlavský, & Ducháč, 2001: 47; Ducháč, 2002c: 19, 82, 143–144; 2006b: 161; Gardini, 2009: 24; Christophoryová, Blick et al., 2004: –; Drogla & Lippold, 2004: 18; 2010: 8. Šťáhlavský & Král, 2004: 50; Šťáhlavský & Ducháč, Chthonius (Ephippiochthonius) tetrachelatus 2005: 83; Šťáhlavský, 2006c: 37, 80; Petrov & Šťáh- (Preyssler): Ducháč, 1988: 105; Ducháč, 1989: 179– lavský, 2007: 16; Harvey, 2009: –; Christophory- 180; Harvey, 1991: 172–173; Krásný, 1993: 39–40; ová, 2010: 8; Christophoryová et al., 2011b: 36.

Checklist of pseudoscorpions from the Czech Republic and Slovakia 7

Suborder IOCHEIRATA Harvey, 1992 Neobisium (Neobisium) beieri Verner, 1958 SR: Neobisium (Neobisium) beieri Verner: Verner, Superfamily NEOBISIOIDEA Chamberlin, 1930 1958b: 198–199; Beier, 1963: 101; Studničková,

1968: 20; Verner, 1971: 25; Harvey, 1991: 347; Har- Family NEOBISIIDAE Chamberlin, 1930 vey, 2009: –; Christophoryová et al., 2011b: 36. Subfamily Neobisiinae Chamberlin, 1930 Neobisium beieri Verner: Krumpál, 1980: 23; Genus Microbisium Chamberlin, 1930 Christophoryová, 2006b: 7; Christophoryová et al.,

Microbisium brevifemoratum (Ellingsen, 1903) 2007c: 11.

CR: Microbisium brevifemoratum (Ellingsen): Chris- Neobisium (Neobisium) brevidigitatum tophoryová et al., 2011b: 36, 38. (Beier, 1928)

Microbisium suecicum Lohmander, 1945 SR: Neobisium (Neobisium) brevidigitatum (Beier): CR: Microbisium suecicum Lohmander: Ducháč, Harvey, 2009: –; Christophoryová et al., 2011b: 36. 1998b: –; Šťáhlavský & Ducháč, 2001: 47; Ducháč, Neobisium brevidigitatum (Beier): Krumpál, 2002c: 151; Blick et al., 2004: –; Drogla & Lippold, 1980: 23; Krumpál & Krumpálová, 2003: 120; Christophoryová, 2006b: 8. 2004: 18; Šťáhlavský & Ducháč, 2005: 83; Christo- phoryová, 2006b: 9; Šťáhlavský, 2006a: 170; Har- Neobisium (Neobisium) carcinoides vey, 2009: –; Christophoryová et al., 2011b: 36. (Hermann, 1804) SR: Microbisium suecicum Lohmander: Chris- CR: Obisium muscorum Leach, 1817: Stecker, 1874: tophoryová, 2009b: 129; Christophoryová & 241; Nosek, 1901: 21; Nosek, 1902: 66, 70; Nosek, Krumpál, 2009: 14; Christophoryová et al., 2011b: 1957: 59; Ducháč, 2002c: 12, 13, 14 (synonymized 36, 38. by Beier, 1963: 115). Obisium carcinoides (Hermann): Stecker, 1874: Genus Neobisium Chamberlin, 1930 240; Nosek, 1901: 21; Nosek, 1902: 64; Ducháč,

Subgenus Neobisium (Blothrus) Schiödte, 1847 2002c: 12, 13. Neobisium muscorum (Leach): Verner, 1957: 88; Neobisium (Blothrus) slovacum Gulička, 1977 Verner, 1959b: 379; Ducháč, 1999a: 105; Ducháč, SR: Neobisium (Blothrus) slovacum Gulička: 2002c: 16; Christophoryová, 2006b: 9, 85; Christo- Gulička, 1977a: 6–8; Gulička, 1977b: 27–28; phoryová et al., 2007b: 19. Gulička, 1985: 111; Harvey, 1991: 383; Ducháč, Neobisium (Neobisium) muscorum (Leach): 1996a: 153–157; Ducháč, 1998c: 112; Ducháč, 1999c: Verner, 1971: 25. 175–180; Ducháč, 2000: 9–10; Ducháč & Mlejnek, Neobisium (Neobisium) carcinoides (Hermann): 2000a: 15–20; Ducháč & Mlejnek, 2000b: 48–49; Studničková, 1968: 16–18; Ducháč, 1988: 105; Krumpál, 2000: 95; Ducháč, 2001a: 8; Ducháč, Ducháč, 1989: 180–181; Ducháč, 1994a: 141–142; 2001b: 1–142; Ducháč, 2002d: 18; Mlejnek & Šťáhlavský, 1996:14–17; Šumpich et al., 1999: 163; Ducháč, 2002: 125–126; Ducháč, 2004: 52–53; Chris- Ducháč, 2002c: 26, 31–32, 44–45; Harvey, 2009: –; tophoryová, 2006b: 7; Christophoryová & Krum- Šťáhlavský & Tuf, 2009: 98–99; Christophoryová, pál, 2006c: 20; Christophoryová et al., 2007c: 11; 2010: 5; Christophoryová et al., 2011b: 36. Harvey, 2009: –; Christophoryová, 2009a: 254, 256; Neobisium carcinoides (Hermann): Ducháč, Christophoryová et al., 2011b: 36. 1993b: 66; Krásný, 1993: 42–45; Ducháč, 1994b: –; Neobisium slovacum Gulička: Gulička, 1982: Ducháč, 1995a: 52; Ducháč, 1995d: 130; Růžička et 174; Ducháč, 1997c: 10–11; Vlk et al., 2001: 68 (as al., 1995: 47; Ducháč, 1997a: 117; Ducháč, 1999a: Neobissium [sic] slovacum); Papáč et al., 2007a: 52; 105–113; Ducháč, 1999b: 14; Šťáhlavský, 2000: 20; Christophoryová, 2008a: 133. Jagoš, 2001: 29–30; Šťáhlavský, 2001: 84–85; Neobisium (Blothrus) cf. slovacum Gulička: Ducháč, 2002a: 58–59; Ducháč, 2002b: 24; Ducháč, Ducháč, 1996b: 51–52. 2002c: 17, 18, 35, 65, 68, 76, 82, 83–91, 93, 102, 103, Neobisium cf. slovacum Gulička: Ducháč, 1997c: 107, 110–111, 113–114, 116–117, 118, 119–120, 122– 10–11. 123, 128–129, 130, 131, 132, 134, 144–147; Ducháč, Neobisium (Blothrus) spelaeum (Schiödte, 1847) 2003: 3; Blick et al., 2004: –; Laška, 2006: 19; Mi- CR (note 8): Blothrus spelaeus Schiödte: Stecker, kula, 2006: 18; Šťáhlavský, 2006a: 170–172; Šťáh- 1875: 168–169. lavský, 2006b: 163; Šťáhlavský, 2006c: 70, 80; Šťáh-

Subgenus Neobisium (Neobisium) lavský, 2007: 60; Šťáhlavský & Krásný, 2007: 430– Chamberlin, 1930 432; Božanić, 2008: 6; Christophoryová, 2010: 8.

8 Christophoryová, J. et al.

Neobisium muscorum germanicum Beier, 1931: Neobisium (Neobisium) carpaticum Beier, 1935 Verner, 1957: 88. SR: Neobisium (Neobisium) carpaticum Beier: Chris- SR: Obisium muscorum Leach: Daday, 1888a: tophoryová, 2006a: 63; Christophoryová, 2006b: 132; Daday, 1888b: 188; Ellingsen, 1910: 400; 24; Christophoryová & Krumpál, 2007b: 29; Har- Nosek, 1957: 84 (synonymized by Beier, 1963: 115). vey, 2009: –; Christophoryová, 2010: 5; Christo- Obisium carcinoides (Hermann): Tömösváry, phoryová et al., 2011b: 36. 1882: 232–233; Daday, 1888a: 131; Daday, 1888b: Neobisium carpaticum Beier: Krumpál, 1980: 24; 187; Daday, 1918: 2. Krumpál, 2000: 96; Krumpál & Krumpálová, 2003: Obisium (Obisium) muscorum Leach: Beier, 120; Christophoryová, 2006b: 8; Christophoryová 1928: 288–289. & Krumpál, 2006b: 4; Christophoryová, 2009b: 130; Neobisium muscorum (Leach): Verner, 1958b: Christophoryová & Mock, 2009: 17. 198; Krumpál & Cyprich, 1988: 42; Fenďa et al., Neobisium (Neobisium) erythrodactylum (L. 1998: 24; Mock et al., 2003: 37; Christophoryová & Koch): Christophoryová & Krumpál, 2005: 79; Krumpál, 2004: 10; Christophoryová, 2006b: 8; Christophoryová & Krumpál, 2006a: 43; Christo- Christophoryová & Krumpál, 2006b: 4; Kováč et phoryová et al., 2007b: 20; Christophoryová, al., 2007: 49. 2008b: 22 (all misidentification).

Neobisium (Neobisium) muscorum (Leach): Neobisium (Neobisium) crassifemoratum Verner, 1971: 25; Christophoryová & Krumpál, (Beier, 1928) 2005: 79; Christophoryová, 2006a: 63; Christo- CR: Neobisium (Neobisium) crassifemoratum (Beier): phoryová, 2006b: 24; Christophoryová & Krumpál, Christophoryová et al., 2011b: 36. 2006a: 43; Christophoryová & Krumpál, 2006c: 20; Neobisium crassifemoratum (Beier): Ducháč, Christophoryová & Krumpál, 2007b: 29; Christo- 2002b: 24; Ducháč, 2002c: 93, 147–148; Ducháč, phoryová et al., 2007b: 20. 2003: 3–4; Blick et al., 2004: –; Laška, 2006: 19–20. Neobisium (Neobisium) carcinoides (Hermann): Neobisium cf. crassifemoratum (Beier): Ducháč, Studničková, 1968: 16–18; Ducháč, 1989: 180–181; 2002c: 132. Šťáhlavský, 1996: 14; Jagoš, 2001: 29–30; Ducháč, SR: Neobisium (Neobisium) crassifemoratum 2002c: 31–32; Christophoryová, 2008b: 22; Christo- (Beier): Verner, 1958a: 109–110; Verner, 1960: 167; phoryová & Krištofík, 2008: 22; Christophoryová, Beier, 1963: 100–101; Rafalski, 1967: 8; Harvey, 2009a: 256; Harvey, 2009: –; Christophoryová, 2009: –; Christophoryová et al., 2011b: 36. 2010: 5; Christophoryová & Krumpál, 2010: 107; Neobisium crassifemoratum (Beier): Krumpál, Christophoryová et al., 2011b: 36. 1980: 24; Krumpál & Cyprich, 1988: 42; Fenďa et Neobisium carcinoides (Hermann): Krumpál, al., 1998: 27; Ducháč, 2003: 3; Krumpál & Krum- 1980: 23–24; Fenďa et al., 1998: 27; Krumpál, 2000: pálová, 2003: 121; Farkač et al., 2004: 221; Christo- 96; Krumpál et al., 2000–2001: 52; Ducháč, 2002c: phoryová, 2006b: 7; Benová & Svatoň, 2009: 153. 144–145; Krumpál & Krumpálová, 2003: 120; Neobisium (Neobisium) erythrodactylum Farkač et al., 2004: 221; Šťáhlavský, 2006c: 80; (L. Koch, 1873) Christophoryová, 2008a: 133; Benová & Svatoň, CR: Obisium erythrodactylum L. Koch: Stecker, 2009: 153; Christophoryová, 2009b: 129–130; Chris- 1874: 240–241; Nosek, 1901: 21; Nosek, 1902: 65; tophoryová & Mock, 2009: 16; Mock et al., 2009: Ducháč, 2002c: 12, 13. 268; Papáč et al., 2009: 37; Christophoryová & Neobisium (Neobisium) erythrodactylum (L. Krumpálová, 2010: 108. Koch): Studničková, 1968: 21; Ducháč, 1989: 181; Neobisium cf. muscorum (Leach): Papáč et al., Ducháč, 1994a: 143; Šťáhlavský, 1996: 11–12; 2007b: 154. Ducháč, 2002c: 32, 46; Harvey, 2009: –; Šťáhlavský Neobisium (Neobisium) carcinoides carcinoides & Tuf, 2009: 99–100; Christophoryová et al., 2011b: (Hermann, 1804) 36. CR: Neobisium (Neobisium) muscorum muscorum Neobisium erythrodactylum (L. Koch): Verner, (Leach, 1817): Hadži, 1939: 193–196. 1957: 88; Verner, 1959b: 379; Ducháč, 1997a: 118; Neobisium (Neobisium) carcinoides carcinoides Ducháč, 1999b: 14; Šťáhlavský, 2000: 21; Jagoš, (Hermann): Harvey, 2009: –. 2001: 31, 46; Šťáhlavský, 2001: 85–86; Ducháč, SR: Neobisium (Neobisium) carcinoides carci- 2002b: 24; Ducháč, 2002c: 16, 17, 68, 77, 82, 93, 148; noides (Hermann): Harvey, 2009: –. Ducháč, 2003: 3, 4; Blick et al., 2004: –; Christo- phoryová, 2006b: 9, 85; Mikula, 2006: 18; Šťáh-

Checklist of pseudoscorpions from the Czech Republic and Slovakia 9 lavský, 2006a: 172; Šťáhlavský, 2006b: 163; Šťáh- 2001: 32–33; Harvey, 2009: –; Christophoryová et lavský, 2006c: 81; Christophoryová et al., 2007b: al., 2011b: 37. 19. Neobisium fuscimanum (C. L. Koch): Krumpál, SR: Obisium erythrodactylum L. Koch: Tömös- 1980: 24; Krumpál, 2000: 96; Krum- váry, 1882: 229–230; Daday, 1888a: 132; Daday, pál & Krumpálová, 2003: 121; Kárpáthegyi & 1888b: 188; Ellingsen, 1910: 397; Daday, 1918: 2. Kontschán, 2005: 5; Christophoryová, 2006b: 7; Obisium (Obisium) erythrodactylum L. Koch: Christophoryová & Mock, 2009: 17; Mock et al., Beier, 1928: 288–289. 2009: 268.

Neobisium (Neobisium) erythrodactylum (L. Neobisium (Neobisium) inaequale Chamberlin, Koch): Šťáhlavský, 1996: 11; Jagoš, 2001: 31; Har- 1930 vey, 2009: –; Christophoryová et al., 2011b: 36. SR: Neobisium inaequale Chamberlin: Krumpál & Neobisium erythrodactylum (L. Koch): Krumpál, Cyprich, 1988: 42; Fenďa et al., 1998: 27; Krištofík 1980: 24; Krumpál & Krumpálová, 2003: 121; et al., 2005: 172 (as Neobium [sic] inaequale); Chris- Krumpál & Christophoryová, 2007: 95. tophoryová, 2010: 2. Not Neobisium (Neobisium) erythrodactylum (L. Neobisium (Neobisium) inaequale Chamberlin: Koch): Christophoryová & Krumpál, 2005: 79; Harvey, 2009: –; Christophoryová et al., 2011b: 37. Christophoryová & Krumpál, 2006a: 43; Christo- Neobisium (Neobisium) jugorum (L. Koch, 1873) phoryová et al., 2007b: 20; Christophoryová, CR (note 9): Obisium jugorum L. Koch: Simon, 2008b: 22 (all misidentification, see Neobisium 1879: 60–61; Caporiacco, 1926: 75. (Neobisium) carpaticum). SR (note 9): Neobisium (Neobisium) jugorum (L. Neobisium (Neobisium) fuscimanum (C. L. Koch, Koch): Verner, 1960: 167; Rafalski, 1967: 11; 1843) Verner, 1971: 25; Christophoryová et al., 2011b: 37. CR: Obisium fuscimanum C. L. Koch: Stecker, 1874: Neobisium jugorum (L. Koch): Christophory- 240; Nosek, 1901: 21; Nosek, 1902: 65; Ducháč, ová, 2006b: 7. 2002c: 12, 13. Neobisium (Neobisium) macrodactylum (Daday, Obisium (Obisium) fuscimanum C. L. Koch: 1888) Beier, 1928: 295. SR (note 10): Neobisium (Neobisium) macrodactylum Neobisium (Neobisium) fuscimanum (C. L. Koch): (Daday): Harvey, 2009: –; Christophoryová et al., Beier, 1932a: 92–93; Rafalski, 1967: 12; Studničk- 2011b: 37. ová, 1968: 19–20; Ducháč, 1989: 181; Ducháč, Neobisium macrodactylum (Daday): Krumpál, 1994a: 142–143; Šťáhlavský, 1996: 12–13; Ducháč, 1980: 24; Christophoryová, 2006b: 8. 2002c: 32, 45–46; Harvey, 2009: –; Christophoryová et al., 2011b: 37. Neobisium (Neobisium) macrodactylum macro- Neobisium fuscimanum (C. L. Koch): Verner, dactylum (Daday, 1888) 1957: 87; Beier, 1963: 112–113; Ducháč, 1995b: 145– SR: Neobisium (Neobisium) macrodactylum macrodac- 151; Ducháč, 1999b: 14; Šťáhlavský, 2000: 21; Jagoš, tylum (Daday): Harvey, 2009: –.

2001: 32–33; Šťáhlavský, 2001: 86; Ducháč, 2002b: Neobisium (Neobisium) polonicum Rafalski, 1936 24; Ducháč, 2002c: 17, 57–63, 68, 82, 93, 107, 111, SR: Neobisium polonicum Rafalski: Ducháč, 1995e: 117, 130, 149–150; Ducháč, 2003: 3, 4; Blick et al., 11–17; Ducháč, 1999d: 207–209; Krumpál, 2000: 95; 2004: –; Kárpáthegyi & Kontschán, 2005: 5; Chris- Krumpál & Krumpálová, 2003: 121–122; Christo- tophoryová, 2006b: 85; Laška, 2006: 20; Šťáhlavský, phoryová, 2006b: 7; Šťáhlavský, 2006c: 81; Vařil, 2006a: 172; Šťáhlavský, 2006b: 163; Šťáhlavský, 2007: 15; Murienne et al., 2008: 174. 2006c: 81. Neobisium (Neobisium) polonicum Rafalski: Neobisium fuscimanum ponticum Beier: Ducháč, Krumpál, 1979: 429–435; Harvey, 2009: –; Christo- 2002c: 16. phoryová et al., 2011b: 37.

Neobisium (Neobisium) fuscimanum ponticum Neobisium (Neobisium) simile (L. Koch, 1873) Beier, 1963: Rafalski, 1967: 12. SR (note 11): Obisium simile L. Koch: Tömösváry, Neobisium fuscimanum fuscimanum (C. L. Koch): 1882: 225–226. Ducháč, 2002c: 16. Neobisium (Neobisium) simile (L. Koch): Chris- SR: Obisium (Obisium) fuscimanum C. L. Koch: tophoryová et al., 2011b: 37. Beier, 1928: 295. Neobisium (Neobisium) sylvaticum (C. L. Koch, Neobisium (Neobisium) fuscimanum (C. L. Koch): 1835) Szent-Ivány, 1941: 87; Šťáhlavský, 1996: 12; Jagoš, 10 Christophoryová, J. et al.

CR: Obisium sylvaticum C. L. Koch: Stecker, 1874: Obisium (Obisium) validum (L. Koch): Daday, 239–240 (as Obisium silvaticum [sic]); Ducháč, 1888a: 130; Daday, 1888b: 186. 2002c: 12 (as Obisium silvaticum [sic]). Neobisium validum (L. Koch): Christophoryová, Obisium dumicola C. L. Koch: Nosek, 1901: 21; 2008a: 133. Nosek, 1902: 64, 70, Ducháč, 2002c: 13, 14 (syn- Genus Roncus L. Koch, 1873 onymized by Beier, 1963: 83). Neobisium sylvaticum (C. L. Koch): Verner, Roncus alpinus L. Koch, 1873 1957: 87; Verner, 1959b: 379; Krásný, 1993: 48; CR (note 13): Roncus alpinus L. Koch: Cârdeiu, Ducháč, 1994b: –; Ducháč, 1995a: 52; Ducháč, 1948: 4. 1997a: 118; Ducháč, 1999b: 14; Šťáhlavský, 2000: Obisium alpinum (L. Koch): Simon, 1879: 65; 21–22; Jagoš, 2001: 32; Šťáhlavský, 2001: 86; Caporiacco, 1922: 67. Ducháč, 2002b: 24; Ducháč, 2002c: 16, 17, 18–19, 65, SR (note 13): Roncus alpinus L. Koch: Tömös- 68, 77, 82, 93, 108, 114, 116–117, 120, 122–123, 128– váry, 1882: 217–218; Daday, 1918: 2; Krumpál 129, 130, 132, 134, 150–151; Ducháč, 2003: 3, 4; & Krumpálová, 2003: 122; Christophoryová & Blick et al., 2004: –; Christophoryová, 2006b: 9, 85; Šťáhlavský, 2007: 12; Christophoryová, 2008a: 133.

Mikula, 2006: 18; Šťáhlavský, 2006a: 172; Šťáh- Roncus lubricus L. Koch, 1873 lavský, 2006b: 163; Christophoryová et al., 2007b: CR: Roncus lubricus L. Koch: Stecker, 1874: 237; 19; Šťáhlavský & Krásný, 2007: 432; Božanić, 2008: Ducháč, 1998b: –; Šťáhlavský, 1998: –; Šťáhlavský, 6. 2000: 22; Šťáhlavský, 2001: 86–87; Ducháč, 2002c: Neobisium (Neobisium) sylvaticum (C. L. Koch): 11, 13; Harvey, 2009: –; Christophoryová et al., Ducháč, 1988: 105; Harvey, 1991: 369–371; Šťáh- 2011b: 37. lavský, 1996: 10–11; Ducháč, 2002c: 26; Harvey, Obisium (Roncus) lubricum (L. Koch): Nosek, 2009: –; Šťáhlavský & Tuf, 2009: 100; Christo- 1902: 65. phoryová, 2010: 5; Christophoryová et al., 2011b: Roncus lubricus lubricus L. Koch: Blick et al., 37. 2004: –. Neobisium (Neobisium) sylvaticum inaculeatum Roncus „lubricus“ L. Koch: Šťáhlavský & Hadži, 1939: Harvey, 1991: 370–371; Harvey, 2009: Ducháč, 2001: 47–48. – (synonymized by Schawaller, 1989: 9). Roncus cf. lubricus L. Koch: Ducháč, 2002c: 20, Neobisium (Neobisium) sylvaticum sylvaticum (C. 151. L. Koch): Studničková, 1968: 14–16. SR: Roncus lubricus L. Koch: Tömösváry, 1882: SR: Obisium sylvaticum C. L. Koch: Tömösváry, 215–217; Verner, 1971: 26; Ducháč, 2002c: 151; 1882: 222–224; Daday, 1918: 2. Krumpál & Krumpálová, 2003: 122–123; Christo- Obisium dumicola C. L. Koch: Tömösváry, 1882: phoryová & Krumpál, 2005: 79; Christophoryová, 226–227; Daday, 1888a: 131; Daday, 1888b: 187 2006a: 63; Christophoryová, 2006b: 8, 24; Christo- (synonymized by Beier, 1963: 83). phoryová & Krumpál, 2006b: 4; Šťáhlavský, 2006c: Obisium (Obisium) dumicola nitidum Daday, 81; Christophoryová & Krumpál, 2007b: 29; Chris- 1988: Daday, 1888a: 131–132; Daday, 1888b: 187– tophoryová et al., 2007b: 20; Christophoryová & 188 (synonymized Beier, 1932a: 137). Šťáhlavský, 2007: 12; Christophoryová, 2008a: 133; Neobisium sylvaticum (C. L. Koch): Ducháč, Christophoryová, 2008b: 22; Christophoryová, 1998c: 112–113; Krumpál, 2000: 97; Krištofík et al., 2009b: 130–131; Harvey, 2009: –; Papáč et al., 2009: 2002: 605; Krumpál & Krumpálová, 2003: 122; 37; Christophoryová et al., 2011b: 37. Christophoryová, 2009b: 130; Christophoryová & Obisium lubricum (L. Koch): Daday, 1888a: Mock, 2009: 17; Papáč et al., 2009: 37; Christo- 128–129; Daday, 1888b: 184–185. phoryová, 2010: 2. Roncus (Roncus) lubricus L. Koch: Szent-Ivány, Neobisium (Neobisium) sylvaticum (C. L. Koch): 1941: 88; Ducháč, 1994a: 143; Ducháč, 2002c: 46. Šťáhlavský, 1996: 10; Jagoš, 2001: 32; Christo- Roncus (Roncus) lubricus lubricus L. Koch: phoryová, 2008b: 22; Christophoryová, 2009a: 256; Beier, 1963: 182. Harvey, 2009: –; Christophoryová, 2010: 5; Chris- Roncus transsilvanicus Beier, 1928 tophoryová et al., 2011b: 37. CR (note 14): Roncus transsilvanicus Beier: Harvey, Neobisium (Neobisium) validum (L. Koch, 1873) 2009: –. SR (note 12): Obisium validum L. Koch: Daday, SR (note 14): Roncus transsilvanicus Christo- 1918: 2. phoryová & Šťáhlavský, 2007: 12; Christophory-

Checklist of pseudoscorpions from the Czech Republic and Slovakia 11 ová, 2008a: 133; Murienne et al., 2008: 174; Harvey, Genus Cheiridium Menge, 1855 2009: –; Christophoryová et al., 2011b: 37. Cheiridium museorum (Leach, 1817) Roncus aff. transsilvanicus Beier: Krumpál & CR: Cheiridium museorum (Leach): Stecker, 1874: Krumpálová, 2003: 123. 232; Nosek, 1901: 21; Nosek, 1902: 49, 70; Verner, 1957: 88; Studničková, 1968: 22; Verner, 1971: 27; Family SYARINIDAE Chamberlin, 1930 Krásný, 1993: 33–34; Ducháč, 1994a: 143; Ducháč, Subfamily Syarininae Chamberlin, 1930 1995d: 130; Šťáhlavský, 1996: 19–20; Ducháč, Genus Syarinus Chamberlin, 1925 1999b: 14 (as Cheridium [sic] museorum); Šťáh- Syarinus strandi (Ellingsen, 1901) lavský, 2000: 4–5; Šťáhlavský, 2001: 87–88; Ducháč, CR: Syarinus strandi (Ellingsen): Ducháč, 1998a: 2002c: 11 (as Chernes [sic] museorum), 13, 14, 18, 19, 506–508; Ducháč, 1998b: –; Ducháč, 1999b: 14; 46, 68, 82, 161; Blick et al., 2004: –; Šťáhlavský, Šťáhlavský & Ducháč, 2001: 48; Ducháč, 2002c: 79– 2006b: 163; Šťáhlavský, 2006c: 78; Šťáhlavský & 81, 82, 152, 159; Blick et al., 2004: –; Drogla & Lip- Krásný, 2007: 432; Harvey, 2009: –; Christophory- pold, 2004: 25; Šťáhlavský & Ducháč, 2005: 83; ová et al., 2011b: 37. Harvey, 2009: –; Christophoryová et al., 2011b: 37. SR: Cheiridium museorum (Leach): Tömösváry,

1882: 210–211; Nosek, 1957: 84; Verner, 1971: 27; Superfamily GARYPOIDEA Simon, 1879 Krumpál & Cyprich, 1988: 42; Ducháč, 1994a: 143; Family LARCIDAE Harvey, 1992 Krištofík et al., 1995: 490 (as Cheridium [sic] Genus Larca Chamberlin, 1930 museorum); Šťáhlavský, 1996: 19; Fenďa et al., 1998: Larca lata (Hansen, 1884) 24; Krumpál et al., 2000–2001: 52; Ducháč, 2002c: CR: Larca lata (Hansen): Ducháč, 1993a: 36; 46; Christophoryová, 2008a: 133; Christophoryová Ducháč, 1993b: 66; Ducháč, 1995d: 130; Judson & & Krištofík, 2008: 22; Harvey, 2009: –; Christo- Legg, 1996: 205; Šťáhlavský, 1996: 18; Ducháč, phoryová, 2010: 1, 2, 5; Christophoryová & Krum- 1998b: –; Ducháč, 1999b: 14; Ducháč, 2002c: 35, 39, pál, 2010: 107; Christophoryová & Krumpálová, 68, 82, 152; Blick et al., 2004: –; Drogla & Lippold, 2010: 108; Christophoryová et al., 2011b: 37. 2004: 26; Šťáhlavský & Ducháč, 2005: 83; Christo- phoryová, 2006b: 9; Šťáhlavský, 2006c: 80; Chris- Superfamily CHELIFEROIDEA Risso, 1826 tophoryová et al., 2007b: 19; Harvey & Volschenk, 2007: 368; Petrov & Šťáhlavský, 2007: 18; Christo- Family ATEMNIDAE Chamberlin, 1931 phoryová & Krištofík, 2008: 22; Murienne et al., Subfamily Atemninae Chamberlin, 1931 2008: 174; Harvey, 2009: –; Christophoryová et al., Genus Atemnus Canestrini, 1884 2011b: 37. Atemnus politus (Simon, 1878) SR: Larca lata (Hansen): Christophoryová & SR: Atemnus politus (Simon): Krumpálová & Krištofík, 2008: 22; Christophoryová & Krumpál, Krumpál, 1993: 36; Christophoryová, 2006b: 8, 85; 2009: 14; Christophoryová & Krumpálová, 2010: Šťáhlavský, 2006c: 78; Christophoryová, 2008a: 108; Christophoryová et al., 2011a: 2–4; Christo- 133; Harvey, 2009: –; Christophoryová et al., phoryová et al., 2011b: 37. 2011b: 37.

Superfamily CHEIRIDIOIDEA Hansen, 1894 Subfamily Miratemninae Beier, 1932 Family CHEIRIDIIDAE Hansen, 1894 Genus Diplotemnus Chamberlin, 1933 Subfamily Cheiridiinae Hansen, 1894 Diplotemnus insolitus Chamberlin, 1933 Genus Apocheiridium Chamberlin, 1924 SR: Diplotemnus insolitus Chamberlin: Harvey, Subgenus Apocheiridium (Apocheiridium) 2009: –; Christophoryová et al., 2011b: 37. Chamberlin, 1924 Diplotemnus piger (Simon, 1878): Verner, 1959a: Apocheiridium (Apocheiridium) ferum (Simon, 61–63; Beier, 1963: 248; Verner, 1971: 28 (syn- 1879) onymized by Dashdamirov & Schawaller, 1993: 7). CR: Apocheiridium ferum (Simon): Ducháč, 1997b: –; Diplotemnus ophthalmicus Redikorzev, 1949: Ducháč, 1998b: –; Ducháč, 1999b: 14; Šťáhlavský & Christophoryová, 2006b: 7; Christophoryová, Ducháč, 2001: 48; Ducháč, 2002c: 82, 127, 159–161; 2008a: 133 (synonymized by Dashdamirov & Blick et al., 2004: –; Šťáhlavský, 2006c: 78. Schawaller, 1993: 7). Apocheiridium (Apocheiridium) ferum (Simon): Diplotemnus insolitus insolitus Chamberlin, 1933 Harvey, 2009: –; Christophoryová et al., 2011b: 37. SR: Diplotemnus insolitus insolitus Chamberlin: Harvey, 2009: –. 12 Christophoryová, J. et al.

Family CHELIFERIDAE Risso, 1826 1999b: 14; Ducháč, 2002c: 68, 82, 162; Blick et al., Subfamily Cheliferinae Risso, 1826 2004: –; Harvey, 2009: –. Dactylochelifer latreillii (Leach): Christophory- Genus Chelifer Geoffroy, 1762 ová et al., 2011b: 37. Chelifer cancroides (Linnaeus, 1758) SR: Chelifer degeerii C. L. Koch, 1835: Tömös- CR: Chelifer cancroides (Linnaeus): Stecker, 1874: váry, 1882: 204–205 (as Chelifer De Geeri) (syn- 236; Nosek, 1901: 21; Nosek, 1902: 51, 70; Verner, onymized by Pavesi, 1884: 455). 1957: 88; Studničková, 1968: 23; Verner, 1971: 31; Dactylochelifer latreillei (Leach): Krumpál & Ducháč, 1989: 181; Harvey, 1991: 487–490; Krásný, Cyprich, 1988: 42; Krištofík et al., 1993: 495; 1993: 35; Ducháč, 1994a: 145; Ducháč, 1995d: 130; Ducháč, 1994a: 145; Šťáhlavský, 1996: 33; Fenďa et Sláma, 1995: 544; Šťáhlavský, 1996: 31–32; Ducháč, al., 1998: 24; Krumpál et al., 2000–2001: 52; Ducháč, 1999b: 14; Šťáhlavský, 2000: 4, 23; Šťáhlavský, 2002c: 48; Krištofík et al., 2002: 605 (as Dactiloche- 2001: 88; Ducháč, 2002c: 11, 13, 13–14, 18, 19, 32, lifer [sic] latreillei); Christophoryová, 2006b: 8; 48, 68, 82, 111, 161–162; Krištofík et al., 2003: 232; Christophoryová et al., 2007a: 16; Krumpál Blick et al., 2004: –; Šťáhlavský, 2006c: 78; Šťáh- & Christophoryová, 2007: 97; Christophoryová, lavský, 2007: 60; Šťáhlavský & Krásný, 2007: 432; 2008a: 133; Christophoryová, 2008b: 22; Christo- Harvey, 2009: –; Christophoryová et al., 2011b: 37. phoryová & Krištofík, 2008: 22; Harvey, 2009: –; Chelifer ixoides C. W. Hahn, 1834: Stecker, 1874: Christophoryová, 2009b: 131; Christophoryová, 236; Ducháč, 2002c: 11 (synonymized by Simon, 2010: 1, 2, 5–6; Christophoryová & Krumpál, 2010: 1878: 155). 107–108; Christophoryová & Krumpálová, 2010: Chelifer serratus Stecker, 1974: Stecker, 1874: 108. 235–236; Nosek, 1901: 21; Ducháč, 2002c: 11, 13 Dactylochelifer latreillii (Leach): Christophory- (synonymized by Beier, 1932b: 236). ová et al., 2011b: 37. not Chelifer cancroides (Linnaeus): Verner, 1957: 88 (partly misidentification, partly see Mesochelifer Dactylochelifer latreillii latreillii (Leach, 1817) ressli). CR: Dactylochelifer latreillii latreillii (Leach): Har- SR: Chelifer cancroides (Linnaeus): Tömösváry, vey, 2009: –. 1882: 206–208; Daday, 1888a: 121; Daday, 1888b: SR: Dactylochelifer latreillii latreillii (Leach): 176; Nosek & Lichard, 1962: 35; Krumpál & Cy- Harvey, 2009: –. prich, 1988: 42; Cyprich et al., 1992: 78; Ducháč, Genus Mesochelifer Vachon, 1940 1994a: 145; Šťáhlavský, 1996: 31; Fenďa et al., 1998: Mesochelifer ressli Mahnert, 1981 24; Cyprich et al., 2000: 190; Krumpál et al., 2000– CR: Mesochelifer ressli Mahnert: Ducháč, 1995c: –; 2001: 52; Ducháč, 2002c: 48; Krumpál & Krum- Šťáhlavský, 1996: 33; Ducháč, 1997b: –; Ducháč, pálová, 2003: 123; Varga & Fedor, 2005: 45; Har- 1999b: 14; Ducháč, 2002c: 67, 68, 82, 124, 162–164; vey, 2009: –; Christophoryová, 2010: 1, 2, 5; Chris- Blick et al., 2004: –; Christophoryová, 2006b: 9; tophoryová & Krumpál, 2010: 107; Christophory- Šťáhlavský, 2006a: 172–173; Šťáhlavský, 2006c: 79; ová & Krumpálová, 2010: 108; Christophoryová et Harvey, 2009: –; Šťáhlavský & Tuf, 2009: 100; al., 2011b: 37. Christophoryová et al., 2011b: 37.

Chelifer cancroides cancroides (Linnaeus, 1758) Chelifer cancroides (Linnaeus): Verner, 1957: 88 CR: Chelifer cancroides cancroides (Linnaeus): Har- (partly misidentification). vey, 2009: –. SR: Mesochelifer ressli Mahnert: Krumpálová & SR: Chelifer cancroides cancroides (Linnaeus): Krumpál, 1993: 37; Ducháč, 1995c: –; Ducháč, Harvey, 2009: –. 2002c: 67; Drogla & Lippold, 2004: 29; Christo- phoryová, 2006b: 8; Harvey, 2009: –; Christo- Genus Dactylochelifer Beier, 1932 phoryová, 2010: 2; Christophoryová et al., 2011b: Dactylochelifer latreillii (Leach, 1817) 37. CR: Chelifer latreillii Leach: Nosek, 1902: 54 (as Che- lifer Latreille sub Schäfferi). Genus Rhacochelifer Beier, 1932

Chelifer Schäfferi C. L. Koch, 1839: Stecker, Rhacochelifer peculiaris (L. Koch, 1873) 1874: 237; Nosek, 1901: 21; Ducháč, 2002c: 11, 13 SR (note 15): Chelifer peculiaris L. Koch: Tömös- (synonymized by Simon, 1879: 22). váry, 1882: 201–202; Daday, 1888a: 121–122; Dactylochelifer latreillei (Leach): Verner, 1957: Daday, 1888b: 176–177; Daday, 1918: 2. 88; Šťáhlavský, 1996: 33; Ducháč, 1998b: –; Ducháč, Rhacochelifer peculiaris (L. Koch): Verner, 1960:

Checklist of pseudoscorpions from the Czech Republic and Slovakia 13

168; Verner, 1971: 31; Christophoryová, 2008a: 133; Božanić, 2008: 6; Harvey, 2009: –; Šťáhlavský & Christophoryová et al., 2011b: 37. Tuf, 2009: 100; Christophoryová, 2010: 5; Christo-

Rhacochelifer quadrimaculatus (Tömösváry, 1882) phoryová et al., 2011b: 37. SR: Chelifer quadrimaculatus Tömösváry: Tömös- Chernes cimicoides (Fabricius): Verner, 1957: 88 váry, 1882: 196–198; Daday, 1918: 2. (misidentification). Rhacochelifer quadrimaculatus (Tömösváry): SR: Chernes hahnii (C. L. Koch): Krumpál & Verner, 1960: 168; Verner, 1971: 31; Harvey, 1991: Cyprich, 1988: 42 (as Chernes hahni); Ducháč, 530; Christophoryová, 2006b: 7; Christophoryová, 1994a: 144 (as Chernes hahni); Krištofík et al., 1996: 2008a: 133; Harvey, 2009: –; Christophoryová, 560 (as Chernes hahni); Šťáhlavský, 1996: 26 (as 2010: 2; Christophoryová et al., 2011b: 37. Chernes hahni); Krumpál et al., 2000–2001: 52 (as Chernes hahni); Ducháč, 2002c: 47 (as Chernes Family CHERNETIDAE Menge, 1855 hahni); Krumpál & Christophoryová, 2007: 96; Christophoryová & Mock, 2009: 17; Harvey, 2009: Subfamily Chernetinae Menge, 1855 –; Christophoryová, 2010: 1, 6; Christophoryová Tribe Chernetini Menge, 1855 & Krumpál, 2010: 108; Christophoryová & Krum- Genus Chernes Menge, 1855 pálová, 2010: 108; Christophoryová et al., 2011b:

Chernes cimicoides (Fabricius, 1793) 37. CR: Chelifer cimicoides (Fabricius): Nosek, 1901: 21; Chernes similis (Beier): Christophoryová et al., Nosek, 1902: 51, 70; Ducháč, 2002c: 13, 14. 2007a: 16; Christophoryová & Krištofík, 2008: 22 (all misidentification). Chernes cimicoides (Fabricius): Stecker, 1874: 234; Verner, 1971: 30; Šťáhlavský, 1996: 25; Chernes nigrimanus Ellingsen, 1897 Ducháč, 1999b: 14; Ducháč, 2002c: 11, 18, 68, 82, CR: Chernes nigrimanus Ellingsen: Ducháč, 1995c: – 186; Krištofík et al., 2003: 232; Blick et al., 2004: –; ; Schmarda, 1995: 61–65; Šťáhlavský, 1996: 26; Šťáhlavský, 2006b: 163; Harvey, 2009: –; Christo- Ducháč, 1998b: –; Ducháč, 1999b: 14; Šťáhlavský, phoryová et al., 2011b: 37. 2001: 73; Ducháč, 2002c: 19, 67, 68, 82, 188; Blick et Chernes Mengei L. Koch, 1873: Stecker, 1874: al., 2004: –; Drogla & Lippold, 2004: 33–34; Šťáh- 234; Ducháč, 2002c: 11 (synonymized by Beier, lavský & Ducháč, 2005: 83; Šťáhlavský, 2006c: 79; 1932b: 155). Harvey, 2009: –; Christophoryová et al., 2011b: 37.

Chelifer Mengei L. Koch: Nosek, 1901: 21; Chernes similis (Beier, 1932) Ducháč, 2002c: 13. CR: Chernes similis (Beier): Šťáhlavský, 1996: 27; not Chernes cimicoides (Fabricius): Verner, 1957: Ducháč, 1997b: –; Ducháč, 1998b: –; Ducháč, 1999b: 88 (misidentification, see Chernes hahnii). 14; Šťáhlavský, 2000: 13; Ducháč, 2002c: 18, 20, 82, SR: Chernes cimicoides (Fabricius): Tömösváry, 127, 188; Blick et al., 2004: –; Šťáhlavský, 2006a: 1882: 188–190; Daday, 1888a: 117–118; Daday, 173–174; Šťáhlavský, 2006c: 79; Harvey, 2009: –; 1888b: 172; Verner, 1960: 168; Verner, 1971: 30; Christophoryová et al., 2011b: 37. Krumpál & Cyprich, 1988: 42; Ducháč, 1994a: 144 SR: Toxochernes similis (Beier): Krumpál, 1980: (as Chernes cimocoides [sic]); Šťáhlavský, 1996: 25; 24; Krumpál et al., 2000–2001: 52. Krumpál et al., 2000–2001: 52; Ducháč, 2002c: 47 Chernes similis (Beier): Krumpál & Cyprich, (as Chernes cimocoides [sic]); Krištofík et al., 2002: 1988: 42; Krumpálová & Krumpál, 1993: 37; Šťáh- 605; Christophoryová, 2006b: 8; Christophoryová, lavský, 1996: 27; Ducháč, 1997b: –; Ducháč, 2002c: 2009b: 131; Harvey, 2009: –; Christophoryová, 127; Christophoryová & Krumpál, 2005: 79; Chris- 2010: 2; Christophoryová & Krumpál, 2010: 108; tophoryová, 2006a: 63; Christophoryová, 2006b: 8, Christophoryová et al., 2011b: 37. 24; Christophoryová & Krumpál, 2006b: 4; Chris-

Chernes hahnii (C. L. Koch, 1839) tophoryová & Krumpál, 2007b: 29; Christophory- CR: Chernes hahnii (C. L. Koch): Ducháč, 1994a: 144 ová et al., 2007b: 20; Harvey, 2009: –; Christo- (as Chernes hahni); Šťáhlavský, 1996: 26 (as Chernes phoryová et al., 2011b: 37. hahni); Ducháč, 1999b: 14 (as Chernes hahni); Šťáh- not Chernes similis (Beier): Christophoryová et lavský, 2000: 25; Šťáhlavský, 2001: 90–91; Ducháč, al., 2007a: 16; Christophoryová & Krištofík, 2008: 2002c: 20, 47, 68, 82 (all as Chernes hahni), 187; Blick 22 (all misidentification, see Chernes hahnii). et al., 2004: –; Vítková et al., 2005: 147; Šťáhlavský, Chernes vicinus (Beier, 1932) 2006a: 173; Šťáhlavský, 2006b: 163; Šťáhlavský, CR: Chernes vicinus (Beier): Šťáhlavský & Ducháč, 2006c: 79; Šťáhlavský & Krásný, 2007: 432–433; 2001: 49; Ducháč, 2002c: 189; Blick et al., 2004: –; 14 Christophoryová, J. et al.

Drogla & Lippold, 2004: 34–35; Šťáhlavský & 1907: 159). Ducháč, 2005: 83; Christophoryová, 2006b: 9; Har- Toxochernes panzeri (C. L. Koch): Krumpál et vey, 2009: –; Christophoryová et al., 2011b: 37. al., 2000–2001: 52. SR: Allochernes vicinus Beier: Krumpál & Cy- Dinocheirus panzeri (C. L. Koch): Krumpál & prich, 1988: 42; Krumpál et al., 2000–2001: 52. Cyprich, 1988: 42; Šťáhlavský, 1996: 30; Krištofík et Chernes vicinus (Beier): Drogla & Lippold, al., 2002: 605; Krumpál & Krumpálová, 2003: 123; 2004: 34–35; Harvey, 2009: –; Christophoryová et Christophoryová & Mock, 2009: 17; Harvey, 2009: al., 2011b: 37. –; Christophoryová, 2010: 2, 6; Christophoryová et al., 2011b: 37. Genus Dendrochernes Beier, 1932 Pselaphochernes scorpioides (Hermann): Dendrochernes cyrneus (L. Koch, 1873) Krištofík et al., 2009: 975 (misidentification). CR: Dendrochernes cyrneus (L. Koch): Ducháč, 1993a: 36; Ducháč, 1995a: 52; Šťáhlavský, 1996: 23; Subfamily Lamprochernetinae Beier, 1932

Ducháč, 1998b: –; Ducháč, 1999b: 14; Ducháč, Genus Allochernes Beier, 1932 2002c: 39, 65, 68, 82, 189–190; Blick et al., 2004: –; Allochernes peregrinus Lohmander, 1939 Šťáhlavský & Ducháč, 2005: 83; Christophoryová, CR: Allochernes peregrinus Lohmander: Ducháč, 2006b: 9; Šťáhlavský, 2006c: 79; Harvey, 2009: –; 1995a: 52; Ducháč, 1995c: –; Šťáhlavský, 1996: 29; Christophoryová et al., 2011b: 37. Ducháč, 1998a: 508; Ducháč, 1998b: –; Ducháč, SR: Chernes cyrneus L. Koch: Tömösváry, 1882: 1999b: 14; Ducháč, 2002c: 65, 67, 68, 82, 164–167; 194–195; Daday, 1918: 1. Blick et al., 2004: –; Šťáhlavský & Ducháč, 2005: 83; Chernes (Chernes) cyrneus hungaricus Daday, Christophoryová, 2006b: 9, 84; Harvey, 2009: –; 1888a: 116–117; Daday, 1888b: 171–172 (syn- Christophoryová et al., 2011b: 37. onymized by Beier, 1929: 346). SR: Allochernes peregrinus Lohmander: Krum- Dendrochernes cyrneus (L. Koch): Krumpál & pálová & Krumpál, 1993: 37; Ducháč, 1995a: 51; Cyprich, 1987: 196; Krumpál & Cyprich, 1988: 42; Ducháč, 1995c: –; Ducháč, 2002c: 64, 67; Drogla & Ducháč, 1993a: 36; Šťáhlavský, 1996: 23; Fenďa et Lippold, 2004: 29–30; Christophoryová, 2006b: 8, al., 1998: 24; Krumpál et al., 2000–2001: 52; Ducháč, 24, 84; Christophoryová & Krumpál, 2006b: 4; 2002c: 39; Christophoryová, 2006b: 8; Christo- Christophoryová & Krumpál, 2007b: 29; Harvey, phoryová et al., 2007a: 16; Harvey, 2009: –; Chris- 2009: –; Christophoryová & Krumpál, 2010: 109; tophoryová, 2010: 1, 2; Christophoryová & Krum- Christophoryová et al., 2011b: 37. pál, 2010: 105, 109; Christophoryová & Krum- Allochernes wideri (C. L. Koch, 1843) pálová, 2010: 108; Christophoryová et al., 2011b: CR: Chelifer Wideri C. L. Koch: Nosek, 1901: 21; 37. Ducháč, 2002c: 13. Dendrochernes cyrneus cyrneus (L. Koch, 1873) Chernes Wideri (C. L. Koch): Stecker, 1874: 233; CR: Dendrochernes cyrneus cyrneus (L. Koch): Har- Ducháč, 2002c: 11. vey, 2009: –. Chelifer (Chernes) Wideri (C. L. Koch): Nosek, SR: Dendrochernes cyrneus cyrneus (L. Koch): 1902: 59. Harvey, 2009: –. Allochernes wideri (C. L. Koch): Verner, 1959b:

379; Ducháč, 1993b: 66; Šťáhlavský, 1996: 28; Šťáh- Genus Dinocheirus Chamberlin, 1929 lavský, 1998: –; Ducháč, 1999b: 14; Šťáhlavský, Dinocheirus panzeri (C. L. Koch, 1837) 2000: 24; Šťáhlavský, 2001: 88–89; Ducháč, 2002c: CR: Dinocheirus panzeri (C. L. Koch): Ducháč, 16, 18, 35, 68, 82, 185; Christophoryová, 2006b: 85; 1993b: 66; Šťáhlavský, 1996: 30; Šťáhlavský, 1998: – Šťáhlavský, 2006a: 173; Šťáhlavský, 2006b: 163; ; Ducháč, 1999b: 14; Šťáhlavský, 2000: 25–26; Šťáh- Christophoryová et al., 2007b: 19; Šťáhlavský & lavský, 2001: 91–92; Ducháč, 2002c: 35, 68, 82, 190; Krásný, 2007: 432; Harvey, 2009: –; Šťáhlavský & Krištofík et al., 2003: 232; Blick et al., 2004: –; Šťáh- Tuf, 2009: 100; Christophoryová, 2010: 6–7, 8; lavský, 2006a: 174; Šťáhlavský, 2006b: 163; Šťáh- Christophoryová et al., 2011b: 37. lavský, 2006c: 79; Christophoryová et al., 2007b: SR: Allochernes wideri (C. L. Koch): Krumpál & 19; Šťáhlavský & Krásný, 2007: 433; Harvey, 2009: Cyprich, 1988: 42; Šťáhlavský, 1996: 28; Christo- –; Šťáhlavský & Tuf, 2009: 100; Christophoryová, phoryová et al., 2007a: 16; Krumpál & Christo- 2010: 8; Christophoryová et al., 2011b: 37. phoryová, 2007: 96; Christophoryová & Krištofík, SR: Chernes rufeolus (Simon, 1879): Tömösváry, 2008: 22; Christophoryová & Mock, 2009: 17; Har- 1882: 190–191 (synonymized by Ellingsen,

Checklist of pseudoscorpions from the Czech Republic and Slovakia 15 vey, 2009: –; Christophoryová, 2010: 6–7; Christo- Ducháč, 1995d: 130; Šťáhlavský, 1996: 21–22; phoryová & Krumpál, 2010: 109; Christophoryová Ducháč, 1999b: 14; Šťáhlavský, 2000: 27; Šťáh- & Krumpálová, 2010: 108; Christophoryová et al., lavský, 2001: 93–94; Ducháč, 2002c: 17, 19, 68, 82, 2011b: 37. 192; Blick et al., 2004: –; Harvey, 2009: –; Christo-

Allochernes wideri phaleratus (Simon, 1879) phoryová et al., 2011b: 37. SR (note 16): Chernes phaleratus (Simon): Daday, SR: Lamprochernes nodosus (Schrank): Krumpál 1888a: 118; Daday, 1888b: 173; Daday, 1918: 2. & Cyprich, 1988: 42; Mašán & Krištofík, 1992: 91; Allochernes wideri phaleratus (Simon): Christo- Krištofík et al., 1995: 490 (as Laprochernes [sic] nodo- phoryová, 2008a: 133. sus); Krištofík et al., 1996: 560 (as Laprochernes [sic] nodosus); Šťáhlavský, 1996: 21; Krumpál et al., Allochernes wideri wideri (C. L. Koch, 1843) 2000–2001: 52; Christophoryová et al., 2007a: 16; CR: Allochernes wideri wideri (C. L. Koch): Blick et Krištofík et al., 2007: 750; Christophoryová, 2008a: al., 2004: –; Harvey, 2009: –. 133; Harvey, 2009: –; Christophoryová, 2010: 1, 2; SR: Allochernes wideri wideri (C. L. Koch): Har- Christophoryová & Krumpál, 2010: 110; Christo- vey, 2009: –. phoryová & Krumpálová, 2010: 108; Christo- Genus Anthrenochernes Lohmander, 1939 phoryová et al., 2011b: 37. Anthrenochernes stellae Lohmander, 1939 not Lamprochernes nodosus (Schrank): Krumpál CR: Anthrenochernes stellae Lohmander: Ducháč, & Christophoryová, 2007: 96; Christophoryová, 1998b: –; Šťáhlavský, 1998: –; Šťáhlavský, 2000: 25; 2010: 7 (all misidentification, see Lamprochernes Šťáhlavský, 2001: 89–90; Šťáhlavský & Ducháč, chyzeri).

2001: 48; Ducháč, 2002c: 20, 185–186; Blick et al., Lamprochernes nodosus nodosus (Schrank, 1803) 2004: –; Drogla & Lippold, 2004: 31; Šťáhlavský & CR: Lamprochernes nodosus nodosus (Schrank): Har- Ducháč, 2005: 83; Christophoryová, 2006b: 9; Har- vey, 2009: –. vey, 2009: –; Christophoryová, 2010: 8; Christo- SR: Lamprochernes nodosus nodosus (Schrank): phoryová et al., 2011b: 37. Harvey, 2009: –.

Genus Lamprochernes Tömösváry, 1882 Genus Lasiochernes Beier, 1932 Lamprochernes chyzeri (Tömösváry, 1882) Lasiochernes pilosus (Ellingsen, 1910) CR: Lamprochernes chyzeri (Tömösváry): Šťáh- SR: Lasiochernes pilosus (Ellingsen): Krumpálová & lavský, 1996: 22; Ducháč, 1999b: 14; Šťáhlavský, Krumpál, 1993: 36; Drogla & Lippold, 2004: 40; 2000: 27; Šťáhlavský, 2001: 92–93; Ducháč, 2002c: Christophoryová, 2006b: 8; Harvey, 2009: –; Chris- 68, 82, 190–192; Blick et al., 2004: –; Harvey, 2009: – tophoryová, 2010: 8; Christophoryová & Krumpál, ; Christophoryová et al., 2011b: 37. 2010: 110; Christophoryová et al., 2011b: 37. SR: Chernes Chyzeri Tömösváry: Tömösváry, 1882: 186–187; Daday, 1888a: 115–116; Daday, Genus Pselaphochernes Beier, 1932

1888b: 170; Daday, 1918: 1. Pselaphochernes dubius (O. P.–Cambridge, 1892) Lamprochernes chyzeri (Tömösváry): Krumpál SR (note 17): Pselaphochernes dubius (O. P.– & Cyprich, 1988: 42; Šťáhlavský, 1996: 22; Krum- Cambridge): Krumpál & Krumpálová, 2003: 123; pál et al., 1997: 105; Christophoryová, 2008a: 133; Christophoryová, 2008a: 133. Christophoryová, 2009b: 131; Harvey, 2009: –; Pselaphochernes scorpioides (Hermann, 1804) Christophoryová & Krumpálová, 2010: 108; Chris- CR: Pselaphochernes scorpioides (Hermann): Ducháč, tophoryová et al., 2011b: 37. 1993b: 66; Ducháč, 1994a: 144–145; Šťáhlavský, Lamprochernes nodosus (Schrank): Krumpál & 1996: 24; Šťáhlavský, 1998: –; Ducháč, 1999b: 14; Christophoryová, 2007: 96; Christophoryová, 2010: Šťáhlavský, 2000: 27–28; Šťáhlavský, 2001: 94; 7 (all misidentification). Ducháč, 2002c: 35, 47–48, 68, 82, 193; Blick et al.,

Lamprochernes nodosus (Schrank, 1803) 2004: –; Christophoryová, 2006b: 83, 85; Šťáh- CR: Chernes bohemicus Stecker, 1874: Stecker, 1874: lavský, 2006a: 174; Šťáhlavský, 2006b: 163; Šťáh- 233; Stecker, 1875: 172; Ducháč, 2002c: 11, 13 (syn- lavský, 2006c: 79; Šťáhlavský & Krásný, 2007: 433; onymized by Beier, 1932b: 83). Božanić, 2008: 6; Harvey, 2009: –; Šťáhlavský & Chelifer bohemicus Stecker: Simon, 1879: 42; Tuf, 2009: 100; Christophoryová, 2010: 7, 8; Chris- Nosek, 1901: 21; Ducháč, 2002c: 13. tophoryová et al., 2011b: 37. Lamprochernes nodosus (Schrank): Minář, 1966: SR: Pselaphochernes scorpioides (Hermann): 271; Harvey, 1991: 589–591; Krásný, 1993: 41; Šťáhlavský, 1996: 24; Christophoryová 16 Christophoryová, J. et al.

& Krumpál, 2005: 79; Christophoryová, 2006a: 63; 1996c). Christophoryová, 2006b: 24; Christophoryová & 8. The attribution of Neobisium spelaeum is Krumpál, 2006a: 43; Christophoryová & Krumpál, based on Stecker´s record (1875) from the sur- 2006b: 4; Christophoryová & Krumpál, 2007b: 29; roundings of Adamov in Moravia (Czech Repub- Christophoryová et al., 2007a: 16; Christophoryová lic) but with no extant specimen available, most et al., 2007b: 20; Krumpál & Christophoryová, likely a misidentification. 2007: 96; Christophoryová, 2008a: 133; Christo- 9. The attribution of Neobisium jugorum in the phoryová & Krištofík, 2008: 22; Christophoryová Czech Republic is based on Stecker (1874) and his & Krumpál, 2009: 15; Harvey, 2009: –; Christo- later citations (Simon 1879, Caporiacco 1926), but phoryová, 2010: 7; Christophoryová & Krumpál, with no extant specimen available. The single re- 2010: 110; Christophoryová & Krumpálová, 2010: cord from Slovakia by Verner (1960) should be re- 108; Christophoryová et al., 2011b: 37. vised as well. not Pselaphochernes scorpioides (Hermann): 10. Neobisium macrodactylum: its attribution in Krištofík et al., 2009: 975 (misidentification, see Slovakia is based on a single record by Krumpál Dinocheirus panzeri). (1980) but with no extant specimen available. 11. Neobisium simile was observed more than a Notes and discussion on the checklist hundred years ago (Tömösváry 1882) but has not 1. The record of Chthonius carinthiacus in Slovakia been recorded since in Slovakia. This record is (Šťáhlavský 1996) is, however, without exact most likely a misidentification. faunistic data and no extant specimen available. 12. Neobisium validum: its Slovakian status is 2. Chthonius diophthalmus has not yet been offi- not accepted, no extant specimen from the older cially approved for the fauna of the Czech Repub- records (Daday 1888a, 1888b, 1918) is available. lic and its attribution declared by Harvey (1991, 13. The attribution of Roncus alpinus in the 2009) “near Brno" (type locality for synonymic C. Czech Republic is based on Stecker (1874) and his ksenemanni) appears a misinterpretation of the Pop later citations (Simon 1879, Caporiacco 1922, Ivan conservation area, situated in Ukraine (Hadži Cârdeiu 1948), with no extant specimen available. 1939). Its Slovakian status is not accepted. Excepting 3. Attribution of Chthonius ressli has been some older records (probably misidentified) from based on a single record in the Czech Republic eastern Slovakia (Tömösváry 1882, Daday 1918) (Šťáhlavský 2006a) as well as in Slovakia (Mock et with no material available, the species has not al. 2004, 2005). been recorded (Krumpál & Krumpálová 2003), 4. The Czech status of Chthonius submontanus even after detailed morphological and mor- is not accepted. It is a misidentification (Ducháč phometric analyses on Roncus species from the 2002c) of C. cf. pygmaeus carinthiacus (in the check- same region (Christophoryová & Šťáhlavský list C. carinthiacus). 2007). Moreover, the distribution area of this spe- 5. The attribution of Chthonius subterraneus is cies seems to be limited to the European Alps based on 2 specimens from Slovakia (Krumpál & (Harvey 2009). Krumpálová 2003), however no extant specimen is 14. Roncus transsilvanicus has not yet been offi- available. cially approved for the fauna of the Czech Repub- 6. The attribution of Chthonius tenuis in the lic, and its attribution declared by Harvey (1991, Czech Republic, based on some older references 2009) “near Brno“ appears a misinterpretation of (e.g. Stecker 1874, Nosek 1901, Ducháč 1999b), ap- the Pop Ivan conservation area, situated in pears disputable as no extant specimen is available Ukraine (Hadži 1939). Its preliminary attribution for identification. The recent papers (Šťáhlavský is based on the recent record by Krumpál & 2000, 2001, 2006a, 2006c, Šťáhlavský & Král 2004) Krumpálová (2003) from eastern Slovakia but with emphasize misidentification of C. carinthiacus no extant specimen available. The detailed (Gardini pers. comm. 2009). However, the species karyological analyses from this region (Christo- may potentially be found in the Czech Republic. phoryová & Šťáhlavský 2007) indicate a new spe- 7. The attribution of Chthonius cf. kewi in the cies. However, until completed, R. transsilvanicus Czech Republic is based on one male with a single will be accepted for Slovakia. posteromarginal microseta on carapax, but with 15. Rhacochelifer peculiaris was observed more chaetotaxy and morphometry similar to C. tetra- than a hundred years ago (Tömösváry 1882, chelatus, common in the same locality (Ducháč Daday 1888a, 1888b, 1918) but has not been re-

Checklist of pseudoscorpions from the Czech Republic and Slovakia 17 corded since. However, the species may poten- hungaricus, Neobisium slovacum, N. beieri). Knowl- tially be found in Slovakia. edge on the distribution of some other species is 16. Allochernes wideri phaleratus was observed often still very poor. more than a hundred years ago (Tömösváry 1882) but has not been recorded since in Slovakia. How- ever, its presence in either the Czech Republic or Acknowledgements. We are grateful to our colleagues Slovakia is possible. and friends for their help, criticism and comments, 17. The only specimen of Pselaphochernes including Christoph Hörweg (Austria), Eliška Ducháčová, dubius found in Slovakia (Krumpál & Krumpálová the late Václav Ducháč, Giulio Gardini (Italy), Mark S. 2003) has not been available for more detailed re- Harvey (Australia), and Juan A. Zaragoza (Spain). The project was financially supported by VEGA 1/0176/09, vision and is likely to be misidentified. The species VEGA 1/0137/11 and MŠMT 0021620828. is known from Spain to Scandinavia (Harvey 2009) and even from Central Europe (Drogla & Lippold 2004), however from none of the neighbouring References countries. Beier, M. (1928): Die Pseudoskorpione des Wiener Recent research on pseudoscorpions has detected Naturhistorischen Museums. I. Hemictenodactyli. Annalen de the existence of some cryptic species in Central Naturhistorischen Museum in Wien 42: 285-314. Beier, M. (1929): Die Pseudoskorpione des Wiener Europe; hence supplementary analyses (e.g. DNA, Naturhistorischen Museums. II. Panctenodactyli. Annalen de cytogenetics) are required in future. For some dis- Naturhistorischen Museum in Wien 43: 341-367. putable taxa (especially Neobisium carcinoides and Beier, M. (1932a): Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich 57: i-xx: 1-258. Roncus lubricus) a broad species concept was ap- Beier, M. (1932b): Pseudoscorpionidea II. Subord. C. Cheliferinea. plied as more useful, because the status of these Tierreich 58: i-xxi: 1-294. species with large distribution area may be signifi- Beier, M. (1963): Ordnung Pseudoscorpionidea (Afterskorpione). Bestimmungsbücher zur Bodenfauna Europas. Lieferung 1. cantly changed soon (according to our preliminary Akademie–Verlag, Berlin. results in molecular analyses). Benová, A., Svatoň, J. (2009): Poznatky o faune pavúkov (Araneae) Altogether the pseudoscorpion fauna of Slo- Národného parku Nízke Tatry. Príroda Nízkych Tatier (Banská Bystrica) 2: 153-158. vakia and the Czech Republic contains 58 species Blick, T., Muster, C., Ducháč, V. (2004): Checkliste der in eight families. Most of them, especially Chtho- Pseudoskorpione Mitteleuropas. Checklist of the pseudo- nius ischnocheles, C. orthodactylus, C. tenuis, C. tetra- scorpions of Central Europe (Arachnida: Pseudoscorpiones), Version 1. Available: http://www.arages.de/checklist.html chelatus, Microbisium suecicum, Neobisium carci- accessed at 2011.03. 01. noides, N. erythrodactylum, N. fuscimanum, N. sylva- Božanić, B. (2008): Mosses as living environment for inveretebrates. ticum, Larca lata, Apocheiridium ferum, Cheiridium Bachelor thesis. Department of Ecology and Environmental Sciences, Faculty of Science, Palacký University, Olomouc. museorum, Chelifer cancroides, Dactylochelifer latreil- Caporiacco, L. di (1922): Saggio sulla fauna aracnologica della lii, Mesochelifer ressli, Allochernes peregrinus, A. Carnia e regioni limitrofe. Memoire della Società Entomologica wideri, Chernes cimicoides, C. hahnii, Dendrochernes Italiana 1: 60-111. Caporiacco, L. di (1926): Secondo saggio sulla fauna aracnologica cyrneus, Dinocheirus panzeri, Lamprochernes chyzeri, della Carnia e regioni limitrofe. Memorie della Società L. nodosus, and Pselaphochernes scorpioides have Entomologica Italiana 5: 70-130. broad distributions across Europe and are known Cârdeiu, F. (1948): Beiträge zur Fauna der Pseudoscorpioniden aus dem Urwald «Slătiora» Bezirk Cîmpulung (Bucovina). Revista from almost all surrounding countries (Harvey Ştiinţifică „V. ADAMACHI“ 34: 1-4. 2009). On the other hand, due to its specific loca- Chamberlin, J.C. (1929): A synoptic classification of the false tion in Central Europe, this region is marginal for scorpions or chela–spinners, with a report on a cosmopolitan collection of the same. Part 1. The Heterosphyronida the distribution area of several species, such as (Chthoniidae) (Arachnida–Chelonethida). Annals and Magazine Chthonius boldorii (most eastern occurrence); Neo- of Natural History (10) 4: 50-80. bisium crassifemoratum, Chernes similis and many Christophoryová, J. (2006a): Spoločenstvá epigeických šťúrikov (rad: Pseudoscorpiones) dubovo-hrabového vegetačného stupňa Carpathians species mentioned below (most west- Malých Karpát a Trnavskej pahorkatiny. pp. 62-64. In: Sirotiak, ern occurrences); Atemnus politus (most northern M., Littera, P. (eds.), Študentská vedecká konferencia. Kartprint, occurrence) and Anthrenochernes stellae (most Bratislava. Christophoryová, J. (2006b): Šťúriky (Pseudoscorpiones) Malých southern occurrence). The diversity of pseudo- Karpát a Trnavskej pahorkatiny. Diploma thesis. Department of scorpions in Slovakia includes several Carpathian Zoology, Faculty of Natural Sciences, Comenius University, elements (Chthonius heterodactylus, Mundochthonius Bratislava. Christophoryová, J. (2008a): Druhové zloženie a rozšírenie šťúrikov carpaticus, Neobisium brevidigitatum, N. carpaticum, (Pseudoscorpiones) Slovenska. pp. 132-134. In: Littera, P., and N. polonicum) and local endemics (Chthonius Budzáková, M. (eds.), Študentská vedecká konferencia. Iris, Bratislava. 18 Christophoryová, J. et al.

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(2004): Príspevok k fenológii Východná, 13–16 September 2007, Ústav zoológie SAV, druhu Neobisium muscorum (Pseudoscorpiones) v južnej časti Bratislava. Malých Karpát. pp. 10-11. In: Krumpálová, Z. (ed.), Východná, Cyprich, D., Fenďa, P., Krumpál, M., Lukáš, J., Bystrík, A. (2000): 9–12 September 2004. Ústav zoológie SAV, Bratislava. Hniezdna fauna zebričky červenozobej (Taeniopygia guttata Christophoryová, J., Krumpál, M. (2005): Communities of castanotis). Tichodroma 13: 189-200. pseudoscorpions (Pseudoscorpiones, Arachnida) in epigeon of Cyprich, D., Krumpál, M., Nagy, R. (1992): Prieskum fauny hniezd oak-hornbeam forests in the Malé Karpaty Mts and Trnavská Columba livia f. domestica L. na juhozápadnom Slovensku. pahorkatina hills (SW Slovakia). Ekológia (Bratislava) 24(2): 76- Entomologické Problémy 22: 75-82. 86. Daday, E. (1888a): A Magyar Nemzeti Muzeum álskorpióinak Christophoryová, J., Krumpál, M. (2006a): Epigeické šťúriky áttekintése. Természetrajzi Füzetek 11: 111-136. 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(2004): Zur Kenntnis der Pseudoskorpion– 20. In: Stloukal, E. (ed.), 12. Feriancove dni, 23–24 November Fauna von Ostdeutschland (Arachnida, Pseudoscorpiones). 2006. Faunima, Bratislava. Arachnologische Mitteilungen 27/28: 1-54. Christophoryová, J., Krumpál, M. (2007a): Species composition, Ducháč, V. (1988): Příspěvek k poznání štírků Českého středohoří. distribution and taxonomic problems of soil pseudoscorpions of Fauna Bohemiae Septentrionalis 13: 103-108. subgenus Ephippiochthonius in Slovakia (Pseudoscorpiones, Ducháč, V. (1989): Příspěvek k faunistice štírků Československa Chthoniidae, genus Chthonius). pp. 14. In: Tajovský, K. (ed.), (Pseudoscorpionidea). Zborník Slovenského Národného Múzea 9th Central European Workshop on Soil Zoology, 17–20 April – Prírodné Vedy 35: 179-182. 2007. Institute of Soil Biology, Biology Centre, Academy of Ducháč, V. (1993a): Zwei neue Afterskorpion–Arten aus der Sciences of the Czech Republic, České Budějovice. Tschechischen Republik. 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