Cambrian Geology and Paleontology

Total Page:16

File Type:pdf, Size:1020Kb

Cambrian Geology and Paleontology SMITHSONIAN MISCELLANEOUS COLLECTIONS PART OF VOLUME LIII CAMBRIAN GEOLOGY AND PALEONTOLOGY No. 3.—CAMBRIAN BRACHIOPODA: DESCRIPTIONS OF NEW GENERA AND SPECIES With Four Plates BY CHARLES D. WALCOTT MWi No. I8I0 CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION October I, 1908 : CAMBRIAN GEOLOGY AND PALEONTOLOGY No. 3.—CA^IBRIAN BRACHIOPODA: DESCRIPTIONS OF NEW GENERA AND SPECIES By CHARLES D. WALCOTT (With Four Plates) This is the eighth paper resuhing from the prehminary studies for Monograph 51 of the U. S. Geological Survey. I expect to use many new generic and specific names in lists of fossils occurring in geo- logic sections and in a forthcoming paper on the classification of the Brachiopoda, and think it is best to describe the fossils before using their names elsewhere. The paper on the classification will be the last of the preliminary papers, as the monograph is now in the editor's hands and should appear in 1909. The previous papers in this series are I. Note on the genus Lingulcpis. American Jour. Sci., 4th ser., Ill, 1897, pp. 404-405. II. Cambrian Brachiopoda : Genera Iphidia and Yorkia, with descriptions of new species of each, and of the genus AcrotJiclc. Proc. U. S. Na- tional Museum, XIX, 1897, pp. 707-718. III. Note on the brachiopod fauna of the quartzitic pebbles of the Car- boniferous conglomerates of the Narragansett Basin, Rhode Island. American Jour. Sci.. 4th ser., VI, 1898, pp. 327-328. IV. Cambrian Brachiopoda : Obolus and Lingtdella, with descriptions of new species. Proc. U. S. National Museum, XXI, 1898, pp. 385-420. V. Cambrian Brachiopoda: Obolella, subgenus Glyptias; Bicia; Obolus, subgenus IVcsfoiiia; with descriptions of new species. Proc. U. S. National Museum, XXIII, 1901, pp. 669-695. VI. Cambrian Brachiopoda: Acrotrcta ; Linnarsson-cUa ; Obolus; with de- scriptions of new species. Proc, U. S. National Museum, XXV, 1902, PP- 577-612. VII. Cambrian Brachiopoda, with descriptions of new genera and species. Proc. U. S. National Museum, XXVIII, 1905, pp. 227-2,^7. There are also a number of Cambrian brachiopoda described in two papers on the Cambrian faunas of China : Caml)rian Faunas of China. Proc. U. S. National Museum, XXIX, 1905, pp. 1-106. Cambrian Faunas of Cliina. Proc. U. S. National Museum, XXX, 1906, PP- 563-595- 53 54 SMITHSONIAN MISCELIvANEOUS COLLECTIONS VOL. 53 Genus MICKWITZIA Schmidt [1888, p. 24] MICKWITZIA OCCIDENS, new. species Plate 7, Figure i There are only crushed and broken specimens of this shell. One of these shows that the apex of the ventral valve was a little above the posterior margin of the shell, very much as in Mickwitzia pre- tiosa. The outline of the valves appears to have been ovate to sub- circular, with the ventral valve moderately convex. The shell is phosphatic or chitinous and built up of three principal layers. The outer layer is thin and thickly set with minute pustules or granules that give the surface a roughened appearance. When the outer layer is exfoliated, which is usually the case, the middle layer presents a smooth, shining surface that is marked by a few concentric striae and numerous fine radiating striae, between which many very minute punctae occur. The inner layer shows minute, irregular, serpentine, rounded ridges, perforated by vertical canals or punctae. An inte- rior of a ventral valve shows the lines of adyance of the antero- lateral muscle scars. The largest shell indicated on the surface of the siliceous shale has a length and width of 12 mm. This species and the generic reference is based on the character of the apex of the ventral valve and the structure and character of the shell. Formation and Locality.—Lower Cambrian : ( i ) Near the base of the section, about 5,500 feet (1,676.4 m.) below the top of the Lower Cambrian, in shaly indurated sandstones, one mile (1.61 km.) east of the Saline Valley road, and 2 to 3 miles (3.22 to 4.83 km.) east-northeast of Waucoba Springs, Inyo County, California. (2) Sandstones on small hill in the salt flat one mile {1.61 km.) northeast of Silver Peak Mill, Silver Peak quadrangle ( U. S. G.S.), Esmeralda County, Nevada} MICKWITZIA PRETIOSA, new species Plate 7, Figure 2 This species is founded on a single specimen of a ventral valve. It has a length of 7 mm. ; width, 6.5 mm. Outline subcircular, slightly convex; apex curved over toward the posterior margin and projecting beyond it. False area short and obscure. Surface marked by radiating, raised lines, that at the front margin show six ' Where there is more than one locality, the one from which the type speci- mens come is italicised. CAMBRIAN BRACHIOPODA—WALCOTT 55 in a distance of two millimeters. Fine papillae are thickly scattered over the surface. They have a tendency to follow concentric lines of growth on some portions of the shell, and on others they appear on low, narrow, serpentine ridges, as in Mickwitsia monilifera (Lin- narsson) [1869, p. 344]. A few large pimctse are scattered here and there over the surface. Inner surfaces and layers of shell unknown. This beautiful shell differs in the details of its surface from M. monilifera; it is also less convex and the apex is nearer the posterior margin. Formation and Locality.—Lower Cambrian : Eophyton sand- stone, at Lugnas, Vestergotland, Sweden. Genus MICROMITRA Meek [1873, p. 479] MICROMITRA HAYDENI, new species Plate 7, Figures 3 and 2i<^ Ventral valve subconical, with a minute beak arching slightly over a strong, arched pseudodeltidium, which is about one-half as long as the height of the valve. Cardinal slope rounded ; a slight angle is indicated by a line where the concentric surface strise bend inward toward the pseudodeltidium across the narrow area; a sharp angle is formed where the convex pseudodeltidium rises abruptly from the area. Dorsal valve moderately convex, most elevated at the small umbo just in advance of the marginal, minute beak; area very low and narrow and without trace of pseudodeltidium as far as now known. Surface marked by fine, concentric, slightly undulating, thread- like strise and a varying number of irregular, more or less inter- rupted, narrow, depressed, rounded, radiating- ridges ; these ridges are usually most numerous at the central portions of the valves. The concentric striae extend across the narrow area and arch over the pseudodeltidium, where they are finer and crowded together, so that all the strise between the apex and the front margin are com- pressed in about one-half the distance on the pseudodeltidium. The adult ventral valve is about 4.5 mm. in length by 5 mm. in width and 2.5 mm. in height, with a pseudodeltidium 1.3 mm. in length. A dorsal valve 2 mm. in length has a height of about 0.5 mm. at the umbo. The shell is rather thick for a species of this size and it is built up of several thin layers or lamellae. Observations.—Micromitra hoydeni differs from the nearest re- lated species, M. scnlptilis (Meek) [1873, p. 479], in having a strong, 56 SMITHSONIAN MISCELIvANEOUS COIvLECTlONS VOL. 53 convex pseudodeltidium, less elevation of the ventral valve, and a thicker shell. .1/. haydciii occurs near the base of the Middle Cam- brian and AI. sculptilis about 2,000 feet (609.6 m.) higher in the section of the Cambrian rocks of Utah and southern Idaho. The speciiic name is given in honor of Dr. F. V. Hayden, geologist and explorer, under whose charge the geology of this region was first studied. Formation and Locality.— ]\Iiddle Cambrian : Limestone of the Langston formation, just above the Cambrian quartzitic sandstone beds, north side of Two Alile Canyon, near its mouth, 2 miles (2)-^^ km.) southeast of JMalad, Oneida County, Idaho. MICROMITRA SCULPTILIS ENDLICHI, new variety This form is represented by a single specimen of a ventral valve. The surface is similar to that of Microiiiitra sciilptilis (Aleek) [1873, p. 479], but the valve is more elongate, less elevated, and larger (5 mm. in diameter) than the specimens of the latter from the type locality. Formation and Locality.—Upper Cambrian : Limestone 2 miles (3.22 km.) north of Aurum, Schell Creek Range, White Pine County, Nevada. Subgenus IPHIDELLA Walcott [1905(7, p. 304] MICROMITRA (IPHIDELLA) LOUISE, new species Plate 7, Figures 4 and 4a In form this species is not unlike Microiiiitra pcalcl (Walcott) [1897/7, p. 712] and the more elongate forms of .1/. {IpliidcUa) paii- iiiila inaladciisis (Walcott) [1905a, p. 306]. It diiTers from both species mentioned in its surface characters. In the latter respect it is more like AI. (I.) iiyssa (see p. 57), but the form of AI. (/.) Ionise is more elongate and the apex of the ventral valve is nearer to the posterior margin ; the shell also appears to have been thicker. The surface characters are exceedingly minute. Under a glass magni- fying twenty diameters, the surface looks much like that of AI. (/.) pannula (White) [1874, p. 6]. The largest ventral valve in the collection has a length of 7.5 mm. and a width of 7 mm. ; elevation, I muL Alicroiuitra (Ipfiidclla) loitisc is the oldest brachiopod known from the Cambrian of the Canadian Rocky Mountains. In the Lakes Louise and Agnes section it is 3,100 feet (944.9 m.) below the sum- mit of the Lower Cambrian and 2,750 feet (838.2 m.) below the CAMBRIAN BRACK lOPODA WALCOTT 57 horizon which, on the basis of the associated faunas, is correlated with that at which .1/. (/.) nyssa occurs in Montana. It occurs in a fine, hard, dark gray, siHceous shale in association with Hyolithes, Cntaiaiia, and a fragment indicating the free cheek of a trilobite.
Recommended publications
  • Trace Fossils and Substrates of the Terminal Proterozoic–Cambrian Transition: Implications for the Record of Early Bilaterians and Sediment Mixing
    Trace fossils and substrates of the terminal Proterozoic–Cambrian transition: Implications for the record of early bilaterians and sediment mixing Mary L. Droser*†,So¨ ren Jensen*, and James G. Gehling‡ *Department of Earth Sciences, University of California, Riverside, CA 92521; and ‡South Australian Museum, Division of Natural Sciences, North Terrace, Adelaide 5000, South Australia, Australia Edited by James W. Valentine, University of California, Berkeley, CA, and approved August 16, 2002 (received for review May 29, 2002) The trace fossil record is important in determining the timing of the appearance of bilaterian animals. A conservative estimate puts this time at Ϸ555 million years ago. The preservational potential of traces made close to the sediment–water interface is crucial to detecting early benthic activity. Our studies on earliest Cambrian sediments suggest that shallow tiers were preserved to a greater extent than typical for most of the Phanerozoic, which can be attributed both directly and indirectly to the low levels of sediment mixing. The low levels of sediment mixing meant that thin event beds were preserved. The shallow depth of sediment mixing also meant that muddy sediments were firm close to the sediment–water interface, increasing the likelihood of recording shallow-tier trace fossils in muddy sed- iments. Overall, trace fossils can provide a sound record of the onset of bilaterian benthic activity. he appearance and subsequent diversification of bilaterian Tanimals is a topic of current controversy (refs. 1–7; Fig. 1). Three principal sources of evidence exist: body fossils, trace fossils (trails, tracks, and burrows of animal activity recorded in the sedimentary record), and divergence times calculated by means of a molecular ‘‘clock.’’ The body fossil record indicates a geologically rapid diversification of bilaterian animals not much earlier than the Precambrian–Cambrian boundary, the so-called Cambrian explosion.
    [Show full text]
  • Decoding the Fossil Record of Early Lophophorates
    Digital Comprehensive Summaries of Uppsala Dissertations from the Faculty of Science and Technology 1284 Decoding the fossil record of early lophophorates Systematics and phylogeny of problematic Cambrian Lophotrochozoa AODHÁN D. BUTLER ACTA UNIVERSITATIS UPSALIENSIS ISSN 1651-6214 ISBN 978-91-554-9327-1 UPPSALA urn:nbn:se:uu:diva-261907 2015 Dissertation presented at Uppsala University to be publicly examined in Hambergsalen, Geocentrum, Villavägen 16, Uppsala, Friday, 23 October 2015 at 13:15 for the degree of Doctor of Philosophy. The examination will be conducted in English. Faculty examiner: Professor Maggie Cusack (School of Geographical and Earth Sciences, University of Glasgow). Abstract Butler, A. D. 2015. Decoding the fossil record of early lophophorates. Systematics and phylogeny of problematic Cambrian Lophotrochozoa. (De tidigaste fossila lofoforaterna. Problematiska kambriska lofotrochozoers systematik och fylogeni). Digital Comprehensive Summaries of Uppsala Dissertations from the Faculty of Science and Technology 1284. 65 pp. Uppsala: Acta Universitatis Upsaliensis. ISBN 978-91-554-9327-1. The evolutionary origins of animal phyla are intimately linked with the Cambrian explosion, a period of radical ecological and evolutionary innovation that begins approximately 540 Mya and continues for some 20 million years, during which most major animal groups appear. Lophotrochozoa, a major group of protostome animals that includes molluscs, annelids and brachiopods, represent a significant component of the oldest known fossil records of biomineralised animals, as disclosed by the enigmatic ‘small shelly fossil’ faunas of the early Cambrian. Determining the affinities of these scleritome taxa is highly informative for examining Cambrian evolutionary patterns, since many are supposed stem- group Lophotrochozoa. The main focus of this thesis pertained to the stem-group of the Brachiopoda, a highly diverse and important clade of suspension feeding animals in the Palaeozoic era, which are still extant but with only with a fraction of past diversity.
    [Show full text]
  • Paterimitra Pyramidalis Laurie, 1986, the First Tommotiid Discovered From
    1 Paterimitra pyramidalis Laurie, 1986, the first tommotiid discovered from 2 the early Cambrian of North China 3 4 Bing Pana, b, Glenn A. Brockc, Christian B. Skovstedd, Marissa J. Bettse, Timothy P. Topperf, 5 Guo-Xiang Lia, * 6 7 a State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and 8 Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China 9 b University of Science and Technology of China, Hefei 230026, China 10 c Department of Biological Sciences, Macquarie University, NSW 2109, Australia 11 d Department of Palaeobiology, Swedish Museum of Natural History, Stockholm, Sweden. 12 e Palaeoscience Research Centre, School of Environmental and Rural Science, University of 13 New England, Armidale, NSW, Australia. 14 f Palaeoecosystems Group, Department of Earth Sciences, Durham University, Durham, UK. 15 * Corresponding author. 16 E-mail: [email protected] (B. Pan), [email protected] (G.A. Brock), 17 [email protected] (C.B. Skovsted), [email protected] (M.J. Betts), 18 [email protected] (T.P. Topper), [email protected] (G.X. Li) 19 20 ABSTRACT 21 The eccentrothecimorph tommotiid Paterimitra pyramidalis Laurie, 1986, was 22 previously only known from lower Cambrian rocks of the Northern Territory and South 23 Australia. Herein, we document the first occurrence of P. pyramidalis from the Xinji 24 Formation in the Shuiyu section at Ruicheng County, Shanxi Province, located at the 25 southwestern margin of the North China Platform. This represents the first report of a 1 26 tommotiid taxon from lower Cambrian strata of the North China Platform.
    [Show full text]
  • Resolving Details of the Nonbiomineralized Anatomy of Trilobites Using Computed
    Resolving Details of the Nonbiomineralized Anatomy of Trilobites Using Computed Tomographic Imaging Techniques Thesis Presented in Partial Fulfillment of the Requirements for the Master of Science in the Graduate School of The Ohio State University By Jennifer Anita Peteya, B.S. Graduate Program in Earth Sciences The Ohio State University 2013 Thesis Committee: Loren E. Babcock, Advisor William I. Ausich Stig M. Bergström Copyright by Jennifer Anita Peteya 2013 Abstract Remains of two trilobite species, Elrathia kingii from the Wheeler Formation (Cambrian Series 3), Utah, and Cornuproetus cornutus from the Hamar Laghdad Formation (Middle Devonian), Alnif, Morocco, were studied using computed tomographic (CT) and microtomographic (micro-CT) imaging techniques for evidence of nonbiomineralized alimentary structures. Specimens of E. kingii showing simple digestive tracts are complete dorsal exoskeletons preserved with cone-in-cone concretions on the ventral side. Inferred stomach and intestinal structures are preserved in framboidal pyrite, likely resulting from replication by a microbial biofilm. C. cornutus is preserved in non- concretionary limestone with calcite spar lining the stomach ventral to the glabella. Neither species shows sediment or macerated sclerites of any kind in the gut, which tends to rule out the possibilities that they were sediment deposit-feeders or sclerite-ingesting durophagous carnivores. Instead, the presence of early diagenetic minerals in the guts of E. kingii and C. cornutus favors an interpretation of a carnivorous feeding strategy involving separation of skeletal parts of prey prior to ingestion. ii Dedication This manuscript is dedicated to my parents for encouraging me to go into the field of paleontology and to Lee Gray for inspiring me to continue.
    [Show full text]
  • Oldest Mickwitziid Brachiopod from the Terreneuvian of Southern France
    Oldest mickwitziid brachiopod from the Terreneuvian of southern France LÉA DEVAERE, LARS HOLMER, SÉBASTIEN CLAUSEN, and DANIEL VACHARD Devaere, L., Holmer, L., Clausen, S., and Vachard, D. 2015. Oldest mickwitziid brachiopod from the Terreneuvian of southern France. Acta Palaeontologica Polonica 60 (3): 755–768. Kerberellus marcouensis Devaere, Holmer, and Clausen gen. et sp. nov., originally described as Dictyonina? sp., from the Terreneuvian of northern Montagne Noire (France) is re-interpreted as the oldest relative to or member of mickwitziid- like stem-group brachiopods. We extracted 170 partial to complete phosphatic internal moulds of two types of adult and one type of juvenile disarticulated valves, rarely externally coated with phosphates, from the calcareous Heraultia Member of the Marcou Formation. They correspond to microbially infested, ventribiconvex, inequivalved, bivalved shells. The ventral interarea is bisected by a triangular sinus. The shell, most probably dominantly organic in origin, is orthogonally pierced throughout its entire thickness by radially-aligned, smooth-walled, cylindrical to hour-glass shaped canals except for the sub-apical planar field (interarea). The through-going canals of K. marcouensis are compared with brachiopods endopunctae and with canals of mickwitziid brachiopods. The absence of striations on K. marcouensis canal walls, typical of mickwitziids, implies that (i) the tubes could have been depleted of setae or; (ii) traces of the microvilli were not preserved on the tube wall (taphonomic bias) or, (iii) the tubes could have been associated with an outer epithelial follicle. Key words: Brachiopoda, Mickwitziidae, shell canals, Cambrian, Terreneuvian, West Gondwana, France. Léa Devaere [[email protected]], Sébastien Clausen [[email protected]], and Daniel Vachard [[email protected]], UMR 8217 Géosystèmes CNRS-Université Lille 1, bâtiment SN5, avenue Paul Lan- gevin, 59655 Villeneuve d’Ascq, France.
    [Show full text]
  • Phenotypic Variation? a Case Study from the Burgess Shale.', Palaeontology., 60 (2)
    Durham Research Online Deposited in DRO: 11 June 2018 Version of attached le: Accepted Version Peer-review status of attached le: Peer-reviewed Citation for published item: Topper, Timothy P. and Strotz, Luke C. and Skovsted, Christian B. and Holmer, Lars E. (2017) 'Do brachiopods show substrate-related phenotypic variation? A case study from the Burgess Shale.', Palaeontology., 60 (2). pp. 269-279. Further information on publisher's website: https://doi.org/10.1111/pala.12281 Publisher's copyright statement: This is the accepted version of the following article: Topper, Timothy P., Strotz, Luke C., Skovsted, Christian B. Holmer, Lars E. (2017). Do brachiopods show substrate-related phenotypic variation? A case study from the Burgess Shale. Palaeontology 60(2): 269-279, which has been published in nal form at https://doi.org/10.1111/pala.12281. This article may be used for non-commercial purposes in accordance With Wiley Terms and Conditions for self-archiving. Additional information: Use policy The full-text may be used and/or reproduced, and given to third parties in any format or medium, without prior permission or charge, for personal research or study, educational, or not-for-prot purposes provided that: • a full bibliographic reference is made to the original source • a link is made to the metadata record in DRO • the full-text is not changed in any way The full-text must not be sold in any format or medium without the formal permission of the copyright holders. Please consult the full DRO policy for further details. Durham University Library, Stockton Road, Durham DH1 3LY, United Kingdom Tel : +44 (0)191 334 3042 | Fax : +44 (0)191 334 2971 https://dro.dur.ac.uk 1 DO BRACHIOPODS SHOW SUBSTRATE-RELATED 2 PHENOTYPIC VARIATION? A CASE STUDY FROM THE 3 BURGESS SHALE 4 5 TIMOTHY P.
    [Show full text]
  • Chapter 5. Paleozoic Invertebrate Paleontology of Grand Canyon National Park
    Chapter 5. Paleozoic Invertebrate Paleontology of Grand Canyon National Park By Linda Sue Lassiter1, Justin S. Tweet2, Frederick A. Sundberg3, John R. Foster4, and P. J. Bergman5 1Northern Arizona University Department of Biological Sciences Flagstaff, Arizona 2National Park Service 9149 79th Street S. Cottage Grove, Minnesota 55016 3Museum of Northern Arizona Research Associate Flagstaff, Arizona 4Utah Field House of Natural History State Park Museum Vernal, Utah 5Northern Arizona University Flagstaff, Arizona Introduction As impressive as the Grand Canyon is to any observer from the rim, the river, or even from space, these cliffs and slopes are much more than an array of colors above the serpentine majesty of the Colorado River. The erosive forces of the Colorado River and feeder streams took millions of years to carve more than 290 million years of Paleozoic Era rocks. These exposures of Paleozoic Era sediments constitute 85% of the almost 5,000 km2 (1,903 mi2) of the Grand Canyon National Park (GRCA) and reveal important chronologic information on marine paleoecologies of the past. This expanse of both spatial and temporal coverage is unrivaled anywhere else on our planet. While many visitors stand on the rim and peer down into the abyss of the carved canyon depths, few realize that they are also staring at the history of life from almost 520 million years ago (Ma) where the Paleozoic rocks cover the great unconformity (Karlstrom et al. 2018) to 270 Ma at the top (Sorauf and Billingsley 1991). The Paleozoic rocks visible from the South Rim Visitors Center, are mostly from marine and some fluvial sediment deposits (Figure 5-1).
    [Show full text]
  • Resume of the Geology of Arizona," Prepared by Dr
    , , A RESUME of the GEOWGY OF ARIZONA by Eldred D. Wilson, Geologist THE ARIZONA BUREAU OF MINES Bulletin 171 1962 THB UNIVBR.ITY OP ARIZONA. PR••• _ TUC.ON FOREWORD CONTENTS Page This "Resume of the Geology of Arizona," prepared by Dr. Eldred FOREWORD _................................................................................................ ii D. Wilson, Geologist, Arizona Bureau of Mines, is a notable contribution LIST OF TABLES viii to the geologic and mineral resource literature about Arizona. It com­ LIST OF ILLUSTRATIONS viii prises a thorough and comprehensive survey of the natural processes and phenomena that have prevailed to establish the present physical setting CHAPTER I: INTRODUCTION Purpose and scope I of the State and it will serve as a splendid base reference for continued, Previous work I detailed studies which will follow. Early explorations 1 The Arizona Bureau of Mines is pleased to issue the work as Bulletin Work by U.S. Geological Survey.......................................................... 2 171 of its series of technical publications. Research by University of Arizona 2 Work by Arizona Bureau of Mines 2 Acknowledgments 3 J. D. Forrester, Director Arizona Bureau of Mines CHAPTER -II: ROCK UNITS, STRUCTURE, AND ECONOMIC FEATURES September 1962 Time divisions 5 General statement 5 Methods of dating and correlating 5 Systems of folding and faulting 5 Precambrian Eras ".... 7 General statement 7 Older Precambrian Era 10 Introduction 10 Literature 10 Age assignment 10 Geosynclinal development 10 Mazatzal Revolution 11 Intra-Precambrian Interval 13 Younger Precambrian Era 13 Units and correlation 13 Structural development 17 General statement 17 Grand Canyon Disturbance 17 Economic features of Arizona Precambrian 19 COPYRIGHT@ 1962 Older Precambrian 19 The Board of Regents of the Universities and Younger Precambrian 20 State College of Arizona.
    [Show full text]
  • Brachiopods from the Lower-Middle Cambrian Láncara Formation of the Cantabrian Mountains, Northwest Spain
    Brachiopods from the Lower-Middle Cambrian Láncara Formation of the Cantabrian Mountains, Northwest Spain THOMAS WOTTE & MICHAL MERGL WOTTE, T. & MERGL, M., 2007:09:03. Brachiopods from the Lower-Middle Cambrian Láncara Formation of the Cantabrian Mountains, Northwest Spain. Memoirs of the Association of Australasian Palaeontologists 33, 101-122. ISSN 0810-8889. Micro- and macrobrachiopods from eight stratigraphic sections of the carbonate upper member of the Middle Cambrian Láncara Formation (comprising lower Beleño facies and upper Barrios facies) in the Cantabrian Mountains are analysed. They comprise ten species, two of which are new, assigned to nine genera, two of which are new. They are: Acrothele primaeva (de Verneuil & Barrande in de Prado, 1860), Acrothele sp., Eoobolidae gen. et sp. indet., Genetreta trilix gen. et sp. nov., Iberotreta sampelayoi gen. et sp. nov., Luhotreta? proclinis (Mergl & Elicki, 2004), Micromitra cf. sculptilis (Meek, 1873), Nisusia vaticina (de Verneuil & Barrande in de Prado, 1860), Trematobolus simplex (Vogel, 1962) and Yorkia zafrensis (Gil-Cid & Mélou, 1986). Stratigraphic distribution patterns of the brachiopod fauna mirror the drowning of the environment. Trematobolus simplex is exclusively found in the carbonates of the Beleño facies, whereas Nisusia vaticina and Yorkia zafrensis are typical representatives of the nodular limestones of the Barrios facies. The species of the genera Trematobolus, Nisusia and Yorkia demonstrate the affinity of the upper Láncara brachiopod association with faunas of the Siberian platform, New South Wales and some Avalonian terranes (Newfoundland, New Brunswick). Thomas Wotte ([email protected]), Geological Institute, Freiberg University of Mining and Technology, Bernhard-von-Cotta Street 2, D-09599 Freiberg, Germany; Michal Mergl ([email protected]), Department of Biology, University of West Bohemia at Plzeň, Klatovská 51, CZ-306 19, Plzeň, Czech Republic.
    [Show full text]
  • The Tommotiid Camenella Reticulosa from the Early Cambrian of South Australia: Morphology, Scleritome Reconstruction, and Phylogeny
    The tommotiid Camenella reticulosa from the early Cambrian of South Australia: Morphology, scleritome reconstruction, and phylogeny CHRISTIAN B. SKOVSTED, UWE BALTHASAR, GLENN A. BROCK, and JOHN R. PATERSON Skovsted, C.B., Bathasar, U., Brock, G.A., and Paterson, J.R. 2009. The tommotiid Camenella reticulosa from the early Cambrian of South Australia: Morphology, scleritome reconstruction, and phylogeny. Acta Palaeontologica Polonica 54 (3): 525–540. DOI: 10.4202/app.2008.0082. The tommotiid Camenella reticulosa is redescribed based on new collections of well preserved sclerites from the Arrowie Basin (Flinders Ranges), South Australia, revealing new information concerning morphology and micro− structure. The acutely pyramidal mitral sclerite is described for the first time and the sellate sclerite is shown to be coiled through up to 1.5 whorls. Based on Camenella, a model is proposed by which tommotiid sclerites are composed of alternating dense phosphatic, and presumably originally organic−rich, laminae. Camenella is morphologically most similar to Lapworthella, Kennardia,andDailyatia, and these taxa are interpreted to represent a monophyletic clade, here termed the “camenellans”, within the Tommotiida. Potential reconstructions of the scleritome of Camenella are discussed and although a tubular scleritome construction was recently demonstrated for the tommotiids Eccentrotheca and Paterimitra, a bilaterally symmetrical scleritome model with the sclerites arranged symmetrically on the dorsal surface of a vagrant animal can not be ruled out. Key words: Tommotiida, Camenella, scleritome, phylogeny, Atdabanian, Botoman, Cambrian, South Australia. Christian B. Skovsted [[email protected]] and Uwe Balthasar [[email protected]], Department of Earth Sciences, Palaeobiology, Uppsala University, Villavägen 16, SE−752 36 Uppsala, Sweden; Glenn A.
    [Show full text]
  • Paleoecology of the Greater Phyllopod Bed Community, Burgess Shale ⁎ Jean-Bernard Caron , Donald A
    Available online at www.sciencedirect.com Palaeogeography, Palaeoclimatology, Palaeoecology 258 (2008) 222–256 www.elsevier.com/locate/palaeo Paleoecology of the Greater Phyllopod Bed community, Burgess Shale ⁎ Jean-Bernard Caron , Donald A. Jackson Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, Ontario, Canada M5S 3G5 Accepted 3 May 2007 Abstract To better understand temporal variations in species diversity and composition, ecological attributes, and environmental influences for the Middle Cambrian Burgess Shale community, we studied 50,900 fossil specimens belonging to 158 genera (mostly monospecific and non-biomineralized) representing 17 major taxonomic groups and 17 ecological categories. Fossils were collected in situ from within 26 massive siliciclastic mudstone beds of the Greater Phyllopod Bed (Walcott Quarry — Fossil Ridge). Previous taphonomic studies have demonstrated that each bed represents a single obrution event capturing a predominantly benthic community represented by census- and time-averaged assemblages, preserved within habitat. The Greater Phyllopod Bed (GPB) corresponds to an estimated depositional interval of 10 to 100 KA and thus potentially preserves community patterns in ecological and short-term evolutionary time. The community is dominated by epibenthic vagile deposit feeders and sessile suspension feeders, represented primarily by arthropods and sponges. Most species are characterized by low abundance and short stratigraphic range and usually do not recur through the section. It is likely that these are stenotopic forms (i.e., tolerant of a narrow range of habitats, or having a narrow geographical distribution). The few recurrent species tend to be numerically abundant and may represent eurytopic organisms (i.e., tolerant of a wide range of habitats, or having a wide geographical distribution).
    [Show full text]
  • PALAEONTOLOGICAL ASSOCIATION 47Th Annual Meeting Department
    PALAEONTOLOGICAL ASSOCIATION 47th Annual Meeting Department of Geology, University of Leicester December 14-17, 2003 ABSTRACTS of TALKS ABSTRACTS of POSTERS Compiled and edited by Mark Purnell Oral presentations Rediscovery of the Gutterford Burn ‘Eurypterid Bed’ Pentland Hills, Midlothian, Scotland Lyall I. Anderson National Museums of Scotland, Chambers St, Edinburgh, EH1 1JF <[email protected]> A programme of field excavation undertaken by National Museums of Scotland staff, volunteers and other interested parties during early July 2003 had three main aims. Firstly to relocate the exact position of the ‘Eurypterid Bed’, a fossiliferous sediment which in the late 1880’s yielded the world’s most diverse assemblage of Silurian chelicerate arthropods, secondly to characterise the likely sedimentary depositional setting and preservational mechanics of this Fossil Konservat-Lagerstätte, and thirdly to investigate the wider relevance of this fossiliferous unit to the more fully known sequences lying stratigraphically higher in the inlier as detailed by the work of Clarkson et al. (2001). After extensive excavation, the ‘Eurypterid Bed’ lithology was located in situ on the banks of the Gutterford Burn stream section. Detailed sedimentary logging and sampling indicated that volcaniclastic sediments played a major role in the formation of the bed; both discrete ashfall bands and ash-rich sediment were discovered in the metre-thick unit along with monograptids. Overlying the ‘eurypterid bed’ occur sporadic horizons yielding dendroid graptolites and numerous (at least 12) thin, discontinuous bands of decalcified marine limestone, rich in invertebrate remains. The fauna within these bands shows a degree of similarity with that identified in the overlying Deerhope Formation.
    [Show full text]