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Paper No: 01 Ecosystem Structures & Functions Module: 26 Desert Ecosystem Development Team Prof. R.K. Kohli Principal Investigator & Prof. V.K. Garg &Prof.Ashok Dhawan Co- Principal Investigator Central University of Punjab, Bathinda Dr. Renuka Gupta, YMCA University of Science and Paper Coordinator Technology, Faridabad, Haryana Dr. Sharda R. Gupta, former Professor Content Writer Kurukshetra University, Kurukshetra Content Reviewer Prof. V.K. Garg, Central University of Punjab, Bathinda Anchor Institute Central University of Punjab 1 Ecosystem Structures & Functions Environmental Sciences 26 Desert Ecosystem Description of Module Subject Name Environmental Sciences Paper Name Ecosystem Structures & Functions Module Name/Title 26 Desert Ecosystem Module Id EVS/ESF-I/26 Pre-requisites To learn about the distribution, abiotic environment, biodiversity, and Objectives functions of the desert ecosystems Aridity Index, Hyperarid and arid regions, hot and dry desert, Keywords cold/temperate desert, coastal desert, biodiversity, adaptations, productivity, foodwebs, land degradation, resource management 2 Ecosystem Structures & Functions Environmental Sciences 26 Desert Ecosystem Module 26: Desert Ecosystem Objectives: Controlling Factors Forming Deserts The distribution and climate of deserts Adaptation of Plants and Animals to Dry conditions Biodiversity and Community Composition • Desert ecosystem functions • Human impact on Desert Ecosystems 3 Ecosystem Structures & Functions Environmental Sciences 26 Desert Ecosystem 26.1. Introduction Dry areas as created by global circulation patterns contain most of the deserts on the Earth. Mostly, deserts occur in specific latitudes (25 to 35° north and south of the equator). Deserts are arid where less than 250 mm of rain falls in a year resulting in low soil water availability, sparse growth of vegetation, and low productivity. However, evaporation exceeds rainfall in these ecosystems. The deserts in the world are not restricted by latitude, longitude, or elevation and show extreme variability in abiotic and biotic components. The causes of formation of deserts are varied; they show disjunct distribution and have independent floral histories. Mostly, deserts are often regions of extreme temperatures where living conditions are harsh; temperature variability is extremely variable. For example, many deserts such as the Sahara in Africa are hot all year-round but others, such as Asia's Gobi desert, become quite cold in winter. It is interesting that deserts are having high biodiversity in spite of their low productivity. The world’s deserts occupy almost 17 percent of the earth’s land surface. The deserts and desert margins are home to about eight per cent of the global human population; some of the poorest and most marginalized people in the world live in deserts. The poor people depend on deserts for sustainable ecosystem services under changing climate. 26.2. Controlling Factors Forming Deserts 26.2.1. Earth’s air circulation patterns: Subtropical deserts are formed because of the circulation patterns of air masses. They are found along the Tropic of Cancer, between 15o and 30 o north of the Equator, or along the Tropic of Capricorn, between 15o and 30o south of the Equator. Included in this group are the Sahara Desert and the Kalahari Desert in Africa. In these regions, hot, moist air rises into the atmosphere near the Equator. As the air rises, it cools and drops its moisture as heavy tropical rains. The resulting cooler, drier air mass moves away from the Equator. As it approaches the tropics, the air descends and warms up again. The descending air hinders the formation of clouds, so very little rain falls on the land below. 26.2.2. Rain shadow effect: A rain shadow is a patch of land that becomes a desert because mountain ranges blocked all rainy weather. On one side of the mountain, wet weather systems drop rain and snow. On the other side of the mountain, the rain shadow side, it’s warm and dry (Fig.26.1). When 4 Ecosystem Structures & Functions Environmental Sciences 26 Desert Ecosystem an air mass moves from a low elevation to a high elevation, it expands and cools. Cool air forms clouds, causing rain and snow, as it rises up a mountain. After the air mass crosses over the peak of the mountain and starts down the other side, the air warms up and the clouds dissipate resulting in less rainfall. Rain shadow deserts exist near the leeward slopes of some mountain ranges. For example, the Gobi Desert, The Tibetan Plateau, the deserts of Nevada east of the Sierra Nevada Mountains. Major rain shadow effects occur in Atacama Desert, located in Chile. 26.2.3. Distance from the ocean. Some land is so far from oceans (where air absorbs most of its water) that deserts form. Interior deserts, which are found in the heart of continents, exist because no moisture-laden winds reach them. The Gobi Desert, in China and Mongolia, lies hundreds of kilometers from the ocean. Winds that reach the Gobi have very low moisture. The Gobi is also in the rain shadow of the Himalayan Mountains to the south. 26.2.4. Near cold ocean currents: Cold ocean currents contribute to the formation of coastal deserts. The cold ocean air warms as it passes over continents. A coastal desert may be almost totally rainless, yet damp with fog. The Atacama Desert, on the Pacific shores of Chile, is a coastal desert. Fig. 1. The rain shadow effect, windward and leeward sides of a mountain range, the windward side receives heavy precipitation while the Leeward side is dry ( adapted from Singh et al., 2015). 5 Ecosystem Structures & Functions Environmental Sciences 26 Desert Ecosystem 26.3. The main Characteristics of Deserts Deserts are dry and arid, have an aridity index (P/PET) lower than 0.20. Humidity (water vapor in the air) is near zero in most deserts. Light rains often evaporate in the dry air, never reaching the ground. Temperature extremes are a characteristic of most deserts. Annual precipitation less than the potential evaporation. Low soil moisture to support primary productivity. Animals and plants that live in deserts have adapted to survive in harsh conditions. Many desert dwellers rely on groundwater, stored in aquifers below the surface. 26.4. The Desert Ecosystems: A Global Perspective Aridity is an important criterion for defining a desert. One of the most common approaches to measure aridity is through an estimator called the Aridity Index, which is the ratio between mean annual precipitation (P) and mean annual potential evapotranspiration (PET) (Thornthwaite 1948). The hyperarid and arid regions of the world are defined as those areas with an aridity index (P/PET) lower than 0.20 (Table26.1). Potential evapotranspiration (PET) is calculated from Thornthwaite’s (1948) equations as a function of mean monthly temperatures and mean monthly number of daylight hours, while precipitation (P) is measured directly from weather stations. Arid and hyper-arid regions have a P/PET ratio of less than 0.20; rainfall supplies less than 20 per cent of the amount of water needed to support optimum plant growth (UNEP 1997). Based on this criterion, aridity is highest in the Saharan and Chilean-Peruvian deserts, followed by the Arabian, East African, Gobi, Australian, and South African Deserts, and it is generally lower in the Thar and North American deserts. 6 Ecosystem Structures & Functions Environmental Sciences 26 Desert Ecosystem Table 26.1. Area of hyper-arid and arid regions of the world (based on UNEP 1997; Safriel et al. 2005) Classification Aridity Index Area of world Area of world (P/PET)* total(%) total(%) (UNEP 1997) (Safriel et al. 2005) Hyperarid < 0.05 7.5 6.6 Arid 0.05–0.20 12.1 10.6 * The ratio of precipitation (P) to potential evapotranspiration (PET) The global distribution of deserts as Hyperarid and arid regions is shown in Fig. 26.2. In the northern hemisphere, the deserts are found in three regions (UNEP, 2006, http://www.unep.org/geo/gdoutlook/112.asp): i. The Mojave, Sonoran, and Chihuahuan Deserts in North America, ii. The Sahara desert forms an immense swathe in Northern Africa and the Somali-Ethiopian deserts in the Horn of Africa. iii. The deserts of Asia, including the Arabian, Mesopotamian, Persian, and Thar deserts that stretch from West Asia into Pakistan and India, as well as the Central Asian deserts in Uzbekistan, Turkmenistan, and the Taklimakan and Gobi deserts in China and Mongolia. The world's largest desert, the Sahara of North Africa, which experiences temperatures as high as 57°C, is a trade wind desert. Mid latitude deserts occur between 30° and 50° N. and S, poleward of the subtropical high pressure zones. These deserts are in interior drainage basins far from oceans and have a wide range of annual temperatures. The Sonoran Desert of southwestern North America- is a typical mid latitude desert. The Rajasthan Desert of India and the Thar Desert of Pakistan are parts of a monsoon desert region west of the range. 7 Ecosystem Structures & Functions Environmental Sciences 26 Desert Ecosystem In the southern hemisphere, the desert chainis formed by (i) the Atacama, Puna, and Monte Deserts in South America, (ii ) the Namib and the Karoo in southern Africa, and (iii) the vast expanse of the Australian Deserts, ( UNEP 2006 and references there in) ( http://www.unep.org/geo/gdoutlook/112.asp). Parts of the Arctic and the Antarctic are classified as deserts. These polar deserts contain great quantities of water, but most of it is locked in glaciers and ice sheets year-round. The largest desert in the world is also the coldest. Almost the entire continent of Antarctica is a polar desert, experiencing little precipitation. Few organisms can withstand the freezing, dry climate of Antarctica Fig. 26.2. The hyperarid and arid regions representing deserts cover about 17 percent of the Earth's land surface and are found on every continent.