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Обзор Типовых Экземпляров Скальных Ящериц Комплекса Darevskia (Caucasica) (Sauria: Lacertidae)

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Обзор Типовых Экземпляров Скальных Ящериц Комплекса Darevskia (Caucasica) (Sauria: Lacertidae) Труды Зоологического института РАН Том 318, № 4, 2014, c. 371–381 УДК 598.113.6 ОБЗОР ТИПОВЫХ ЭКЗЕМПЛЯРОВ СКАЛЬНЫХ ЯЩЕРИЦ КОМПЛЕКСА DAREVSKIA (CAUCASICA) (SAURIA: LACERTIDAE) И.В. Доронин Зоологический институт Российской академии наук, Университетская наб. 1, 199034 Санкт-Петербург, Россия; e-mail: [email protected] РЕЗЮМЕ В работе приводится информация о современном месте хранения типовых экземпляров ящериц комплекса Darevskia (caucasica) по состоянию на август 2014 г. Для стабилизации номенклатуры выделяется лектотип Lacerta saxicola daghestanica Darevsky, 1967. Дается описание внешней морфологии паралектотипа L. s. da- ghestanica, лектотипа и паралектотипа Lacerta caucasica Méhely, 1909. Приводится история описания форм комплекса. Ключевые слова: комплекс Darevskia (caucasica), скальные ящерицы, типовые экземпляры REVIEW OF TYPE SPECIMENS OF ROCK LIZARDS OF DAREVSKIA (CAUCASICA) COMPLEX (SAURIA: LACERTIDAE) I.V. Doronin Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia; e-mail: [email protected] ABSTRACT The paper provides information on the current status of the type specimens of Darevskia (caucasica) complex as they were in August 2014. In order to stabilize nomenclature of this complex, the lectotype of Lacerta saxicola daghestanica Darevsky, 1967 is designated. The description of external morphology of paralektotype of a L. s. da- ghestanica, lektotype and paralektotype of Lacerta caucasica Méhely, 1909 is given. History of description of these taxa is discussed. Keywords: Darevskia (caucasica) complex, rock lizards, type specimens ВВЕДЕНИЕ название, замененное на nomen nova Lacerta sa- xicola daghestanica Darevsky, 1967 (= Darevskia da- По современным представлениям (Darevsky ghestanica), Lacerta caucasica alpina Darevsky, 1967 and Roitberg 1999; Arnold et al. 2007; Лотиев и (= Darevskia alpina) и Lacerta caucasica vedenica Доронин [Lotiev and Doronin] 2011) в объеме над- Darevsky et Roitberg, 1999 (= Darevskia cau ca sica видового комплекса Darevskia (caucasica) были vedenica). описаны следующие таксоны: Lacerta caucasica Изучение этих скальных ящериц в ряде регио- Méhely, 1909 (= Darevskia caucasica), Lacerta saxi- нов затруднено тем, что ареалы морфологически cola gracilis Méhely, 1909 – преоккупированное близких видов перекрываются, а также наличием 372 И.В. Доронин предполагаемой зоны вторичной интеграда- М. Франценом (Michael Franzen) и Л. Акер (Linda ции, что относится прежде всего к D. caucasica Acker). и D. daghestanica (Fu et al. 1995; Darevsky and Сокращения названий научных учреждений: Roitberg 1999; Ройтберг [Roitberg] 1999а; Roitberg ZISP – Зоологический институт Российской 1999b). Ситуация осложняется отсутствием ре- академии наук, Санкт-Петербург, Россия; SMF – визии номенклатурного типа (есть синтипы из Зенкенбергский музей (Senckenberg Museum), разных локалитетов) для D. daghestanica и, как Франкфурт-на-Майне, Германия; GNM – Наци- следствие, отсутствием ограниченной типовой ональный музей Грузии им. Симона Джанашиа территории (= типового местонахождения, type (бывший Кавказский музей), Тбилиси, Грузия; locality). Цель данной публикации – ревизия ти- MN KNU – Музей природы Харьковского на- повых экземпляров обозначенных таксонов. ционального университета им. В.Н. Каразина, Харьков, Украина. МАТЕРИАЛ И МЕТОДЫ СИСТЕМАТИКА При описании внешней морфологии экземпля- ров скальных ящериц была взята традиционная 1. Lacerta saxicola Subsp. gracilis Méhely, 1909: 555 схема, используемая рядом авторов (Даревский [Darevsky] 1967; Arnold et al. 2007). Промеры, мм: Этот таксон был описан как подвид «n. subsp.» L. – длина тела от кончика морды до клоакальной Лайошом Мехели по 15 экз. из коллекции GNM: щели; L. cd. – длина нерегенерированного хвоста; 10 половозрелых самок, самцов и ювенильных Pil. – расстояние от кончика морды до заднего края особей, добытых в Дагестане, Россия, 07.1885 г. теменных щитков; Lt. c. – наибольшая ширина го- (№ 26 n1) (coll. Г.И. Радде), две половозрелых сам- ловы, измеренная на уровне барабанных щитков; ки из с. Лаваши 11.06.1894 г. (= Леваши), Дагестан, Al. c. – высота головы в районе затылочного щитка. Россия (coll. Г.И. Радде и Е.Г. Кениг) (№ 26 h), Фолидоз: Lab. – количество верхнегубных щитков один неполовозрелый самец и одна самка из (слева – справа); F. l. – количество верхнегубных Евдокимовска (= Евдокимовский пост), Чечня, щитков спереди от подглазничного; S. l. – количе- Россия, 11.07.1894 (coll. Г.И. Радде и Е.Г. Кениг) ство нижнегубных щитков (слева – справа); Mas. – (№ 26 e), один самец из г. Сухум-Кале (= Сухум, наличие и размер центральновисочного щитка; Сухуми), Абхазия, 05.1893 (coll. Г.И. Радде и Mas./Tym. – количество чешуй в наиболее узком Е.Г. Кениг) (№ 26 s), (Méhely 1909). месте между центральновисочным и барабанным В тексте описания не был обозначен тип, что щитками (слева – справа); Sup. gran. – количество дает основание считать перечисленные экземпля- гранул между верхнересничными и надглазнич- ры синтипами (International Code of Zoological ными щитками (слева – справа), а также наличие Nomenclature 1999) (Кодекс ст. 73.2). разрывов в их ряду; Sup. – количество чешуй Lacerta saxicola gracilis – преоккупированное вдоль края теменного за верхневисочным щитком; название (Lacerta gracilis Eichwald, 1841 – млад- G. – количество горловых чешуй по линии между ший синоним Eremias velox (Pallas, 1771) (Щербак серединой воротника и соединением нижнече- [Szczerbak] 1974; Барабанов [Barabanov] 2009)). люстных щитков; Col. – количество увеличенных В связи с этим Даревский ([Darevsky] 1967) ввел щитков в горловом воротнике; Sq. – количество новое замещающее название nomen novum Lacerta спинных чешуй в одном поперечном ряду вокруг saxicola daghestanica Darevsky, 1967. Согласно Ко- середины тела; P. fm. – количество бедренных пор дексу (ст. 72) номенклатурный тип (в данном слу- (слева – справа); Pr. an.1 – количество преаналь- чае – синтипы) для Lacerta saxicola daghestanica ных щитков в первом ряду; Pr. an.2 – количество тот же, что и для Lacerta saxicola gracilis. увеличенных преанальных щитков. В настоящее время два синтипа из 10, собран- Все промеры выполняли штангенциркулем с ных в 1885 г. в Дагестане (без указания более точ- точностью до 0.1 мм. Изучение фолидоза проведе- но под стереоскопическим микроскопом МСП-1. Параметры фолидоза лектотипа Lacerta 1В каталоге Кавказского музея под этим номером указа- cauca sica взяты с фотографий, предоставленных но не 10, а 11 экз. (Radde 1899). Обзор типовых экземпляров Darevskia (caucasica) 373 (старый № 26 h) (Рис. 3), Российская Федерация, Дагестан, с. Леваши (= Лаваши), 42°25´N 47°19´E (Рис. 4), 11.06.1894. Coll.: Г.И. Радде и Е.Г. Кениг. Lectotype (here designated) Lacerta sa xi- cola daghestanica Darevsky, 1967. GNM 263 (old number 26 h) (Fig. 3), Russian, Dagestan, Levashinsky District, Levaschi (=Lavaschi) Village, 42°25´N 47°19´E (Fig. 4), 11.06.1894. Coll.: G.I. Radde and E.G. Konig. Рис. 1. Паралектотип Lacerta saxicola gracilis Méhely, 1909 Описание лектотипа Lacerta saxicola daghe- (GNM 267) плохой сохранности. sta nica Darevsky, 1967. Половозрелая самка. Fig. 1. Paralectotype Lacerta saxicola gracilis Méhely, 1909 (GNM Длина туловища от конца морды до клоакальной 267) of poor preservation. щели – 49 мм. Длина от клоакальной щели до кончика хвоста – 42 мм; хвост поврежден. Рас- ной локализации), из коллекции GNM находятся стояние от конца морды до заднего края теменных в MN KNU (№ 27217-27218). Их принадлежность щитков – 10.5 мм; наибольшая ширина головы в к типовой серии подтверждается местом и вре- районе барабанного щитка – 6.7 мм; высота голо- менем сбора и указанием места первоначального вы в районе затылочного щитка – 4.2 мм. Ширина хранения – Кавказского музея. Они приведены лобоносового щитка в 1.4 раза превышает его в каталоге герпетологической коллекции музея длину. Межчелюстной щиток не касается лобо- (Зиненко и Гончаренко [Zinenko and Goncharenko] носового. Швы между предлобными и лобным 2011), но не помещены в каталог типовых экзем- щитками не вогнуты внутрь лобного. Между пляров (Vedmederya et al. 2009). Можно пред- верхнересничными и надглазничными щитками, положить, что эти ящерицы были получены из разделяя их, расположен полный ряд из 11 слева Кавказского музея для изучения Александром Михайловичем Никольским (1858–1942), кото- рый жил и работал в Харькове с 1903 г. При работе с герпетологической коллекцией GNM в мае 2014 г. было обнаружено только два синтипа: один (самец?) из Евдокимовска (Рис. 1), который сильно поврежден (работа с его морфо- логическими параметрами невозможна) и одна половозрелая самка из с. Леваши, которую я обо- значаю в качестве лектотипа. Отмечу, что этот экземпляр снабжен этикеткой Л. Мехели (Рис. 2). Лектотип (обозначается здесь) Lacerta sa- xi cola daghestanica Darevsky, 1967. GNM 263 Рис. 2. Этикетка Л. Мехели из коллекции Национального Fig. 3. Лектотип Lacerta saxicola daghestanica (GNM 263): A – музея Грузии им. Симона Джанашиа. общий вид; B – голова, вид слева; С – анальная область. Fig. 2. Label of L. Méhely from the collection of the Georgian Fig. 3. Lectotype of Lacerta saxicola daghestanica (GNM 263): National Museum named Simon Janashia. A – general view; B – left lateral view of the head; C – anal region. 374 И.В. Доронин Рис. 4. Скальные выходы в с. Леваши – место обитания Darevskia daghestanica на типовой территории (фото. Л.Ф. Мазанаевой). Fig. 4. Rock outcrops in the Levaschi Village, a habitat of Darevskia daghestanica in the type locality (photo. by L.F. Mazanaeva). и 10 справа зернышек.
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