Nimbadon, a I:,Iew Genus and Three New Species Of
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SUPPLEMENTARY INFORMATION for a New Family of Diprotodontian Marsupials from the Latest Oligocene of Australia and the Evolution
Title A new family of diprotodontian marsupials from the latest Oligocene of Australia and the evolution of wombats, koalas, and their relatives (Vombatiformes) Authors Beck, RMD; Louys, J; Brewer, Philippa; Archer, M; Black, KH; Tedford, RH Date Submitted 2020-10-13 SUPPLEMENTARY INFORMATION FOR A new family of diprotodontian marsupials from the latest Oligocene of Australia and the evolution of wombats, koalas, and their relatives (Vombatiformes) Robin M. D. Beck1,2*, Julien Louys3, Philippa Brewer4, Michael Archer2, Karen H. Black2, Richard H. Tedford5 (deceased) 1Ecosystems and Environment Research Centre, School of Science, Engineering and Environment, University of Salford, Manchester, UK 2PANGEA Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales, Australia 3Australian Research Centre for Human Evolution, Environmental Futures Research Institute, Griffith University, Queensland, Australia 4Department of Earth Sciences, Natural History Museum, London, United Kingdom 5Division of Paleontology, American Museum of Natural History, New York, USA Correspondence and requests for materials should be addressed to R.M.D.B (email: [email protected]) This pdf includes: Supplementary figures Supplementary tables Comparative material Full description Relevance of Marada arcanum List of morphological characters Morphological matrix in NEXUS format Justification for body mass estimates References Figure S1. Rostrum of holotype and only known specimen of Mukupirna nambensis gen. et. sp. nov. (AMNH FM 102646) in ventromedial (a) and anteroventral (b) views. Abbreviations: C1a, upper canine alveolus; I1a, first upper incisor alveolus; I2a, second upper incisor alveolus; I1a, third upper incisor alveolus; P3, third upper premolar. Scale bar = 1 cm. -
Marsupialia: Ektopodontidae): Including a New Species Ektopodon Litolophus
Records of the Western Australian Museum Supplement No. 57: 255-264 (1999). Additions to knowledge about ektopodontids (Marsupialia: Ektopodontidae): including a new species Ektopodon litolophus Neville S. Pledge!, Michael Archer, Suzanne J. Hand2and Henk Godthelp2 1 South Australian Museum, North Terrace, Adelaide, SA 5000; email: [email protected] 2 School of Biological Science, University of New South Wales, Sydney, NSW 2052 Abstract - Information about the extinct phalangeroid family Ektopodontidae has been increased following the discovery of new material from several localities. A new species, Ektopodon litolophus, described on the basis of an Ml from the Leaf Locality, Lake Ngapakaldi, Lake Eyre Basin, is characterized by the extremely simple structure of the crests. Ektopodontids are recorded for the first time from the northern half of the Australian continent through discovery of a tooth fragment at Wayne's Wok Site, Riversleigh World Heritage area, northwestern Queensland. Comparisons of Ml of Olllnia and Ektopodon species now allow evolutionary trends of simplification to be discerned. INTRODUCTION million years; Woodburne et al. 1985), following Ektopodon is a genus of extinct possum-like preliminary analyses by W.K. Harris of pollen from marsupials established by Stirton et al. (1967) on the Etadunna Formation at Mammalon Hill, Lake isolated teeth found at the Early to Middle Miocene Palankarinna. Subsequent work with Leaf Locality (Kutjamarpu Local Fauna) at Lake Ngapakaldi, northeastern South Australia (Figure 1). Further specimens from this locality were described and interpreted by Woodburne and Clemens (1986b), together with new, slightly older Oligocene species in the plesiomorphic genus CJmnia (c. illuminata, C. sp. cf. C. -
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13 STUDIES IN TASMANIAN MAMMALS, LIVING AND EXTINCT. Number I. Nototherium mitchelli. (A Marsupial Rhinoceros.) lierium. mitchelli, Owen, British Association for Advancement of Science, Report 1844, p. ?Zygomaturus trilobus, De Vis, Proceedings Royal S of Queensland. 1888, Vol. V., pt. .1 p. 111. By I!. 11. Scott (Curator, Launceston Museum); and Clive E. Lord (Curator, Tasmanian Museum). (Received 3rd May, L920. Read 10th May, 1920.) The discovery at Smithton, during the present year, <• of a nearly complete i Nototherium mitchelli forms the occasion for a revision of many of our ideas - - markable marsupial animals, since 1 1 * I ntarv remains hitherto available for study have 1 to yield the sequence of evidence we now possess, be only— inf nded to place upon record the hat. Nototherium mitchelli was an extinct marsupial rhino< I that the four genera, Nototherium, Zygo- maturus, Euowenia, and Sthenomerus, with their - species, are accordingly under revision— and will later on be dealt with in detail. ma > Material 1 in ibids anything like speculation ; . hut it is within the mark to observe thai two groups of these animals have be instinctively felt (by all workers) to have existed, quite irrespective of sex ques- tion- —one a platyrhine and the other a latifrons type, and that it now appears that they were also a horned, and a hornless group, and that Nototherium mitchelli belonged to the former, or cerathine group, and that some other species constituted the acerathine group, in which the 14 STUDIES IN TASMANIAN MAMMALS, LIVING AND EXTINCT, weapons were reduced to very small things, or actually missing. -
A New Family of Diprotodontian Marsupials from the Latest Oligocene of Australia and the Evolution of Wombats, Koalas, and Their Relatives (Vombatiformes) Robin M
www.nature.com/scientificreports OPEN A new family of diprotodontian marsupials from the latest Oligocene of Australia and the evolution of wombats, koalas, and their relatives (Vombatiformes) Robin M. D. Beck1,2 ✉ , Julien Louys3, Philippa Brewer4, Michael Archer2, Karen H. Black2 & Richard H. Tedford5,6 We describe the partial cranium and skeleton of a new diprotodontian marsupial from the late Oligocene (~26–25 Ma) Namba Formation of South Australia. This is one of the oldest Australian marsupial fossils known from an associated skeleton and it reveals previously unsuspected morphological diversity within Vombatiformes, the clade that includes wombats (Vombatidae), koalas (Phascolarctidae) and several extinct families. Several aspects of the skull and teeth of the new taxon, which we refer to a new family, are intermediate between members of the fossil family Wynyardiidae and wombats. Its postcranial skeleton exhibits features associated with scratch-digging, but it is unlikely to have been a true burrower. Body mass estimates based on postcranial dimensions range between 143 and 171 kg, suggesting that it was ~5 times larger than living wombats. Phylogenetic analysis based on 79 craniodental and 20 postcranial characters places the new taxon as sister to vombatids, with which it forms the superfamily Vombatoidea as defned here. It suggests that the highly derived vombatids evolved from wynyardiid-like ancestors, and that scratch-digging adaptations evolved in vombatoids prior to the appearance of the ever-growing (hypselodont) molars that are a characteristic feature of all post-Miocene vombatids. Ancestral state reconstructions on our preferred phylogeny suggest that bunolophodont molars are plesiomorphic for vombatiforms, with full lophodonty (characteristic of diprotodontoids) evolving from a selenodont morphology that was retained by phascolarctids and ilariids, and wynyardiids and vombatoids retaining an intermediate selenolophodont condition. -
The Chinchilla Local Fauna: an Exceptionally Rich and Well-Preserved Pliocene Vertebrate Assemblage from Fluviatile Deposits of South-Eastern Queensland, Australia
The Chinchilla Local Fauna: An exceptionally rich and well-preserved Pliocene vertebrate assemblage from fluviatile deposits of south-eastern Queensland, Australia JULIEN LOUYS and GILBERT J. PRICE Louys, J. and Price, G.J. 2015. The Chinchilla Local Fauna: An exceptionally rich and well-preserved Pliocene verte- brate assemblage from fluviatile deposits of south-eastern Queensland, Australia. Acta Palaeontologica Polonica 60 (3): 551–572. The Chinchilla Sand is a formally defined stratigraphic sequence of Pliocene fluviatile deposits that comprise interbed- ded clay, sand, and conglomerate located in the western Darling Downs, south-east Queensland, Australia. Vertebrate fossils from the deposits are referred to as the Chinchilla Local Fauna. Despite over a century and a half of collection and study, uncertainties concerning the taxa in the Chinchilla Local Fauna continue, largely from the absence of stratigraph- ically controlled excavations, lost or destroyed specimens, and poorly documented provenance data. Here we present a detailed and updated study of the vertebrate fauna from this site. The Pliocene vertebrate assemblage is represented by at least 63 taxa in 31 families. The Chinchilla Local Fauna is Australia’s largest, richest and best preserved Pliocene ver- tebrate locality, and is eminently suited for palaeoecological and palaeoenvironmental investigations of the late Pliocene. Key words: Mammalia, Marsupialia, Pliocene, Australia, Queensland, Darling Downs. Julien Louys [[email protected]], Department of Archaeology and Natural History, School of Culture, History, and Languages, ANU College of Asia and the Pacific, The Australian National University, Canberra, 0200, Australia. Gilbert J. Price [[email protected]], School of Earth Sciences, The University of Queensland, Brisbane, Queensland, 4072, Australia. -
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https://doi.org/10.24199/j.mmv.1978.39.06 7 July 1978 RAEMEOTHERIUM YATKOLAI, gen. et sp. nov., A PRIMITIVE DIPROTODONTID FROM THE MEDIAL MIOCENE OF SOUTH AUSTRALIA By Thomas H. Rich, 1 Michael Archer, 2 and Richard H. Tedford* 1 National Museum of Victoria. 2 Queensland Museum, Brisbane 3 American Museum of Natural History, New York Abstract A new genus and species, Raemeotherium yatkolai, from the medial Miocene Namba Formation in South. Australia, is the most primitive member of the Diprotodontidae yet des- cribed. On its most anterior lower molar is a well-developed paracristid and protocristid. This, together with a strong cristid obliqua on all lower molars suggests derivation of diprolodontids from selenodont rather than quadritubercular ancestors. Introduction NMV National Museum of Victoria, Melbourne. SAM South Australian Museum, Adelaide. In September 1976, Mr Ian Stewart (Na- tional Museum of Victoria) discovered the di- Terminology and Measurements protodontid dentary here described as Rae- Mandibular terminology follows Stirton meotherium yatkolai (SAM P19764). It was (1967), dental terminology is modified from found at the south end of Lake Pinpa, South Archer (1976) (see fig. 1), and enumeration of Australia, in the medial Miocene Namba For- the teeth follows Archer (in press) (see fig. 1, mation. Only one or two centimetres away at 2). The system of tooth enumeration used here the same level was a mandible of a species of departs radically from all previously proposed. pseudocheirine similar generically to that Ted- In it, the permanent cheek teeth of diproto- ford et al. (1976, p. 56) characterized as dontids are P3 M2, M3, M4, M5. -
Systematic and Palaeobiological Implications of Postcranial Morphology in the Diprotodontidae (Marsupialia)
Systematic and palaeobiological implications of postcranial morphology in the Diprotodontidae (Marsupialia) Aaron B. Camens School of Earth and Environmental Sciences Discipline of Ecology and Evolutionary Biology The University of Adelaide South Australia A thesis submitted in partial fulfilment of the degree of Doctor of Philosophy at the University of Adelaide February 2010 II Declaration This work contains no material which has been accepted for the award of any other degree or diploma in any university or other tertiary institution to Aaron Camens and, to the best of my knowledge and belief, contains no material previously published or written by another person, except where due reference has been made in the text. I give consent to this copy of my thesis when deposited in the University Library, being made available for loan and photocopying, subject to the provisions of the Copyright Act 1968. The author acknowledges that copyright of published works contained within this thesis (as listed below) resides with the copyright holder(s) of those works. I also give permission for the digital version of my thesis to be made available on the web, via the University’s digital research repository, the Library catalogue, the Australasian Digital Theses Program (ADTP) and also through web search engines, unless permission has been granted by the University to restrict access for a period of time. Publications in this thesis include: Camens, A. B. and Wells, R.T. 2009. Diprotodontid footprints from the Pliocene of Central Australia. Journal of Vertebrate Paleontology 29: 863-869. Copyright held by Taylor and Francis. Camens, A. B. and Wells, R.T. -
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https://doi.org/10.24199/j.mmv.1973.34.03 9 May 1973 FOSSIL VERTEBRATE FAUNAS FROM THE LAKE VICTORIA REGION, S.W. NEW SOUTH WALES, AUSTRALIA By Larry G. Marshall Zoology Department, Monash University, Victoria* Abstract Fossil vertebrate localities and faunas in the Lake Victoria region of S.W. New South Wales, Australia, are described. The oldest fossil bearing deposit, the late Pliocene or early Pleistocene Moorna Formation and associated Chowilla Sand have yielded specimens of Neoceratodus sp., Emydura macquarrii, several species of small dasyurid, specimens of Glaucodon cf. G. ballaratensis, a species of Protemnodon which compares closely with P. cf. P. otibandus from the late Pliocene or early Pleistocene Chinchilla Sand in SE. Queensland, specimens of Lagostrophus cf L. fasciatus, species of Petrogale, Macropus, Osphranter, Sthenurus, Bettongia, Diprotodon, Lasiorhinus, a peramelid, at least two species of pseudomyine rodents and a species of Rattus cf. R. lutreolus. It is shown that the holotype of Zygomaturus victoriae (Owen) 1872 may have been collected from these sediments. The late Pliocene or early Pleistocene Blanchetown Clay has yielded species of Neoceratodus, Thylacoleo, Phas- colonus, Bettongia, Sthenurus, a diprotodontid, and a rodent. The late Pleistocene Rufus Formation has yielded species of Dasycercus, Sarcophilus, Thylacinus, Phascolonus, Lasiorhinus, Bettongia, Procoptodon, Onychogalea, Macropus, and Leporillus. A large species of macropod and a species of Phascolonus were collected from the late Pleistocene Monoman Formation. The lunette on the E. side of Lake Victoria has yielded a large, diverse fauna of late Pleistocene —Holocene age, that includes such extinct species as Protemnodon anak, P. brehus, Procoptodon goliah, Sthenurus andersoni, S. -
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Memoirs of the National Museum of Victoria 9 April 1969 https://doi.org/10.24199/j.mmv.1969.29.03 A LOWER MANDIBLE OF ZYGOMATURUS GILLI FROM THE SANDR1NGHAM SANDS, BEAUMARIS, VICTORIA, AUSTRALIA By Michael O. Woodburne Department of Geological Sciences, University of California, Riverside, California, U.S.A. Introduction Stirton (1957) described three diprotodontid specimens which had been recov- ered from the Black Rock Sandstone of the Brighton Group (Kcnley 1967), on the P°rt PhilliP Bay near Beaumaris, Victoria. One of the specimens, M.U.G.D.S^rV^ No. 2020, subsequently became the type of Zygomaturus gilli (Stirton 1967) and the second, N.M.V. PI 5905, was also assigned to that species (ibid.). The third specimen, a fragment with the symphysis and part of the horizontal ramus containing the alveoli of P , base 3 of M u and most of M 2 , was also discussed by Stirton (1957), but at that time information on Tertiary diprotodontids was insufficient for a close determination of its affinities. In 1967, Mr Colin Macrae found the rear portion of a mandible containing Ma and M4 which proved to fit perfectly onto the former specimen. With this surprising discovery, a nearly com- plete mandible is represented, lacking only the angle, coronoid process, and anterior tip of the symphysis. Now that this individual is more completely represented, it is possible to compile a sufficiently adequate description to allow an analysis of its affinities. Such is the purpose of the present report. The specimen originally des- cribed by Stirton is catalogued as N.M.V. -
Kenny TRAVOUILLON Phd Thesis Complete
PALAEOECOLOGICAL AND BIOCHRONOLOGICAL STUDIES OF RIVERSLEIGH, WORLD HERITAGE PROPERTY, OLIGO-MIOCENE FOSSIL LOCALITIES, NORTH-WESTERN QUEENSLAND, AUSTRALIA Kenny J mes TRAVOUILLON Thesis submitted for the degree of Doctor of Philoso.hy t the University of Ne0 South W les, Austr li August, 2008 i N1 d2ordre Ann3e 2008 THESE .r3sent3e devant l2UNIVERSITE CLAUDE BERNARD - LYON 1 .our l2obtention du DIPLOME DE DOCTORAT 8 rr9t3 du 7 o;t 2006 et rr9t3 du 6 j nvier 2005) Souten nce : Novembre 2008 . r M. Kenny J mes TRAVOUILLON TITRE : ETUDES PALEOECOLOGIQUES ET BIOCHRONOLOGIQUES DES GISEMENTS FOSSILIFERES DE L2OLIGO-MIOCENE DE RIVERSLEIGH, WORLD HERITAGE PROPERTY, NORD-OUEST DU QUEENSLAND, AUSTRALIE Directeur de thBse : Dr Serge Legendre Prof Mich el Archer Dr SuC nne H nd JURY : Dr. Serge Legendre Directeur de ThBse Pr. Mich el Archer Directeur de ThBse Pr. Christo.he Lecuyer ED min teur Dr. Gilles Esc rguel ED min teur Dr. So.hie Montuire R ..orteur Pr. Phili. Gingerich R ..orteur Dr. M nuel Hern ndeC Fern ndeC Ra..orteur ii THE UNIVERSITY OF NEW SOUTH WALES ThesisEDissert tion Sheet Surname or Family name: Tr vouillon First name: Kenny Other name/s: J mes Abbreviation for degree as given in the University calendar: PhD School: BEES Faculty: Science itle: PALAEOECOLOGICAL AND BIOCHRONOLOGICAL STUDIES OF RIVERSLEIGH, WORLD HERITAGE PROPERTY, OLIGO-MIOCENE FOSSIL LOCALITIES, NORTH-WESTERN QUEENSLAND, AUSTRALIA Abstr ct 350 0ords m Dimum: Riversleigh, World Heritage Property, located in North-western Queensland, Australia, contains over 200 fossil bearing localities from the Oligo-Miocene. The study presented here aims at finding new methods to improve the accuracy of palaeoecological and biochronological studies and describe the palaeoenvironmental and chronological settings of the Riversleigh fossil deposits. -
A Survey of Cenozoic Mammal Baramins
The Proceedings of the International Conference on Creationism Volume 8 Print Reference: Pages 217-221 Article 43 2018 A Survey of Cenozoic Mammal Baramins C Thompson Core Academy of Science Todd Charles Wood Core Academy of Science Follow this and additional works at: https://digitalcommons.cedarville.edu/icc_proceedings DigitalCommons@Cedarville provides a publication platform for fully open access journals, which means that all articles are available on the Internet to all users immediately upon publication. However, the opinions and sentiments expressed by the authors of articles published in our journals do not necessarily indicate the endorsement or reflect the views of DigitalCommons@Cedarville, the Centennial Library, or Cedarville University and its employees. The authors are solely responsible for the content of their work. Please address questions to [email protected]. Browse the contents of this volume of The Proceedings of the International Conference on Creationism. Recommended Citation Thompson, C., and T.C. Wood. 2018. A survey of Cenozic mammal baramins. In Proceedings of the Eighth International Conference on Creationism, ed. J.H. Whitmore, pp. 217–221. Pittsburgh, Pennsylvania: Creation Science Fellowship. Thompson, C., and T.C. Wood. 2018. A survey of Cenozoic mammal baramins. In Proceedings of the Eighth International Conference on Creationism, ed. J.H. Whitmore, pp. 217–221, A1-A83 (appendix). Pittsburgh, Pennsylvania: Creation Science Fellowship. A SURVEY OF CENOZOIC MAMMAL BARAMINS C. Thompson, Core Academy of Science, P.O. Box 1076, Dayton, TN 37321, [email protected] Todd Charles Wood, Core Academy of Science, P.O. Box 1076, Dayton, TN 37321, [email protected] ABSTRACT To expand the sample of statistical baraminology studies, we identified 80 datasets sampled from 29 mammalian orders, from which we performed 82 separate analyses. -
A Tiny New Marsupial Lion (Marsupialia, Thylacoleonidae) from the Early Miocene of Australia
Palaeontologia Electronica palaeo-electronica.org A tiny new marsupial lion (Marsupialia, Thylacoleonidae) from the early Miocene of Australia Anna K. Gillespie, Michael Archer, and Suzanne J. Hand ABSTRACT Microleo attenboroughi, a new genus and species of diminutive marsupial lion (Marsupialia: Thylacoleonidae), is described from early Miocene freshwater limestones in the Riversleigh World Heritage Area, northwestern Queensland, Australia. A broken palate that retains incomplete cheektooth rows demonstrates that this new, very small marsupial lion possessed the elongate, trenchant P3 and predominantly subtriangular upper molars characteristic of thylacoleonids, while other features of the premolar sup- port its placement in a new genus. Phylogenetic analysis suggests that Microleo atten- boroughi is the sister taxon to all other thylacoleonids, and that Thylacoleonidae may lie outside Vombatomorphia as the sister taxon of all other wombat-like marsupials including koalas. However, given limited data about the cranial morphology of M. atten- boroughi, Thylacoleonidae is concluded here, conservatively, to be part of the vom- batomorphian clade. This new thylacoleonid brings to three the number of marsupial lion species that have been recovered from early Miocene deposits at Riversleigh and indicates a level of diversity previously not seen for this group. It is likely that the differ- ent size and morphology of the three sympatric taxa reflects niche partitioning and hence reduced competition. Thylacoleonids may have been the dominant arboreal predators of Cenozoic Australia. Anna K. Gillespie*, Palaeontology, Geobiology and Earth Archives (PANGEA) Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney 2052, Australia; [email protected], *corresponding author Michael Archer, Palaeontology, Geobiology and Earth Archives (PANGEA) Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney 2052, Australia; [email protected] Suzanne J.