The Journal of Threatened Taxa (JoTT) is dedicated to building evidence for conservaton globally by publishing peer-reviewed artcles OPEN ACCESS online every month at a reasonably rapid rate at www.threatenedtaxa.org. All artcles published in JoTT are registered under Creatve Commons Atributon 4.0 Internatonal License unless otherwise mentoned. JoTT allows unrestricted use, reproducton, and distributon of artcles in any medium by providing adequate credit to the author(s) and the source of publicaton.
Journal of Threatened Taxa Building evidence for conservaton globally www.threatenedtaxa.org ISSN 0974-7907 (Online) | ISSN 0974-7893 (Print) Communication
Mantids (Insecta: Mantodea) of Uttar Pradesh, India
Ramesh Singh Yadav & G.P. Painkra
26 May 2021 | Vol. 13 | No. 6 | Pages: 18578–18587 DOI: 10.11609/jot.5958.13.6.18578-18587
For Focus, Scope, Aims, and Policies, visit htps://threatenedtaxa.org/index.php/JoTT/aims_scope For Artcle Submission Guidelines, visit htps://threatenedtaxa.org/index.php/JoTT/about/submissions For Policies against Scientfc Misconduct, visit htps://threatenedtaxa.org/index.php/JoTT/policies_various For reprints, contact
The opinions expressed by the authors do not refect the views of the Journal of Threatened Taxa, Wildlife Informaton Liaison Development Society, Zoo Outreach Organizaton, or any of the partners. The journal, the publisher, the host, and the part- Publisher & Host ners are not responsible for the accuracy of the politcal boundaries shown in the maps by the authors.
Member
Threatened Taxa Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 ISSN 0974-7907 (Online) | ISSN 0974-7893 (Print) OPEN ACCESS htps://doi.org/10.11609/jot.5958.13.6.18578-18587
#5958 | Received 12 April 2020 | Final received 07 March 2021 | Finally accepted 30 March 2021
COMMUNICATION
Mantds (Insecta: Mantodea) of Utar Pradesh, India
Ramesh Singh Yadav 1 & G.P. Painkra 2
1 Government School Dehariya, Zamania, Ghazipur, Utar Pradesh 232331, India. 2 Hindi: 2017Raj Mohini2019 Devi College of Agriculture & Research Staton,, Ambikapur, Chhatsgarh, 497001, India. सन से 1 [email protected]क% अविध के दौरान पूव0(correspondingउ2र 3देश author),भारत 2 [email protected]म8 म9:टड जीव? के िलए कई सवCDण :कए गए थे। सवCDण के दौरान एकIJत सामिKय? म8 छः कु ल ? और नौ उ प कु ल ? से 17 वंश? क% कु ल 21 3जाितयां 3ाR हुई ह9। मैनटUडU कु ल म8 सबसे अिधक संWया म8 म9:टड जोव? ने योगदान :दया। कु ल िमलाकर, वतYमान Abstract: Several surveys of mantd fauna from eastern Utar Pradesh, India were carried out during the period 2017–2019. The collected materialsउ2र-3देश include(राZय 21 speciesके Iवभाजन of 17 generaके fromबाद six) familiesराZय andसे पहलीnine subfamilies.बार म8:टस The Familyक% 10 Mantdae नई 3जाितयां contributed[रकॉडY the highestक% numbersगई ह9, ofजब:क mantdटेनोडेरा fauna. In को]टेिलस total, 10 species(^लां ofका mantsडY, 1853) are reportedदेश म8 forपहली the frstबार tmeउ2र from-3देश the presentसे दजY Utarक% Pradeshगई है। (afer division of the state) while Tenodera costalis (Blanchard, 1853) is recorded for the frst tme from the country.
Keywords: Insecta, Mantdae, new country record.
Hindi: सन 2017 से 2019 क% अविध के दौरान पूव0 उ2र 3देश, भारत म8 म9:टड जीव? के िलए कई सवCDण :कए गए थे। सवCDण के दौरान एकIJत सामिKय? म8 छः कु ल ? और नौ उ प कु ल ? से 17 वंश? क% कु ल 21 3जाितयां 3ाR हुई ह9। मैनटUडU कु ल म8 सबसे अिधक संWया म8 म9:टड जोव? ने योगदान :दया। कु ल िमलाकर, वतYमान उ2र-3देश (राZय के Iवभाजन के बाद) राZय से पहली बार म8:टस क% 10 नई 3जाितयां [रकॉडY क% गई ह9, जब:क टे नोडेरा को]टेिलस (^लांकाडY, 1853) देश म8 पहली बार उ2र-3देश से दजY क% गई है।
Hindi: सन 2017 से 2019 क% अविध के दौरान पूव0 उ2र 3देश, भारत म8 म9:टड जीव? के िलए कई सवCDण :कए गए थे। सवCDण के दौरान एकIJत सामिKय? म8 छः कु ल ? और नौ उपकुल? से 17 वंश? क% कु ल 21 3जाितयां 3ाR हुई ह9। मैनटUडU कु ल म8 सबसे अिधक संWया म8 म9:टड जोव? ने योगदान :दया। कु ल िमलाकर, वतYमान उ2र-3देश (राZय के Iवभाजन के बाद) राZय से पहली बार म8:टस क% 10 नई 3जाितयां [रकॉडY क% गई ह9, जब:क टेनोडेरा को]टेिलस (^लांकाडY, 1853) देश म8 पहली बार उ2र-3देश से दजY क% गई है। Editor: Anonymity requested. Date of publicaton: 26 May 2021 (online & print)
Citaton:Yadav, R.S. & G.P. Painkra (2021). Mantds (Insecta: Mantodea) of Utar Pradesh, India. Journal of Threatened Taxa 13(6): 18578–18587. htps://doi. org/10.11609/jot.5958.13.6.18578-18587
Copyright: © Yadav & Painkra 2021. Creatve Commons Atributon 4.0 Internatonal License. JoTT allows unrestricted use, reproducton, and distributon of this artcle in any medium by providing adequate credit to the author(s) and the source of publicaton.
Funding: None.
Competng interests: The authors declare no competng interests.
Author details: Ramesh Singh Yadav is an independent researcher cum educator in the Government School Dehariya, Zamania, Ghazipur, U.P., India. Presently he involved in teaching as well as research too. He actvely partcipate in collaboratve work for wildlife from diferent states along with various other community services. He has published more than 20 artcles in various journals of natonal and internatonal reputes with several new records and authored fve books on entomology/zoology. His areas of research interest include ecology and systematc zoology. G.P. Painkra is a senior scientst of Entomology, Department of Entomology at IGKVV-RMD College of Agriculture and Research Staton, Ambikapur, Chhatsgarh, India. He advised 10 MSc and one PhD students. He recognized as a loving scientst in farmer community due to his excellent research in apiculture. He published two books and 79 research papers in reputed journals.
Author contributons: RSY involved in survey, fauna collecton, captured photographs and identfcaton as well as write up of the manuscript. GPP partcipated in design of the research, supervision of research actvites and overview the manuscript.
Acknowledgements: The frst author wishes to extend his grattude to Dr. H.V. Ghate, former head, Department of Zoology, Modern College, Shivajinagar, Pune and Dr. T.K. Mukherjee, associate professor (retred), Presidency University, Kolkata for identfcaton of certain mantd species.
18578 Mantds of Utar Pradesh, India Yadav & Painkra J TT INTRODUCTION Table 1. Coordinates of the survey sites.
Site Coordinates
Mantds are one of the important groups of predatory 1 Karmahari, Ghazipur 25.35140N & 82.64810E insects of diferent agroecosystems. Mantds are also 2 Chahaniya, Chandauli 25.41890N & 83.21180E called preying mantds due to their preying ability. They 3 Chakka bandh Dam, Ghazipur 25.42040N & 83.55740E are sometme confused with the mantsfies of the order 4 Rohuna, Ghazipur 25.41590N & 83.55980E Neuroptera. Mantds are grouped in the superorder 0 0 Dictyoptera because of their more closeness to cockroaches 5 Maujhi, Chandauli 25.4255 N & 83.5580 E and termites while forelegs of mantds are raptorial type 6 Mohammadabad, Ghazipur 25..61680N & 83.75310E which is not identcal to the forelegs of cockroaches and 7 Dehariya, Ghazipur 25.29910N & 83.65620E termites. Globally, 2,384 species under 434 genera of 15 families are known (Ehrmann & Roy 2002) while Ote et al (2020) listed 2,400 species from 430 genera; in which more MATERIALS AND METHODS than 169 species, 71 genera, and 11 families are known from India of which 60 species are endemic to the country The present exploraton was made from a series of (Mukherjee et al. 2014). Mantds are a key agent of natural surveys of the several locatons in Utar Pradesh, India from and biological control in most of the terrestrial ecosystems 2017 to 2019. The study sites were located in diferent due to their sophistcated behaviour and modifcatons. areas of eastern Utar Pradesh (Table 1). The mantds They are easily identfed by their raptorial forelegs, were collected by sweeping net, light trap, and hand- triangular head, visible eyes, and large cerci. picking methods. Afer photography, most of the mantds Probably, frst taxonomic work on mantds was initated were released to avoid the loss of biodiversity issue. The by Linnaeus (1758) while in India, remarkable contributon collectons were made during morning and night hours, to mantds came in light when the Wood-Mason joined i.e., 07.00–10.00 h and 20.00–22.00 h, respectvely. Some as director of Indian Museum, Kolkata (Yadav 2017) and of the collected materials were killed in the killing botle published a Catalogue of the Mantodea of Indian Museum containing ethyle acetate. Subsequently, materials were collectons by Wood-Mason (1889 & 1891). Subsequently cleaned with hair brush, pinned and relaxed (as per need) Bolivar (1899) and Werner (1931, 1933 & 1935) contributed on the handmade stretching board of thermocol. It was to the Indian mantds. Kirby (1904) has created a synonimic kept for more than 60–72 hours for drying to avoid spoilage catalogue of the 82 species of mantds from India whereas, of the specimens. The identfcaton was made based on Nadkerni (1965) amassed 55 more mantds. Some notable the morphological and genitalic features. The specimens works on the taxonomy and distributon of mantds were collected were deposited in the Department of Agricultural done by Mukherjee & Hazra (1983), Mukherjee et al. (1992, Entomology, R.M.D. College and Research Staton, 1995, 2014, 2015, 2017), Mukherjee & Shisodia (1999 Ambikapur, Chhatsgarh, India. The classifcatons were & 2000), Chaturvedi & Hedge (2000), Rao et al. (2005), made according to Mukherjee et al. (2014). The coordinates Sureshan et al. (2006), Jadhav et al. (2006), Mukherjee & of the survey sites are presented in Table 1. Hazra (2007), Vyjayandi (2007), Jadhav (2008), Chandra (2009), Sureshan (2009), Sureshan & Sambath (2009), Mukherjee & Ghate (2010), Vyjayandi et al. (2010), Ghate RESULTS et al. (2012), and Chaterjee & Mukherjee (2013). The most remarkable work on Indian mantds was presented Biodiversity of surveyed mantds from Utar Pradesh are by Mukherjee et al. (1995) which was more consolidated presented here in fgure 1 and their checklist is as below. account on this group tll date. Utar Pradesh has remained less studied for Mantodea. More recently Yadav et al (2018) Systematc checklist of the mantds studied from Utar have recorded some mantds from this state. More diversity Pradesh of the mantds remain unexplored. Therefore, we explored Systematc account the mantd fauna extensively from Utar Pradesh afer the Class: Insecta division of Utarakhand. Order: Mantodea Latreille, 1802 A. Family: Hymenopodidae Giglio-Tos, 1915 Subfamily: Acromantnae Giglio-Tos, 1915 Tribe: Anaxarchini Giglio-Tos, 1919 1. Euantssa pulchra Fabricius, 1787*
Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 18579 J TT Mantds of Utar Pradesh, India Yadav & Painkra 2. Odontomants montana Giglio-Tos, 1915* Tribe: Hymenpodini Giglio-Tos, 1915 3. Creobroter apicalis Saussure, 1869* Tribe: Acromantni Runner De Watenwyl, 1893 4. Ephestasula pictpes (Wood-Mason, 1879)** 5. Hestasula brunneriana Saussure, 1871*
B. Family: Liturgusidae Giglio-Tos, 1915 Subfamily: Listurgusinae Giglio-Tos, 1915 Tribe: Humbertellini Brunner De Watenwyl, 1893 6. Humbertella ceylonica Saussure, 1869 7. Humbertella similis Giglio-Tos, 1917 Figure 1. Biodiversity of mantds of Utar Pradesh. Relatve abundance (in percentage) C. Family: Tarachodidae Handlirsch, 1930 Subfamily: Schizocephalinae Saussure, 1869 8. Schizocephala bicornis (Linné,̇ 1758)** :541. 2002. Euantssa pulchra Ehrmann, Mantodea der Welt D. Family: Mantdae Latreille,̇ 1802 244. Subfamily: Amelinae Westwood, 1889 2014. Euantssa pulchra Mukherjee et al. Priamus 30: 7. 9. Amants saussurei (Bolivar, 1897)* Materials examined: RMDCRS 001-002, 1 male, 1 10. Elmants trincomaliae (Saussure, 1869)* female, 21.x.2017, Udharanpur (Ghazipur) Utar Pradesh, Subfamily: Tenoderinae Brunner De Watenwyl, 1893 India; on Theveta peruviana, coll. R.S. Yadav. 11. Tenodera fasciata (Oliver,1792)* Diagnosis: Vertex without protuberance; eyes oblong; 12. Tenodera costalis (Blanchard, 1853) ¶ pronotum broad, fore femur simple with four sharp- 13. Hierodula coarctata Saussure, 1869 long external and four discoidal spines; mid and hind legs Subfamily: Mantnae Burmeister, 1838 have without lobes; wings brightly coloured, longer than 14. Mants religiosa religiosa Linnaeus, 1758** abdomen, with dark line along the outer border (Vyjayandi 15. Statlia maculata contnentalis Werner, 1935 2007). 16. Statlia nemoralis (Saussure, 1870) Distributon: India (Andhra Pradesh, Tamil Nadu, West Bengal, & Utar Pradesh (new record)), China, and Sri Lanka E. Family: Toxoderidae Saussure, 1869 (Mukherjee et al. 2014). Subfamily: Oxyothespinae Gigli̇ o-Tos,̇ 1916 Measurement: Body length 25mm 17. Cheddikulama straminea Henry, 1932** Remark: The specie is more common in bushes but not Subfamily: Toxoderinae Saussure, 1869 frequent in my collecton. 18. Aethalochroa ashmoliana (Westwood, 1841)* 19. Toxoderopsis taurus Wood-mason, 1889* Odontomants montana Giglio-Tos, 1915 (Image 2 & 3) 1915. Odontomants montana Giglio-Tos. Bull. Soc. Ent. F. Family Empusidae Burmeister, 1838; Ital. 46: 100. Subfamily Empusinae Burmeister, 1838 2002. Odontomants montana Ehrmann. Mantodea der 20. Empusa (Empusa) gutula (Thunberg, 1815) Welt 244. 21. Empusa spinosa Krauss, 1902 * 2014. Odontomants montana Mukherjee et al. Priamus Note: Asterisk marks (*), (**) and (¶) indicatons are 30: 8. indicated as species frstly recorded from Utar Pradesh, Materials examined: RMDCRS 003, 1female, 09.ix.2017, probably earlier recorded from Utar Pradesh and species Chakka bandh Dam (Ghazipur) Utar Pradesh, India; on frst tme recorded from country respectvely. broad leaved shrub, coll. R.S. Yadav. Diagnosis: Vertex without protuberance; frontal sclerite Family: Hymenopodidae Giglio-Tos, 1915 narrowed in the form of a groove, superior border wavy, Subfamily: Hymenopodinae, Giglio-Tos, 1915 a litle arched in the middle; inferior border with a slightly Tribe: Anaxarchini Giglio-Tos, 1919 elevated ridge; margin of pronotum bearing small tubercular Euantssa pulchra Fabricius, 1787 (Image 1) spines. In forelegs external spines of femora blackish near 1927. Euantssa pulchra Gigilio-Tos, Das Tierriech 50 their tps and proximal two of them closer, tbiae with 10
18580 Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 Mantds of Utar Pradesh, India Yadav & Painkra J TT internal spines of same colour (Sureshan et al. 2006). Java, Nepal, and Bhutan (Mukherjee et al. 2014, 2017). Distributon: India (Odisha & Utar Pradesh (new Measurement: Body length 34mm record)), Malayasia, and Sumatra (Mukherjee et al. 2014). Remark: The species was very atractve and colourful. Measurement: Immature mantd (nymph) Remark: The species was rare in the collecton. Tribe: Acromantni Runner De Watenwyl, 1893 Ephestasula pictpes (Wood-Mason, 1879) (Image 5) Tribe: Hymenpodini Giglio-Tos, 1915 1879. Hestas pitpes Wood-Mason, Proc. Asiatc Soc. Creobroter apicalis Saussure, 1869 (Image 4) Bengal: 258. 1869. Creobroter apicalis Saussure Mill. Schweiz. 1951. Ephestasula pictpes Roonwal & Bhasin, Indian Entomol. Ges., 3: 73. Forester, 77(5): 313-318. 1927. Creobroter apicalis Giglio-Tos Das Tierreich., 50: 1995. Ephestasula pictpes Mukherjee et al., Oriental 558. Insects 29(1): 217. 2014. Creobroter apicalis: Mukherjee et al., Priamus, 2014. Ephestasula pictpes Mukherjee et al., Priamus, 30: 9. 30: 12. 2017. Creobroter apicalis Mukherjee, Iyer & Chaterjee 2015. Ephestasula pictpes Majumder et al. Zool. Surv. J. Threat. Taxa, 9(2): 9832. India, 115(4): 368 Materials examined: RMDCRS 004-005, 2 female, Materials examined: RMDCRS 006, 1 male, 23.ix.2017, 16.x.2017, Udharanpur (Ghazipur) Utar Pradesh, India; on Karmahari (Ghazipur) Utar Pradesh, India; on bushes, coll. Theveta peruviana, coll. R.S. Yadav. R.S. Yadav. Diagnosis characters: Female: Body medium to large, Diagnosis: External edge of fore femur dentculated, body greenish with yellow spot on the fore wing; winged internal face of forefemur spines black, inferior internal forms; head trapezoidal, vertex of head with a small mid lobes contain three yellow to white spots encased by black dorsal spine, squarish with prominent lateral lobes; eyes patch; black patch absents at upper border and very thin conical, black band with white tps, projectng upwards; below near base; tarsi internally black; pronotum dorso- antennae slender, fliform; ocelli conspicuous; pronotum medially black, saddle shape, shorter than fore coxae, supra coxal dialaton Distributon: India (Chhatsgarh, Madhya Pradesh, well pronounced, prozona laterally dentculated, spatulate Maharashtra, Odisha, Utar Pradesh (earlier record from with central indistnct carina, metazonal constricton Dehradun, Utarakhand) (Mukherjee et al. 1995, 2014), well pronounced; Coxae of fore legs triangular dorsally Utar Pradesh, (new record)) and Nepal. with eight obtuse marginal spines, middorsal carina with Measurement: Body length 34mm spines, internal apical lobes converging; femur longer than Remark: The species was brown in colour. coxa, with four external, four discoidal, (third longest and stoutest,) with six longer internal and seven shorter internal Hestasula brunneriana Saussure, 1871 (Image 6 & 7) spines, two distal longer internal spines enclose two shorter 1871. Hestasula brunneriana Saussure, Mem. Soc. spines; tbia with 16 smaller, depressed external spines, Phys. Hist. Nat. Geneve, 21: 330. 14 longer internal closely arranged spines; metatarsus as 2001. Hestasula brunneriana Ghate, Ranade, Kaur and long as all other tarsal segments together; middle and hind Marathe, J. Bombay femur twice as long as mid tbia, with semi-circular, distal Nat. Hist. Soc. 98 (3): 473-476. ventral lobe, with two genicular spines; tbia with three 2007. Hestasula brunneriana Vyjayandi. Zool. Surv. distal genicular spines; wings longer than abdomen; costal India, Occ. Paper .267: 39. and anal areas of forewing transparent, forewing with 2014. Hestasula brunneriana Mukherjee et al., yellow band bordered by two black semi-circular rings like Priamus, 30: 12. an eye spot and placed in the middle of the both wings, this Materials examined: RMDCRS 007-009, 1 male, 2 eye mark enclosed black dots, usually two; base of forewing female, 09.ix.2017, Chakka bandh Dam (Ghazipur) Utar with a yellow spot also, the hind wing with pink at base, Pradesh, India, on white colour fowering bushes, coll. R.S. yellowish in costal area and brownish in discoidal and anal Yadav. areas. Diagnosis: Vertex with protuberance; fore femur Distributon: India (Andhra Pradesh, Arunachal Pradesh, arched, foliaceous, external edge smooth with three black Assam, Karnataka, Kerala, Madhya Pradesh, Maharashtra, spots on superior margin and one near the spine in middle Manipur, Meghalaya, Odisha, Sikkim, Tamil Nadu, West Mukherjee et al. 1995; Ghate et al. 2001; Vyjayandi 2007). Bengal, & Utar Pradesh (new record)), Bangladesh, China, Distributon: India (Andhra Pradesh, Bihar, Maharashtra,
Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 18581 J TT Mantds of Utar Pradesh, India Yadav & Painkra Meghalaya, Odisha, West Bengal, & Utar Pradesh (New black at tps only. Irregular veinules in the costal area of fore record)), Pakistan, and Sri Lanka (Mukherjee et al. 2014). wing, wings smoky and longer than body. Measurement: Body length 29mm Distributon: India (Himachal Pradesh, Jammu & Remark: Inner side of fore femur is orange in colour and Kashmir, Kerala, Madhya Pradesh, Odisha, & Utar Pradesh), ornamented with three inner black spots. Nepal, and Sri Lanka (Mukherjee et al. 2014). Measurement: Body length 37mm Family: Liturgusidae Giglio-Tos, 1915; Subfamily: Remark: Brown coloured species. Listurgusinae Giglio-Tos, 1915 Humbertella ceylonica Saussure, 1869 (Image 8) Family: Tarachodidae Handlirsch, 1930; Subfamily: 1869. Humbertella ceylonica Saussure Mit. Schweiz. Schizocephalinae Saussure, 1869 Entomol. Ges.,3:62. Schizocephala bicornis (Linné,̇ 1758) (Image 10&11) 1891. Theopompa sepentrionum Wood-Mason. A 1927. Schizocephala bicornis Giglio-Tos, Das Tierreich, catalogue of Mantblac 2:61. 50: 237. 2007. Humbertella ceylonica Vyjayandi. Zool. Surv. 2007. Schizocephala bicornis Vyjayandi. Zool. Surv. India, India, Occ. Paper .267: 73. Occ. Paper .267: 121. 2014. Humbertella ceylonica Mukherjee et al., Priamus, 2014. Schizocephala bicornis Mukherjee et al., Priamus, 30: 14. 30: 16. Materials examined: RMDCRS 0010-012, 3 female, Materials examined: RMDCRS 0016, 1 female, 09.vi.2017, Zamania (Ghazipur) Utar Pradesh, India, on 07.vii.2018, Dehariya (Ghazipur), Utar Pradesh, India, on light trap, coll. R.S. Yadav. grasses, coll. R.S. Yadav. Diagnosis: Small black species with triangular head Diagnosis: Long and slender mantd, green coloured. having fve lobes in vertex and blackish frontal sclerite along Antenna thickened toward base; eyes anteriorly prolonged with contnuous black band; pronotum short and rhomboid forming cone shape. Fore femur with four external and in shape; fore femur slightly bulged, externally 4 discoidal three discoidal in which second is longest one; tbia and internally 5 large (completely black) and 5 short spines shortened with six external spines. Forewing very small and with three blackish band on inner side of femur as well as on opaque in female (Majumder et al. 2015). tbia black; anal vein of fore wing three branched in female. Distributon: India (Andhra Pradesh, Bihar, Chhatsgarh Distributon: India (Andhra Pradesh, Assam, Bihar, (Bilaspur Korba, Raipur, Koriya), Kerala, Madhya Pradesh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Maharashtra, Odisha, Tamil Nadu, West Bengal, & Utar Tamil Nadu, Utar Pradesh, West Bengal, & Utar Pradesh Pradesh (new record)), Africa, Nepal, Thailand, Sri Lanka, (new record)), Myanmar, Nepal, and Sri Lanka (Mukherjee and Sunda Island. et al. 2014). Measurement: Body length 145mm Measurement: Body length 34mm Remark: Largest mantd in the collecton looks like a Remark: Wood coloured black species. stck insect.
Humbertella similis Giglio-Tos, 1917 (Image 9) Family: Mantdae Latreille,̇ 1802, Subfamily: Amelinae 1897. Humbertella indica Bolivar Ann. Soc. Enlolnol. Westwood, 1889 France, 66: 303. Amants saussurei (Bolivar, 1897) (Image 12) 2007. Humbertella similis Vyjayandi. Zool. Surv. India, 1927. Amants saussurei Giglio-Tos, Das Tierreich, 50: Occ. Paper .267: 70 171. 1917. Humbertella similis GigIio-tos Bull. Soc. Entomol. 2014. Amants saussurei Mukherjee et al. Priamus, 30: Ital., 48: 83. 25. 2014. Humbertella similis Mukherjee et al., Priamus, 2015. Amants saussurei Majumder et al. Zool. Surv. 30: 15. India, 115(4): 376. Materials examined: RMDCRS 013-015, 1 male, 2 Materials examined: RMDCRS 0017-0018, 2 female, female, 23.vi.2018, Chahaniya, (Chandauli), Utar Pradesh, 24.iii.2018, Maujhi (Chandauli), Utar Pradesh, India, on India, on light trap, coll. R.S. Yadav. grasses, coll. R.S. Yadav. Diagnosis: Small brown species with frontal sclerite Diagnosis: Small in size, brown coloured mants similar brown with less arched superior edges; pronotum with less to bark with scatered black spots on vertex, frontal sclerite prominent protuberances; in fore femur internal spines with two black spots; pronotum with black line. First tarsal
18582 Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 Mantds of Utar Pradesh, India Yadav & Painkra J TT segment brown. Stgma with black spot at either ends Distributon: India (Arunachal Pradesh, Assam, (Majumder et al. 2015). Maharashtra, Manipur, Meghalaya, Odisha, West Bengal, Distributon: India (Andhra Pradesh, Chhatsgarh, & Utar Pradesh (new record)), Borneo, China, Flores, Maharashtra, Karnataka, Kerala, Tamil Nadu, & Utar Java, Malaysia, Moluccas, Myanmar, Philippines, Sri Lanka, Pradesh (new record)). Sumba, Sulawesi, Talaud Islands, and Thailand. Measurement: Body length 13mm Measurement: Body length 87mm Remark: Smaller one mantd. Remark: Larger one mantd.
Elmants trincomaliae (Saussure, 1869) Tenodera costalis (Blanchard, 1853) (Image 14) 1869. Gonypeta trincomaliae Saussure, Mit. Schweiz. 1853. Mants costalis Blanchard, Voy. Pole Sud, Zool. Entomol. Ges., 3: 63-64. Iv(I): 353 1915a. Elmants trincomaliae Giglio-Tos, Bull. Soc. 1904. Tenodera costalis Kirby, Syn. Cat. Orth. 1: 238. Entomol. Ital., 46: 161. 1927. Tenodera costalis Giglio-Tos, Orthoptera. 2014. Elmants trincomaliae Mukherjee et al. Priamus, Mantdae. Das Tierreich/animal kingdom, 414. 30: 25. Materials examined: RMDCRS 0021-0022, 1 male, 1 2015. Elmants trincomaliae Majumder et al. Zool. Surv. female, 10.iv.2018, Karmhari (Ghazipur), Utar Pradesh, India, 115(4): 377. India, on grasses, coll. R.S. Yadav. Materials examined: RMDCRS 0019, 1 male, 24.III.2018, Diagnosis: Body slender, sof green in colour, much Zamania (Ghazipur), Utar Pradesh, India, on stones, coll. elongated; frontal sclerite much wider, rather a double R.S. Yadav. curved edge on the upper side; antennae medium-long, Diagnosis: Small brownish insect. In foreleg, proximal brownish to body colored; Fore coxae smooth or partal two external spines close beset; foretbia with 10 external spines in male, In female fnely dentculate, the apex of spines. Hind metatarsus longer than rest segments together the femur spines black; fore femora with four discoidal and (Majumder et al. 2015). four external spines, laterally smooth; 1st discoidal spines Distributon: India (Andhra Pradesh, Chhatsgarh shorter than 2nd. Both wings well developed, slightly longer (Bastar), Karnataka, Kerala, Maharashtra, Tamil Nadu, & than body; hind femora with apical spine. Pronotum litle Utar Pradesh (new record)) and Sri Lanka. dilated anteriorly keeled in its middle, fnely dentculate Measurement: Body length 14 mm on its edges, especially in its anterior porton, metazona Remark: Smaller one mantd. of the pronotum slightly longer or in similar length to the fore coxae; fore wing long, rather narrow, acuminate, all the Tenodera fasciata (Oliver, 1792) (Image 13) anterior part slightly darker, discoidal area of male is hyaline, 1912. Tenodera fasciata Giglio-Tos. Bull. Soc. Entomol. in female one third the densely veined, the hind wings are Ital., 43: 45. adorned with black basal spot, costal area blood-stained, 2009. Tenodera fasciata Sureshan, Rec. Zool. Sur. India. discoidal area transverse venules dark/smoky, anal area Occasional Paper No. 305: 37. dark/smoky, hyaline areas with irregular fenestrae (Giglio- 2007. Tenodera fasciata Mandal et al. Zoos’ Print Tos 1912 & 1927). Journal 22(6): 2719. Distributon: India (Utar Pradesh (new record)). 2014. Tenodera fasciata Mukherjee et al. Priamus, 30: Measurement: Body length 89mm, pronotum 30. 33.32mm; pronotum width 4.92mm; metazona 27.20mm; 2017. Tenodera fasciata Raut & Gaikwad, J. Threat. Taxa, Fore wing 48.38mm. 9(6): 10351–10354 Remark: First tme recorded from India. More detail Materials examined: RMDCRS 0020, 1 male, 10.iv.2018, study and comparison is needed. Zamania (Ghazipur), Utar Pradesh, India, on grasses, coll. R.S. Yadav. Hierodula coarctata Saussure, 1869 (Image 15) Diagnosis: Body slender, much elongated; antenna very 1869. Hierodula coarctata Saussure, Mit. schweiz. ent. much elongated, pronotum nearly equal or longer than fore Ges. 3:67 coxae. Fore femora with four discoidal and four external 1927. Parahierodula coarctata Giglio-Tos. Tierreich spines, laterally smooth; 1st discoidal spines shorter than 2nd. 50:458 1935. Hierodula coarctata Beier. Genera Insect. 203: Both wings well developed; hind femora with apical spine; 84 anal cerci conical and hind wing with reddish transverse 2010. Hierodula coarctata Mukherjee and Ghate, J. veinlets in costal area (Raut & Gaikwad 2017). Threat. Taxa, 2(9): 1167-1171.
Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 18583 J TT Mantds of Utar Pradesh, India Yadav & Painkra 2014. Hierodula coarctata Mukherjee et al. Priamus, spot, fore femora without black spot; a distnct transverse 30: 30. pink line on the vertex, prominent in fresh specimen. Claw 2018b. Hierodula coarctata Yadav et al., J. Exp. Zool. groove of femora in the middle; fore wing semi- hyaline and India Vol. 21, No. 2, pp. 745-747. both wings shorter to body. Materials examined: RMDCRS 0023-0025, 1 female, Distributon: India (Karnataka, Kerala, Madhya Pradesh, 12.iv.2019, Rohuna (Ghazipur), Utar Pradesh, India, on light Maharashtra, Manipur, Odisha, Utar Pradesh (new record), bulb. 2 male, 1 female, 07.ix.2019, Zamania (Ghazipur), on & West Bengal), Asia, Africa, Europe, North America, Canada, shrub plant, coll. R.S. Yadav. Jordan, Lebanon, and United Arab Emirates (Mukherjee et Diagnosis: The prosternum and mesosternum with al. 2014). red coloured transverse band were found. The supra coxal Measurement: Body length 57mm dilaton of pronotum was broader than male and prozonal Remark: Species shows aggression when disturbed. tubercles on edges were comparatvely stronger than the male. The metazona with distnct median carina was there. Statlia maculata (Thunberg, 1784) The forefemur was dark green with dark patches on inner 1985. Statlia maculata Mukherjee & Hazra, Rec. zool. side. The forefemur has black coloured larger spines and Surv. India, 82(1-4): 34. discoidal spines also. Wings were mostly similar with male. 1927. Statlia maculata Giglio-Tos. Das nerreich, 50: 410. Male: The body colour of male was green. The head was 2014. Statlia maculata Mukherjee et al., Priamus, 30: triangular; eyes were rounded with prominent ocelli. The 38 supra coxal dilaton of the pronotum was somewhat wide. Materials examined: RMDCRS 0028-0030, 2 male, 1 The prozona has some fner tubercles. The metazonal female, 23.VI.2018, Chahaniya (Chandauli), Utar Pradesh, carinaton was not clear. Externally, forefemur was green in India, on bushes, coll. R.S. Yadav. colour. Larger spines of forefemur were black in colour. The Diagnosis: Body brownish, pronotum slender, longer meso and meta thoracic legs are yellowish green in colour. than fore coxa, claw groove of fore femora situated above The forewing was green in colour and veins were green with middle, four external and four discoidal spines, inner disc yellow coloured subcosta. There were yellowish to white with pale yellow and black patches; tbiae with seven colour stgma surrounded by brownish colour patches on external spines; femora with shining yellow patch, ofen the forewing. bordered by a black patch, larger internal spines of femora Distributon: India (Andhra Pradesh, Bihar, Maharshtra, not entrely black, wings as long as abdomen. West Bengal, & Utar Pradesh), Nepal, Java, Pakistan, & Distributon: India (Andaman Island, Andhra Pradesh, Australia? (Mukherjee et al. 2014; Yadav et al. 2018). Arunachal Pradesh, Assam, Bihar, Himachal Pradesh, Measurement: Body length 47mm Madhya Pradesh, Maharashtra, Meghalaya, Kerala, Odisha, Remark: Species may show gradual changes in their Sikkim, Utar Pradesh, & West Bengal) Annam, Borneo, colouraton. China, Japan, Java, Labuan, Myanmar, Malaysia, Maluku Islands, Nepal, New Guinea, Palawan, Sri Lanka, and Subfamily: Mantnae Burmeister, 1838 Sumatra, (in Staatliches Museum furNaturkunde, Karlsruhe, Mants religiosa religiosa Linnaeus, 1758 (Image 16) Germany (SMNK): Laos, Pakistan, Philippines, Thailand, & 1758. Gryllus (Mants) religiosus Linnaeus. Sys. Nat. 10: Vietnam) (Mukherjee et al. 2014). p.426 Measurement: Body length 55mm 1927. Mants religiosa Giglio-Tos, 1927. Das Tierreich, Remark: Brown coloured species, male & female are 50: 406. more or less same size. 2009. Mants religiosa Sureshan, Rec. Zool. Sur. India. Occasional Paper No. 305: 30. Statlia nemoralis (Saussure, 1870) 2014. Mants religiosa religiosa Mukherjee et al. 1927. Statlia nemoralis Giglio-Tos. Das nerreich, 50: Priamus, 30: 37. 411. Materials examined: RMDCRS 0026-0027, 1 male, 1 2014. Statlia nemoralis Mukherjee et al., Priamus, 30: female, 22.viii.2018, Rohuna (Ghazipur), Utar Pradesh, 38 India, on paddy, coll. R.S. Yadav. Materials examined: RMDCRS 0031-0032, 1 male,1 Diagnosis: Pronotum a litle longer than fore coxa, female, 23.xi.2017, Rohuna (Ghazipur), Utar Pradesh, India, prosternum has two small tubercles just to the base; shorter on bushes, coll. R.S. Yadav. when compared to that of Statlia Stal, fore coxa internally Diagnosis: Body Straw yellow with grass green shade, with a basal black spot which ofen encloses an oval yellow pronotum a shorter than fore coxa, claw groove of fore
18584 Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 Mantds of Utar Pradesh, India Yadav & Painkra J TT femora situated above middle, with yellow and a black external spines and eight internal spines, wing is shorter patch in front of it, four external and four discoidal spines, than abdomen, middle and hind femora without geniuclar inner disc with pale yellow and black patches; tbiae with spines, cerci foliaceous, longer than wide. seven external spines; prosternum without a black patch Distributon: India (Andhra Pradesh, Chhatsgarh, posteriorly, wings as long as abdomen. Kerala, Maharashtra, Orissa, West Bengal, & Utar Pradesh Distributon: India (Arunachal Pradesh, Himachal (new record)), Iran, Malaysia, and Sri Lanka. Pradesh, Maharashtra, Manipur, Kerala, Tamil Nadu, West Measurement: Body length 112mm Bengal, & Utar Pradesh (new record)), Borneo, China, Remark: One of the larger species. Japan, Java, Korea, Myanmar, Malayasia, Philippines, Sumatra, Taiwan, and Vietnam (Mukherjee et al. 2014). Toxoderopsis taurus Wood-mason, 1889 Measurement: Body length 58mm 1927. Toxoderopsis taurus Giglio-Tos. Das Tierreich, 50: Remark: Male & female were mostly similar. 569. 2009. Toxoderopsis taurus Sureshan, Rec. zool. Surv. Cheddikulama straminea Henry, 1932 (Image 17) India, Occ. Paper (305): 34. 1995. Cheddikulama straminea Mukherjee et al., 2014. Toxoderopsis taurus Mukherjee et al., Priamus, Oriental Insects 29(1): 202, 317, 320. 30: 2, 3, 47. 2007. Cheddikulama straminea Vyjayandi, (267): IX, 3, Materials examined: RMDCRS 0037, 1 female, 134, 154. 12.IX.2019, Dehariya (Ghazipur), Utar Pradesh, India, on 2014. Cheddikulama straminea Mukherjee et al., logs, coll. R.S. Yadav. Priamus, 30: 45 Diagnosis: Body black brown in colour, middle and hind Materials examined: RMDCRS 0033-0034, 1 male,1 tbiae not carinate; eyes with a distnctly projectng spine, female, 02.viii.2017, Dehariya (Ghazipur), Utar Pradesh, median lobe of vertex higher than laterals, hind wings India, on logs, coll. R.S. Yadav. hyaline, middle and hind femora with genicular spines; cerci Diagnosis: Body straw yellow coloured, head pentagonal, long, fat, crested at tp. fatened; occiput produced into two angular lobes; conical Distributon: India (Andhra Pradesh, Bihar, Maharashtra, eyes with tubercle, pronotum slender as twice longer than Madhya Pradesh, Odisha, & Utar Pradesh (new record)) fore coxa; hindlegs much longer than middle legs; wings and Pakistan (Mukherjee et al. 2014). ornate; forewing with parallel venaton; hindwing with Measurement: Body length 88mm purple blotch and concentrically arranged purple patches; Remark: Sometme confuse with Aethalochroa cerci short. ashmoliana. Distributon: India (Kerala and Utar Pradesh) and Sri Lanka (Vyjayandi 2007; Mukherjee et al. 2014). Family Empusidae Burmeister, 1838; Subfamily Empusinae Measurement: Body length 59mm Burmeister, 1838 Remark: Male & female were mostly similar. Empusa (Empusa) gutula (Thunberg, 1815) (Image 19) 1889. Gongylus gutula Thunberg, 1815: 5: 294. Aethalochroa ashmoliana (Westwood, 1841) (Image 2002. Empusa (Empusa) gutula Ehrmann, Mantodea 18) der Welt: 127. 1951. Aethalochroa ashmoliana Roonwal & Bhasin, 2009. Empusa gutula Sureshan, Rec. zool. Surv. India, Indian Forester, 77(5): 313-318. Occ. Paper (305): 16 1904a. Aethalochroa ashmoliana Kirby,̇ 1904: Brit. Mus. 2014. Empusa gutula Mukherjee et al., Priamus, 30: 2, (Nat. Hist.), Vol. 1 1: 308 3, 51. 2014. Aethalochroa ashmoliana syn. As Loxomants Materials examined: RMDCRS 0038-0039, 1 female,1 indica Mukherjee et al., Priamus, 30: 2, 3, 47. female, 05.viii.2019, Dehariya (Ghazipur), Utar Pradesh, 2015. Aethalochroa ashmoliana Majumder et al. Zool. India, on paddy, coll. R.S. Yadav. Surv. India, 115(4): 382. Diagnosis: Fore femora without lobe and superior edge Materials examined: RMDCRS 0035-0036, 1 male, almost straight; middle and hind femora without dorsal 23.XI.2017, 1 female, 12.X.2019 Dehariya (Ghazipur), Utar lobes, vertex prolonged into a conical protuberance, armed Pradesh, India, on logs, coll. R.S. Yadav. medially and laterally by triangular sharp spines, apex litle Diagnosis: Body dark in coloured middle and hind expanded and bifog with a median carina extended into tbiae dorsally carinated, eyes prominent with a very small sharp point; antennae simple in female and pectnate in dorsal tubercle, upper edge of vertex concave, fore leg male; pronotum slender, slightly spined; forewing litle
Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 18585 J TT Mantds of Utar Pradesh, India Yadav & Painkra
Image 1. Observed mantds during feld surveys in the province of the Utar Pradesh: 1—Euantssa pulchra | 2 & 3—Odontomants montana nymph & adult | 4—Creobroter apicalis | 5—forefemur of Ephestasula pictpes | 6 & 7—ootheca of Hestasula brunneriana & adult | 8— Humbertella ceylonica | 9—Humbertella similis | 10 & 11—Schizocephala bicornis | 12—Amants saussurei (nymph) | 13—Tenodera fasciata | 14—Tenodera costalis | 15—Hierodula coarctata | 16—Mants religiosa religiosa | 17—Cheddikulama straminea | 18—Aethalochroa ashmoliana | 19—Empusa (empusa) gutula | 20 & 21—Empusa spinosa. © R.S. Yadav
longer than body, stgma with two brown spots at corner, coll. R.S. Yadav. costal area green, opaque. Diagnosis: Longer than Empusa gutula, yellowish-green, Distributon: India (Andhra Pradesh, Odisha, Rajasthan, slender body; vertex of head with a conical protuberance, & Utar Pradesh), Algeria, Angola, Cameroon, Egypt, Kenya, bifurcated at top; pronotum slender, long with distnct Libya, Madagascar, Mauritania, Morocco, Namibia, Senegal, spines, longer than fore coxae; upper edge of fore femur Somalia, Tanzania, Transversal, Tunisia, Gambia, South Africa, is staright, and simple fore femur (Majumder 2015); wing and Tanzania (Mukherjee et al. 2014). extended beyond the abdomen; abdominal segments Measurement: Body length 58mm usually expanded laterally. Remark: Species recorded from the state previously. Distributon: India (Chandigarh, Chhatsgarh, & Utar Pradesh (new record)) (Mukherjee et al. 2014, Majumder et Empusa spinosa Krauss, 1902 (Image 20 & 21) al. 2015) 1964. Empusa spinosa Gupta, Current Sci. 33: 370. Measurement: Body length 59mm 2002. Empusa spinosa Ehrmann, Mantodea der Welt, Remark: Very less studied species in the country. 130. 2014. Empusa spinosa Mukherjee et al., Priamus, 30: 2, 3, 51. CONCLUSION AND SUMMARY 2015. Empusa spinosa Majumder et al. Zool. Surv. India, 115(4): 382. The 21 species of mantds discussed here, 10 species Materials examined: RMDCRS 0040-0042, 1 male and are frstly recorded from Utar Pradesh and one species frst 1 female, 15.x.2017, Udharanpur (Ghazipur), Utar Pradesh, tme reported from the country. Further more intensive feld India, on Tectona. 1 female, 05.viii.2019, Dehariya (Ghazipur), work may enrich mantd fauna of the Utar Pradesh, India.
18586 Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 Mantds of Utar Pradesh, India Yadav & Painkra J TT REFERENCES India (Insecta: Mantodea). Oriental Insects 29(1): 185–358. Mukherjee, T.K., G. Iyer & P. Chaterjee (2017). Twenty-three new records Blanchard, E. (1853). Descripton des insectes – Orthopteres, pp. 349–376. of Mantodea (Insecta) from some states of India. Journal of Threatened In: Jaquinot, M. (ed.), Voyage au Pole Sud et dans l’Oceanie sur les Taxa 9(2): 9829–9839. htp://doi.org/10.11609/jot.1936.9.2.9829-9839 corvetes L’Astrolabe et La Zelee. Zoologie IV (1). Trunot & Co., Paris. Mukherjee, T.K. & M.S. Shishodia (1999). Mantodea of Patalkot Bolivar, I. (1899). Les Orthoptères de St.-Joseph`s College à Trichinopoly Chhindwara dist. Madhya Pradesh, India. Records of Zoological Survey (Sud de l`Inde). Annales de la Société entomologique de France 68: 761- of India 97(4): 45-48. 812, 2pls. Mukherjee, T.K. & M.S. Shishodia (2000). Insecta: Mantodea. Zoological Chandra, K. (2009). Insecta: Mantodea. Fauna of Bandhavgarh Tiger Survey of the India, Fauna of Renuka Wetland (Himachal Pradesh). Reserve (Madhya Pradesh), Zoological Survey of India. Conservaton Wetland Ecosystem Series 2: 63–66. Area Series 40: 59–61. Mukherjee, T.K., R. Ehrmann & P. Chaterjee (2014). Checklist of Mantodea Chaterjee, P. & T.K. Mukherjee (2013). Mants indica Mukherjee, 1995: (Insecta) from India. Priamus (Serial Publicaton of the Centre for a synonym of Statlia nemoralis (Saussure, 1870) (Insecta: Mantodea). Entomological Studies Ankara) (Suppl.) (30): 1–61. Journal of Threatened Taxa 5(14): 4907–4909. htps://doi.org/10.11609/ Mukherjee, T.K. & A.K. Hazra (1985). New record of a desertcolous Mantd JoTT.o3289.4907-9 Family (Mantodea: Eremiaphilidae) from India. Entomon 10(3): 245–247. Chaturvedi N., & V. Hedge (2000). Mantd fauna of Sanjay Gandhi Natonal Nadkerni, N.T. (1965). A note on mantds and tetgonids in the collecton of Park, Mumbai, with some new records for Maharashtra State. Journal of Bombay Natural History Society. Journal of the Bombay Natural History the Bombay Natural History Society 97: 295–297. Society 62(1): 76–83. Ehrmann, R. & R. Roy (2002). Systematsche Aufstellung der Gatungen. In: Ote, D., L. Spearman & B.D.S. Martn (2020). Mantodea Species File Mantodea: Gotesanbeterinnen der Welt. Natur und Tier-Verlag. Online. Version 5.0/5.0. [12.iii.2020].
Threatened Taxa Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2021 | 13(6): 18578–18587 18587 The Journal of Threatened Taxa (JoTT) is dedicated to building evidence for conservaton globally by OPEN ACCESS publishing peer-reviewed artcles online every month at a reasonably rapid rate at www.threatenedtaxa.org. All artcles published in JoTT are registered under Creatve Commons Atributon 4.0 Internatonal License unless otherwise mentoned. JoTT allows allows unrestricted use, reproducton, and distributon of artcles in any medium by providing adequate credit to the author(s) and the source of publicaton.
ISSN 0974-7907 (Online) | ISSN 0974-7893 (Print)
May 2021 | Vol. 13 | No. 6 | Pages: 18411–18678 Date of Publicaton: 26 May 2021 (Online & Print) www.threatenedtaxa.org DOI: 10.11609/jot.2021.13.6.18411-18678
Conservaton Applicaton Review
First atempt at rehabilitaton of Asiatc Black Bear cubs to the wild in Thailand Termites (Blatodea: Isoptera) of southern India: current knowledge on distributon and – Robert Steinmetz, Worrapan Phumanee, Rungnapa Phoonjampa & Suthon Weingdow, systematc checklist Pp. 18411–18418 – M. Ranjith & C.M. Kalleshwaraswamy, Pp. 18598–18613
Communicatons Short Communicatons
Status of Sumatran Tiger in the Berbak-Sembilang landscape (2020) Populaton status and distributon of Ibisbill Ibidorhyncha struthersii (Vigors, 1832) (Aves: – Tomi Ariyanto, Yoan Dinata, Dwiyanto, Erwan Turyanto, Waluyo Sugito, Sophie Kirklin & Rajan Charadriiformes: Ibidorhynchidae) in Kashmir Valley, India Amin, Pp. 18419–18426 – Iqram Ul Haq, Bilal A. Bhat, Khursheed Ahmad & Asad R. Rahmani, Pp. 18614–18617
The diversity of small mammals in Pulau Perhentan Kecil, Terengganu, Malaysia A new fsh species of genus Garra (Teleostei: Cyprinidae) from Nagaland, India – Aminuddin Baqi, Isham Azhar, Ean Wee Chen, Faisal Ali Anwarali Khan, Chong Ju Lian, – Sophiya Ezung, Bungdon Shangningam & Pranay Punj Pankaj, Pp. 18618–18623 Bryan Raveen Nelson & Jayaraj Vijaya Kumaran, Pp. 18427–18440 Occurrence of Tamdil Leaf-liter Frog Leptobrachella tamdil (Sengupta et al., 2010) (Amphibia: Paterns, perceptons, and spatal distributon of human-elephant (Elephas maximus) incidents in Megophryidae) from Manipur, India and its phylogenetc positon Nepal – Ht. Decemson, Vanlalsiammawii, Lal Biakzuala, Mathipi Vabeiryureilai, Fanai Malsawmdawngliana – Raj Kumar Koirala, Weihong Ji, Yajna Prasad Timilsina & David Raubenheimer, Pp. 18441–18452 & H.T. Lalremsanga, Pp. 18624–18630
Assessing spato-temporal paterns of human-leopard interactons based on media reports in Further additons to the Odonata (Insecta) fauna of Asansol-Durgapur Industrial Area, Paschim northwestern India Bardhaman, India – Kaushal Chauhan, Arjun Srivathsa & Vidya Athreya, Pp. 18453–18478 – Amar Kumar Nayak & Subhajit Roy, Pp. 18631–18641
Bat diversity in the Banpale forest, Pokhara, Nepal during spring season A note on the ecology and distributon of Litle Bloodtail Lyriothemis acigastra Brauer, 1868 – Prabhat Kiran Bhatarai, Basant Sharma, Anisha Neupane, Sunita Kunwar & Pratyush Dhungana, (Insecta: Odonata: Libellulidae) in Kerala, India Pp. 18479–18489 – Jeevan Jose, Muhamed Sherif & A. Vivek Chandran, Pp. 18642–18646
A patho-microbiological study of tssue samples of the Greater Adjutant Leptoptlos dubius (Aves: Viewpoint Ciconiiformes: Ciconiidae) that died in Deeporbeel Wildlife Sanctuary, Assam, India – Derhasar Brahma, Parikshit Kakat, Sophia M. Gogoi, Sharmita Doley, Arpita Bharali, Biswajit Duta, A unique archetype of conservaton in Himachal Pradesh, western Himalaya, India Taibur Rahman, Saidul Islam, Arfan Ali, Siraj A. Khan, Sailendra Kumar Das & Nagendra Nath Barman, – Rupali Sharma, Monika Sharma, Manisha Mathela, Himanshu Bargali & Amit Kumar, Pp. 18490–18496 Pp. 18647–18650
Vaduvur and Sitheri lakes, Tamil Nadu, India: conservaton and management perspectve Notes – V. Gokula & P. Ananth Raj, Pp. 18497–18507 A camera trap record of Asiatc Golden Cat Catopuma temminckii (Vigors & Horsfeld, 1827) A new species of shieldtail snake (Squamata: Uropeltdae: Uropelts) from the Bengaluru uplands, (Mammalia: Carnivora: Felidae) in State Land Forest, Merapoh, Pahang, Malaysia India – Muhamad Hamirul Shah Ab Razak, Kamarul Hambali, Aainaa Amir, Norashikin Fauzi, Nor Hizami – S.R. Ganesh, K.G. Punith, Omkar D. Adhikari & N.S. Achyuthan, Pp. 18508–18517 Hassin, Muhamad Azahar Abas, Muhammad Firdaus Abdul Karim, Ai Yin Sow, Lukman Ismail, Nor Azmin Huda Mahamad Shubli, Nurul Izzat Adanan, Ainur Izzat Bakar, Nabihah Mohamad, Nur A looming exotc reptle pet trade in India: paterns and knowledge gaps Izyan Fathiah Saimeh, Muhammad Syafq Mohmad Nor, Muhammad Izzat Hakimi Mat Naf & Syafq – A. Pragatheesh, V. Deepak, H.V. Girisha & Monesh Singh Tomar, Pp. 18518–18531 Sulaiman, Pp. 18651–18654
Legal or unenforceable? Violatons of trade regulatons and the case of the Philippine Sailfn Lizard Reappearance of Dhole Cuon alpinus (Mammalia: Carnivora: Canidae) in Gujarat afer 70 years Hydrosaurus pustulatus (Reptlia: Squamata: Agamidae) – A.A. Kazi, D.N. Rabari, M.I. Dahya & S. Lyngdoh, Pp. 18655–18659 – Sarah Heinrich, Adam Toomes & Jordi Janssen, Pp. 18532–18543 Matng behavior of Eastern Spoted Skunk Spilogale putorius Linnaeus, 1758 (Mammalia: Conservaton breeding of Northern River Terrapin Batagur baska (Gray, 1830) in Sundarban Tiger Carnivora: Mephitdae) revealed by camera trap in Texas, USA Reserve, India – Alexandra C. Avrin, Charles E.Pekins & Maximillian L. Allen, Pp. 18660–18662 – Nilanjan Mallick, Shailendra Singh, Dibyadeep Chaterjee & Souritra Sharma, Pp. 18544–18550 Record of Indian Roofed Turtle Pangshura tecta (Reptlia: Testudines: Geoemydidae) from Koshi Discovery of two new populatons of the rare endemic freshwater crab Louisea yabassi Mvogo Tappu Wildlife Reserve, Nepal Ndongo, von Rintelen & Cumberlidge, 2019 (Brachyura: Potamonautdae) from the Ebo Forest – Ashmita Shrestha, Ramesh Prasad Sapkota & Kumar Paudel, Pp. 18663–18666 near Yabassi in Cameroon, Central Africa, with recommendatons for conservaton acton – Pierre A. Mvogo Ndongo, Thomas von Rintelen, Christoph D. Schubart, Paul F. Clark, Additonal distributon records of Zimiris doriae Simon, 1882 (Araneae: Gnaphosidae) from India Kristna von Rintelen, Alain Didier Missoup, Christan Albrecht, Muriel Rabone, Efole Ewoukem, – Dhruv A. Prajapat, Pp. 18667–18670 Joseph L. Tamesse, Minete Tomedi-Tabi Eyango & Neil Cumberlidge, Pp. 18551–18558 Notes on new distributon records of Euaspa motokii Koiwaya, 2002 (Lepidoptera: Lycaenidae: Checklists of subfamilies Dryptnae and Panagaeinae (Insecta: Coleoptera: Carabidae) from the Theclinae) from Bhutan Indian subcontnent – Jigme Wangchuk, Dhan Bahadur Subba & Karma Wangdi, Pp. 18671–18674 – V.A. Jithmon & Thomas K. Sabu, Pp. 18559–18577 New distributon records of two litle known plant species, Hedychium longipedunculatum A.R.K. Mantds (Insecta: Mantodea) of Utar Pradesh, India Sastry & D.M. Verma (Zingiberaceae) and Mazus dentatus Wall. ex Benth. (Scrophulariaceae), – Ramesh Singh Yadav & G.P. Painkra, Pp. 18578–18587 from Meghalaya, India – M. Murugesan, Pp. 18675–18678 An assessment of genetc variaton in vulnerable Borneo Ironwood Eusideroxylon zwageri Teijsm. & Binn. in Sarawak using SSR markers Publisher & Host – Sit Fatmah Md.-Isa, Christna Seok Yien Yong, Mohd Nazre Saleh & Rusea Go, Pp. 18588–18597
Member
Threatened Taxa