Butterflies and Dragonflies
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The Wing Venation of Odonata
International Journal of Odonatology, 2019 Vol. 22, No. 1, 73–88, https://doi.org/10.1080/13887890.2019.1570876 The wing venation of Odonata John W. H. Trueman∗ and Richard J. Rowe Research School of Biology, Australian National University, Canberra, Australia (Received 28 July 2018; accepted 14 January 2019) Existing nomenclatures for the venation of the odonate wing are inconsistent and inaccurate. We offer a new scheme, based on the evolution and ontogeny of the insect wing and on the physical structure of wing veins, in which the veins of dragonflies and damselflies are fully reconciled with those of the other winged orders. Our starting point is the body of evidence that the insect pleuron and sternum are foreshortened leg segments and that wings evolved from leg appendages. We find that all expected longitudinal veins are present. The costa is a short vein, extending only to the nodus, and the entire costal field is sclerotised. The so-called double radial stem of Odonatoidea is a triple vein comprising the radial stem, the medial stem and the anterior cubitus, the radial and medial fields from the base of the wing to the arculus having closed when the basal sclerites fused to form a single axillary plate. In the distal part of the wing the medial and cubital fields are secondarily expanded. In Anisoptera the remnant anal field also is expanded. The dense crossvenation of Odonata, interpreted by some as an archedictyon, is secondary venation to support these expanded fields. The evolution of the odonate wing from the palaeopteran ancestor – first to the odonatoid condition, from there to the zygopteran wing in which a paddle-shaped blade is worked by two strong levers, and from there through grade Anisozygoptera to the anisopteran condition – can be simply explained. -
Lesson: Key Details Lesson Topic: Identify and Explain Key Details Question 1
Lesson: Key Details Lesson Topic: Identify and explain key details Question 1: Read the text below. Sea Stars Written by Lindsey Crowe There are many different kinds of sea stars. For example, some sea stars have five legs and some have ten legs. Some have skinny arms and some have wide arms. Sea stars are found in the ocean. Some sea stars live in warm oceans. Some live in cold oceans. They can be found in deep and shallow waters. Some people call sea stars star fish. Sea stars are not fish. They do not have gills, fins, or bones. Which of the following key details supports the idea that there are many different kinds of sea stars? There are many different kinds of sea stars. Sea stars are found in the ocean. Some sea stars have five legs and some have ten legs. They do not have gills, fins, or bones. Question 2: Read the text below. There are many different kinds of sea stars. For example, some sea stars have five legs and some have ten legs. Some have skinny arms and some have wide arms. Why is the underlined sentence an important key detail? because it explains why the author took pictures and wrote a text about sea stars because it is an important fact that we need to know to understand about different kinds of sea stars because it explains what kind of sea star is in the photograph because it gives the names of different types of sea stars Question 3: Read the text below. -
Superior National Forest
Admirals & Relatives Subfamily Limenitidinae Skippers Family Hesperiidae £ Viceroy Limenitis archippus Spread-wing Skippers Subfamily Pyrginae £ Silver-spotted Skipper Epargyreus clarus £ Dreamy Duskywing Erynnis icelus £ Juvenal’s Duskywing Erynnis juvenalis £ Northern Cloudywing Thorybes pylades Butterflies of the £ White Admiral Limenitis arthemis arthemis Superior Satyrs Subfamily Satyrinae National Forest £ Common Wood-nymph Cercyonis pegala £ Common Ringlet Coenonympha tullia £ Northern Pearly-eye Enodia anthedon Skipperlings Subfamily Heteropterinae £ Arctic Skipper Carterocephalus palaemon £ Mancinus Alpine Erebia disa mancinus R9SS £ Red-disked Alpine Erebia discoidalis R9SS £ Little Wood-satyr Megisto cymela Grass-Skippers Subfamily Hesperiinae £ Pepper & Salt Skipper Amblyscirtes hegon £ Macoun’s Arctic Oeneis macounii £ Common Roadside-Skipper Amblyscirtes vialis £ Jutta Arctic Oeneis jutta (R9SS) £ Least Skipper Ancyloxypha numitor Northern Crescent £ Eyed Brown Satyrodes eurydice £ Dun Skipper Euphyes vestris Phyciodes selenis £ Common Branded Skipper Hesperia comma £ Indian Skipper Hesperia sassacus Monarchs Subfamily Danainae £ Hobomok Skipper Poanes hobomok £ Monarch Danaus plexippus £ Long Dash Polites mystic £ Peck’s Skipper Polites peckius £ Tawny-edged Skipper Polites themistocles £ European Skipper Thymelicus lineola LINKS: http://www.naba.org/ The U.S. Department of Agriculture (USDA) prohibits discrimination http://www.butterfliesandmoths.org/ in all its programs and activities on the basis of race, color, national -
Monitoring Dragonfly Migration in North America Protocols for Citizen Scientists
Monitoring Dragonfly Migration in North America Protocols for Citizen Scientists Migratory Dragonfly Partnership Blank on purpose Monitoring Dragonfly Migration in North America Protocols for Citizen Scientists Migratory Dragonfly Partnership Canada • United States • Mexico www.migratorydragonflypartnership.org © 2014 by The Migratory Dragonfly Partnership The Migratory Dragonfly Partnership uses research, citizen science, education, and outreach to under- stand North American dragonfly migration and promote conservation. MDP steering committee members represent a range of organizations, including: Ontario Ministry of Natural Resources; Peggy Notebaert Nature Museum; Pronatura Veracruz; Rutgers University; Slater Museum of Natural History, University of Puget Sound; Smithsonian Conservation Biology Institute; St. Edward's University; U. S. Forest Service International Programs; U. S. Geological Survey; Vermont Center for Ecostudies; and the Xerces Society for Invertebrate Conservation. Migratory Dragonfly Partnership Project Coordinator, Celeste Mazzacano [email protected] 628 NE Broadway, Suite 200, Portland, OR 97232 Tel (855) 232-6639 Fax (503) 233-6794 www.migratorydragonflypartnership.org Acknowledgements Funding for the Migratory Dragonfly Partnership's work is provided by the U.S. Forest Service Inter- national Programs. We thank the photographers who generously allowed use of their images. Copyright of all photographs remains with the photographers. Front and Back Cover Photographs Common Green Darner (Anax junius) male. Photograph © John C. Abbott/Abbott Nature Photography. CONTENTS Summary Page 1 1. Introduction Page 3 1.1 Objectives and Goals Page 3 Box 1: Citizen Science Projects, page 4. 2. Citizen Science Projects Page 5 2.1 Migration Monitoring Page 5 2.1.1 Fall Migration Observations Page 5 - Objectives, page 5. Box 2: MDP Monitoring Projects, page 6. -
Wetlands Invertebrates Banded Woollybear(Isabella Tiger Moth Larva)
Wetlands Invertebrates Banded Woollybear (Isabella Tiger Moth larva) basics The banded woollybear gets its name for two reasons: its furry appearance and the fact that, like a bear, it hibernates during the winter. Woollybears are the caterpillar stage of medium sized moths known as tiger moths. This family of moths rivals butterflies in beauty and grace. There are approximately 260 species of tiger moths in North America. Though the best-known woollybear is the banded woollybear, there are at least 8 woollybear species in the U.S. with similar dense, bristly hair covering their bodies. Woollybears are most commonly seen in the autumn, when they are just about finished with feeding for the year. It is at this time that they seek out a place to spend the winter in hibernation. They have been eating various green plants since June or early July to gather enough energy for their eventual transformation into butterflies. A full-grown banded woollybear caterpillar is nearly two inches long and covered with tubercles from which arise stiff hairs of about equal length. Its body has 13 segments. Middle segments are covered with red-orange hairs and the anterior and posterior ends with black hairs. The orange-colored oblongs visible between the tufts of setae (bristly hairs) are spiracles—entrances to the respiratory system. Hair color and band width are highly variable; often as the caterpillar matures, black hairs (especially at the posterior end) are replaced with orange hairs. In general, older caterpillars have more black than young ones. However, caterpillars that fed and grew in an area where the fall weather was wetter tend to have more black hair than caterpillars from dry areas. -
Dakota Skipper (Hesperia Dacotae)
Plan for the Controlled Propagation, Augmentation, and Reintroduction of Dakota skipper (Hesperia dacotae) April 30th 2017 Authors: Phil Delphey, Fish & Wildlife Biologist, U.S. Fish & Wildlife Service, Bloomington, MN Erik Runquist, Butterfly Conservation Biologist, Minnesota Zoo, Apple Valley, MN Cale Nordmeyer, Butterfly Conservation Specialist, Minnesota Zoo, Apple Valley, MN Plan for the Controlled Propagation, Augmentation, and Reintroduction of Dakota skipper 2017 Contents I. Executive Summary ............................................................................................................................... 4 II. Introduction .......................................................................................................................................... 5 III. Definitions of Terms Used in this Plan .............................................................................................. 5 IV. Justification for Captive Rearing, Augmentation and Reintroduction .............................................. 6 A. Status of the species ......................................................................................................................... 6 B. Likely Population Trends ................................................................................................................. 10 C. Threats ............................................................................................................................................ 11 D. Contributions to the Species’ Recovery ......................................................................................... -
Life Cycle of a Dragonfly, and Will Be Broken Down Into Sections Based on the Chronological Life Stages of This Insect
The Dragonfly Life Cycle Explained A BRIEF DISCUSSION ON THE LIFE CYCLE OF THE COMMON DRAGONFLY Kimberly Malcom | Pennsylvania State University, ENGL 202C: Technical Writing | 2016 Cover Photo: Mitchell, Forrest L and Lasswell, James L., A Dazzle of Dragonflies [Online Image]. Retrieved October 31, 2016 from http://www.audubon.org/magazine/july-august-2012/chasing-dragonflies-and-damselflies Audience and Purpose This document will explain the life cycle of a dragonfly, and will be broken down into sections based on the chronological life stages of this insect. Each section will be further broken down to include a photograph of the stage being discussed, and an explanation of what each stage entails. This document was prepared for the general public, and is meant to be a basic informational description of the life process of the common dragonfly for anyone with a curiosity to learn more about these unique insects. Introduction The dragonfly is a large, colorful, predatory insect generally found in or near watery locations in both the Northern and Southern Hemispheres. There are more than 5,000 known species of dragonflies, and fossil evidence suggests that they’ve been on the earth for many years. They have long, thin, colorful bodies, six legs, large eyes, and two pairs of transparent wings that allow them to propel themselves up, down, forward, backward and side to side Fotolia_1704510_xs [Online Image]. Retrieved October 31, 2016 from http://labs.blogs.com/its_alive_in_the_lab/2009/04/why-dragonfly.html while in flight. They are proficient fliers, and tend to only catch prey and eat while flying. -
Gardening with the Masters Growing, Gardening and Gaining Knowledge August/September 2020
Gardening With The Masters Growing, Gardening and Gaining Knowledge August/September 2020 Editor’s Corner WHAT’S HAPPENING By Marcia Winchester, Cherokee County Master Gardener As the worldwide health concerns continue with the COVID-19 virus, the Cherokee County Master Gardeners also continue to evaluate our monthly events and activities, and continue to follow UGA containers featuring pitcher plant bogs, Extension protocols and state hydrangeas, hostas, and even a number guidelines. of native azaleas. I tuck in a few annuals, and this gives me blooms all year to We continue to ensure the enjoy while sitting at my kitchen table. wellbeing of our members and community. Should we I can view a climbing hydrangea reschedule any of our events, (Schizophragma hydrangeoides) 30 feet you will be notified here, in our up a tulip poplar (Liriodendron tulipifera) newsletter, on our website, or from both the bedroom and family room. on Facebook. I also have a Magnolia liliflora ‘Ann’ that Please feel free to visit any of has beautiful deep pink/purple flowers our web links posted below. that I can see from both rooms. I have They will keep you up to date a grouping of a large clay pot planted with all our activities, events with ferns with a ceramic mushroom and changes, and help you to that I see when I’m practicing yoga. stay informed. A dear friend gave me a combination container with a beautiful 10 foot tall deep pink hibiscus that blooms in mid-summer, and when it stops in late September, an equally tall and dramatic Confederate rose (Hibiscus mutabilis) blooms until frost. -
Aravalli Range of Rajasthan and Special Thanks to Sh
Occasional Paper No. 353 Studies on Odonata and Lepidoptera fauna of foothills of Aravalli Range, Rajasthan Gaurav Sharma ZOOLOGICAL SURVEY OF INDIA OCCASIONAL PAPER NO. 353 RECORDS OF THE ZOOLOGICAL SURVEY OF INDIA Studies on Odonata and Lepidoptera fauna of foothills of Aravalli Range, Rajasthan GAURAV SHARMA Zoological Survey of India, Desert Regional Centre, Jodhpur-342 005, Rajasthan Present Address : Zoological Survey of India, M-Block, New Alipore, Kolkata - 700 053 Edited by the Director, Zoological Survey of India, Kolkata Zoological Survey of India Kolkata CITATION Gaurav Sharma. 2014. Studies on Odonata and Lepidoptera fauna of foothills of Aravalli Range, Rajasthan. Rec. zool. Surv. India, Occ. Paper No., 353 : 1-104. (Published by the Director, Zool. Surv. India, Kolkata) Published : April, 2014 ISBN 978-81-8171-360-5 © Govt. of India, 2014 ALL RIGHTS RESERVED . No part of this publication may be reproduced, stored in a retrieval system or transmitted in any form or by any means, electronic, mechanical, photocopying, recording or otherwise without the prior permission of the publisher. This book is sold subject to the condition that it shall not, by way of trade, be lent, resold hired out or otherwise disposed of without the publisher’s consent, in any form of binding or cover other than that in which, it is published. The correct price of this publication is the price printed on this page. Any revised price indicated by a rubber stamp or by a sticker or by any other means is incorrect and should be unacceptable. PRICE Indian Rs. 800.00 Foreign : $ 40; £ 30 Published at the Publication Division by the Director Zoological Survey of India, M-Block, New Alipore, Kolkata - 700053 and printed at Calcutta Repro Graphics, Kolkata - 700 006. -
Out of the Orient: Post-Tethyan Transoceanic and Trans-Arabian Routes
Systematic Entomology Page 2 of 55 1 1 Out of the Orient: Post-Tethyan transoceanic and trans-Arabian routes 2 fostered the spread of Baorini skippers in the Afrotropics 3 4 Running title: Historical biogeography of Baorini skippers 5 6 Authors: Emmanuel F.A. Toussaint1,2*, Roger Vila3, Masaya Yago4, Hideyuki Chiba5, Andrew 7 D. Warren2, Kwaku Aduse-Poku6,7, Caroline Storer2, Kelly M. Dexter2, Kiyoshi Maruyama8, 8 David J. Lohman6,9,10, Akito Y. Kawahara2 9 10 Affiliations: 11 1 Natural History Museum of Geneva, CP 6434, CH 1211 Geneva 6, Switzerland 12 2 Florida Museum of Natural History, University of Florida, Gainesville, Florida, 32611, U.S.A. 13 3 Institut de Biologia Evolutiva (CSIC-UPF), Passeig Marítim de la Barceloneta, 37, 08003 14 Barcelona, Spain 15 4 The University Museum, The University of Tokyo, Hongo, Bunkyo-ku, Tokyo 113-0033, Japan 16 5 B. P. Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii, 96817-0916 U.S.A. 17 6 Biology Department, City College of New York, City University of New York, 160 Convent 18 Avenue, NY 10031, U.S.A. 19 7 Biology Department, University of Richmond, Richmond, Virginia, 23173, USA 20 8 9-7-106 Minami-Ôsawa 5 chome, Hachiôji-shi, Tokyo 192-0364, Japan 21 9 Ph.D. Program in Biology, Graduate Center, City University of New York, 365 Fifth Ave., New 22 York, NY 10016, U.S.A. 23 10 Entomology Section, National Museum of the Philippines, Manila 1000, Philippines 24 25 *To whom correspondence should be addressed: E-mail: [email protected] Page 3 of 55 Systematic Entomology 2 26 27 ABSTRACT 28 The origin of taxa presenting a disjunct distribution between Africa and Asia has puzzled 29 biogeographers for centuries. -
Simultaneous Control of Head and Thoracic Temperature by the Green Darner Dragonfly Anax Junius (Odonata: Aeshnidae)
The Journal of Experimental Biology 198, 2373–2384 (1995) 2373 Printed in Great Britain © The Company of Biologists Limited 1995 SIMULTANEOUS CONTROL OF HEAD AND THORACIC TEMPERATURE BY THE GREEN DARNER DRAGONFLY ANAX JUNIUS (ODONATA: AESHNIDAE) MICHAEL L. MAY Department of Entomology, Cook College, New Jersey Agricultural Experiment Station, Rutgers University, New Brunswick, NJ 08903, USA Accepted 24 July 1995 Summary Anax junius is a large dragonfly that regulates thoracic during unrestrained flight in the field, Th is regulated temperature (Tth) during flight. This species, like several actively by increasing hemolymph circulation from the other intermittently endothermic insects, achieves control warm thorax at low Ta. Concurrent measurements of of Tth at least in part by increasing circulation of abdominal temperature (Tab) confirm that the abdomen is hemolymph to the abdomen at high air temperature (Ta), used as a ‘thermal window’ at Ta>30 ˚C but apparently not thus facilitating heat loss from the thorax. In this paper, I at lower Ta; thus, some additional mechanism(s) must exist demonstrate that heat transfer to the head is also under for regulation of Tth at low Ta. active control, very probably owing to temperature- sensitive alteration of hemolymph circulation. As a result, head temperature (Th) is strikingly elevated above Ta Key words: Anax junius, Anisoptera, body temperature, circulatory during endothermic warm-up and flight. Furthermore, control, dragonfly, green darner, heat exchange, thermoregulation. Introduction Numerous insects regulate Tth (most recently and The primary aim of this study is to investigate the sources comprehensively reviewed by Heinrich, 1993), among them of variation of Th, its mechanism of control and its responses the subject of this paper Anax junius (Drury) (Heinrich and to environmental temperature and internal variables in A. -
Green Darner Anax Junius
TEXAS PARKS AND WILDLIFE Common Green Darner Anax junius TPWD PHOTO Dragonflies haven’t changed much over the last 300 million years. Today they are one of the most easily recognized of all insects. The common green darner is the largest, most abundant and widespread dragonfly species in North America. Common Green Darner Anax junius APPEARANCE HABITAT Length: 3 inches (7.6cm) Common green darners prefer permanent and temporary ponds, Wingspan: 4.5 inches (11.4cm) lakes, bays, estuaries and slow-moving streams and riparian areas (land adjacent to a body of water). Distinguishing Characteristics • Green head and thorax • Abdomen is yellow and brown on females; long, slender and BEHAVIOR bluish on males • Large compound eyes During the reproductive stage, the common green darner seeks • Strong jaws fresh water ponds. Males return first, then females who have • Spiny legs already developed a batch of eggs. If the female is receptive, • Wings clear with yellowish tint toward tips and strongly mating begins. They often mate in flight. Immediately after mat- veined with net-like pattern ing, common green darner females lay their eggs, one at a time, in tiny slits in submerged aquatic plants. Males work hard to protect their territories. Some have been clocked chasing intrud- LIFE HISTORY ers away at 35 miles per hour. Common green darners are one of the few dragonflies that migrate in the spring and fall. Range: North America, the West Indies and Scientists believe that they migrate with seasonal warm fronts. South America Diet: Larvae eat fish eggs, tadpoles and other small aquatic animals.