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New Data on Four Species of Ammonites Created by V. Uhlig in 1883 (Ammonitida, Turrilitina, Hamulinoidea)

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New Data on Four Species of Ammonites Created by V. Uhlig in 1883 (Ammonitida, Turrilitina, Hamulinoidea) See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/275330954 New data on four species of ammonites created by V. Uhlig in 1883 (Ammonitida, Turrilitina, Hamulinoidea) Article · January 2014 CITATIONS READS 0 143 1 author: Jean Vermeulen 52 PUBLICATIONS 514 CITATIONS SEE PROFILE All in-text references underlined in blue are linked to publications on ResearchGate, Available from: Jean Vermeulen letting you access and read them immediately. Retrieved on: 13 November 2016 Ann. Mus. Hist. nat. Nice, XXIX : 1-25, 2014 New data on four species of ammonites created by V. Uhlig in 1883 (Ammonitida, Turrilitina, Hamulinoidea) Nouvelles données sur quatre espèces d'ammonites créées par V. Uhlig en 1883 (Ammonitida, Turrilitina, Hamulinoidea) 1 2 3 JEAN VERMEULEN , ALEXANDER LUKENEDER , CHRISTIAN MEISTER 4 & ZDENEK VASICEK ___________________________________________________________________________ Abstract — This work presents paleontological and Résumé — Ce travail apporte des précisions stratigraphical precise details about four species of paléontologiques et stratigraphiques sur quatre espèces ammonites created by Uhlig. Anahamulina lorioli d'Ammonites créées par Uhlig. Anahamulina lorioli (UHLIG, 1883) is a Barremian species of the lower half (UHLIG, 1883) est une espèce Barrémienne de la part of the Zone of Kotetishvilia nicklesi; it is moitié inférieure de la Zone à Kotetishvilia nicklesi ; characterized by a significant development of the elle se caractérise par un développement important de dorsolateral tubercles on the retroversum. In the la tuberculation latéro-dorsale sur le retroversum. genus Amorina VERMEULEN, 2005 are distinguished Dans le genre Amorina VERMEULEN, 2005 sont three groups of species which are characterized by distingués trois groupes d'espèces qui se distinguent their ornamental characteristics and their par leurs caractéristiques ornementales et par la stratigraphical ranges. Amorina hoheneggeri (UHLIG, répartition stratigraphique de leurs espèces. Amorina 1883) is a Barremian species which has kept primitive hoheneggeri (UHLIG, 1883) est une espèce characters of the older Amorina group. It is barrémienne qui a conservé des caractères primitifs characterized by a little expressed constriction on the présents chez les espèces du groupe le plus ancien du bend and by a pair of constrictions bordered ahead by genre Amorina VERMEULEN, 2005 ; elle se caractérise a varix, on the oldest half of the retroversum. Amorina par une constriction peu exprimée sur le coude et par subcincta (UHLIG, 1883) is a Lower Barremian species une paire de constrictions bordées en avant par une which is classified in the group of the species close to varice, sur la moitié la plus âgée du retroversum. the type species of the genus; it is characterized by a Amorina subcincta (UHLIG, 1883) est une espèce du regular ribbing on all the shell and by the presence of Barrémien inférieur qui se classe dans le groupe des a constriction, bordered by two varixes, which is espèces proches de l'espèce-type du genre ; elle se located between the end of the shaft and the beginning caractérise par une costulation régulière sur toute la of the retroversum. Terrestrina quenstedti (UHLIG, coquille et par la présence d'une constriction, bordée 1883) is present in the Barremian 1, sensu Vermeulen, par deux varices, qui se situe entre la fin de la hampe 2007, p. 165, in Taveraidiscus hugii and Kotetishvilia et le début du retroversum. Terrestrina quenstedti nicklesi Zones. It is characterized by a very thin and (UHLIG, 1883) est présente dans le Barremian 1, sensu regular ribbing, with regular spaced out ribs, and by Vermeulen, 2007, p. 165, dans les Zones à the absence of varix and constriction on the shaft, the Taveraidiscus hugii et à Kotetishvilia nicklesi ; elle se bend and the known part of the retroversum. caractérise par une costulation très fine, régulière, d'espacement régulier, et par l'absence de varice et de constriction sur la hampe, sur le coude et sur la partie connue du retroversum. Key words — Ammonitida, Turrilitina, Mots-clés — Ammonitida, Turrilitina, Hamulinoidea, Hamulinoidea, Ptychoceratoidea, Anahamulinidae, Ptychoceratoidea, Anahamulinidae, Crétacé inférieur. Lower Cretaceous. ___________________________________________________________________________ 1234 1Grand rue, 04330 Barrême, France. 2 Natural History Museum, Geological-Paleontological Department, Burgring 7 A, 1010 Wien, Austria. 3 Natural History Museum, Department of Geology and Paleontology, 1 route de Malagnou, CP 6434, CH-1211, Geneva 6, Switzerland. 4 Institute of Geonics, Academy of Sciences of the Czech Republic, Studentska 1768, CZ-708 00 Ostrava- Poruba, Czech Republic. New data on four species of ammonites created by V. Uhlig in 1883 (Ammonitida, Turrilitina, Hamulinoidea) INTRODUCTION In 1883, Uhlig created new species of hamulinid shaped ammonites which are classified today in the family Anahamulinidae BREISTROFFER, 1952. Some of them were revised by Sarasin and Schöndelmayer in 1902, and then more recently by Vasicek in 1972 and 1973. Despite these works, many species of Anahamulinidae created by Uhlig remain to be revised. The aim of this work is to bring new paleontological and stratigraphical data to the species Anahamulina lorioli (UHLIG, 1883), Amorina hoheneggeri (UHLIG, 1883), Amorina subcincta (UHLIG, 1883) and Terrestrina quenstedti (UHLIG, 1883). 1. BIOSTRATIGRAPHY The biostratigraphical framework of late Hauterivian used herein is that proposed by Vermeulen et al. (2013a) and, for the Lower Barremian, is that proposed by Vermeulen et al. (2014). These biozonations are close to that proposed by the Kilian Group (2006) in Neuchâtel, Switzerland. 1.1. Late Hauterivian The index species of the zones of Late Hauterivian are Subsaynella sayni (PAQUIER, 1900), Plesiospitidiscus ligatus (ORBIGNY, 1841), Balearites balearis (NOLAN, 1894), Ropoloceras seitzi (SARKAR, 1955), Balearites mortilleti (PICTET & LORIOL, 1858) and Pseudothurmannia angulicostata SPATH, 1923. Table 1: Biozonation of Late Hauterivian. Zones Subzones and Horizons Pseudothurmannia angulicostata Balearites mortilleti Ropoloceras seitzi Balearites krenkeli Balearites balearis Balearites binelli Balearites balearis Plesiospitidiscus ligatus Subsaynella begudensis Subsaynella sayni 2 VERMEULEN J., LUKENEDER A, MEISTER C & VASICEK Z. 1.2. Lower Barremian The index species of the zones of Lower Barremian are Taveraidiscus hugii (OOSTER, 1860), Kotetishvilia nicklesi (HYATT, 1903), Nicklesia pulchella (ORBIGNY, 1841), Kotetishvilia compressissima (ORBIGNY, 1841) and Coronites darsi VERMEULEN, 1995. Table 2: Biozonation of Lower Barremian. Zones Subzones and Horizons Holcodiscus uhligi Coronites darsi Subtorcapella defayae Kotetishvilia compressissima Holcodiscus fallax Leptohamulina lukenederi Nicklesia pulchella Kotetishvilia nicklesi Anahamulina lorioli Psilotissotia colombiana Taveraidiscus hugii Taveraidiscus hugii 3. SYSTEMATIC STUDY Order Ammonitida ZITTEL, 1884 Suborder Turrilitina BEZNOZOV & MICHAILOVA, 1983 This suborder includes the whole heteromorphic ammonites, possessing a bifid, or dissymmetrical trifid lateral lobe (Vermeulen, 2006), which is always distinguished from those with trifid symmetrical lateral lobe, classified in Ancyloceratina WIEDMANN, 1966. The super families, Turrilitoidea GILL, 1871, Hamulinoidea GILL, 1871, and Scaphitoidea GILL, 1871 nom. transl. WRIGHT & WRIGHT, 1951 are classified in the suborder Turrilitina BEZNOZOV & MICHAILOVA, 1983. 3 New data on four species of ammonites created by V. Uhlig in 1883 (Ammonitida, Turrilitina, Hamulinoidea) Superfamily Hamulinoidea GILL, 1871 nom. transl. VERMEULEN, LAZARIN, LÉPINAY, LEROY & MASCARELLI, 2014 Type genus: Hamulina ORBIGNY, 1850. Some precisions on the nomenclatural status of the genus Ptychoceras ORBIGNY, 1842 and of the derived taxa are given in Vermeulen et al. (2013a); they imply the nomenclatural status nomen nudum of the genus Ptychoceras ORBIGNY, 1842. The now used superfamily Hamulinoidea GILL, 1871 nom. transl. includes the families Ptychoceratidae GILL, 1871 nomen nudum, Hamulinidae GILL, 1871, Anahamulinidae BREISTROFFER, 1952, and Macroscaphitidae HYATT, 1900. The family Hamulinidae GILL, 1871 is divided into two subfamilies, the Megacrioceratinae VERMEULEN, 2006 nom. transl. VERMEULEN, 2010, and the Hamulininae GILL, 1871 nom. transl. VERMEULEN, CECCA & KRUTA, 2007. The subfamily Megacrioceratinae VERMEULEN, 2006, derives most probably from the Crioceratitidae GILL, 1871 and it is the root (Vermeulen, 2009, 2010) of the Hamulininae GILL, 1871, of the Macroscaphitidae HYATT, 1900, and of the Anahamulinidae BREISTROFFER, 1952. Family Anahamulinidae BREISTROFFER, 1952 Type genus: Anahamulina HYATT, 1900. The four subfamilies Anahamulininae BREISTROFFER, 1952 nom. transl. VERMEULEN, CECCA & KRUTA, 2007, Ptychohamulininae VERMEULEN, 2010, Leptohamulininae VERMEULEN, 2010 and Pacaudininae VERMEULEN, LAZARIN, LÉPINAY, LEROY & MASCARELLI, 2013 divide the family Anahamulinidae BREISTROFFER, 1952 (Vermeulen et al. 2007; Vermeulen, 2010; Vermeulen et al. 2013b). Stratigraphical ranges: the representatives of the family Anahamulinidae BREISTROFFER, 1952 appear in the Upper Hauterivian and disappear in the Upper Albian. Distinctive criteria: the family Anahamulinidae BREISTROFFER, 1952 is distinguished from the family Hamulinidae GILL, 1871 by its species of small sizes, non tuberculated or less tuberculated, and by its stratigraphical range, from
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