DOCSLIB.ORG

Nomenclatural Changes in Coleus and Plectranthus (Lamiaceae): a Tale of More Than Two Genera

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Nomenclatural Changes in Coleus and Plectranthus (Lamiaceae): a Tale of More Than Two Genera Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera Article Published Version Creative Commons: Attribution 4.0 (CC-BY) Open access Paton, A., Mwyanyambo, M., Govaerts, R. H.A., Smitha, K., Suddee, S., Phillipson, P. B., Wilson, T. C., Forster, P. I. and Culham, A. (2019) Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera. PhytoKeys, 129. pp. 1-158. ISSN 1314–2003 doi: https://doi.org/10.3897/phytokeys.129.34988 Available at http://centaur.reading.ac.uk/86484/ It is advisable to refer to the publisher’s version if you intend to cite from the work. See Guidance on citing . To link to this article DOI: http://dx.doi.org/10.3897/phytokeys.129.34988 Publisher: Pensoft All outputs in CentAUR are protected by Intellectual Property Rights law, including copyright law. Copyright and IPR is retained by the creators or other copyright holders. Terms and conditions for use of this material are defined in the End User Agreement . www.reading.ac.uk/centaur CentAUR Central Archive at the University of Reading Reading’s research outputs online A peer-reviewed open-access journal PhytoKeys 129:Nomenclatural 1–158 (2019) changes in Coleus and Plectranthus: a tale of more than two genera 1 doi: 10.3897/phytokeys.129.34988 RESEARCH ARTICLE http://phytokeys.pensoft.net Launched to accelerate biodiversity research Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera Alan J. Paton1, Montfort Mwanyambo2, Rafaël H.A. Govaerts1, Kokkaraniyil Smitha3, Somran Suddee4, Peter B. Phillipson5, Trevor C. Wilson6, Paul I. Forster7, Alastair Culham8 1 Science Directorate, Royal Botanic Gardens Kew, Richmond, Surrey,TW9 3AB, UK 2 National Herbarium and Botanic Gardens of Malawi, Zomba, Malawi 3 Department of Botany, University of Calicut, Kerala- 673 635, India 4 The Forest Herbarium (BKF), Department of National Parks, Wildlife and Plant Conservation, Chatuchack, Bangkok 10900,Thailand 5 Missouri Botanical Garden, P.O. Box 299, St. Louis, MO 63166, U.S.A. and Institut de Systématique, Evolution et Biodiversité (UMR 7205 –CNRS MNHN UPMC EPHE), Muséum national d’Histoire naturelle, Sorbonne Universités, C.P. 39, rue Cuvier 57, 75231 Paris CEDEX 05, France 6 National Herbarium of New South Wales, Royal Botanic Gardens & Domain Trust, Mrs Macquaries Road, Sydney, NSW 2000, Australia and School of Life and Environmental Sciences, Science Road, University of Sydney, NSW 2006, Australia 7 Queensland Herbarium, Department of Environment & Science, Brisbane Botanic Gardens, Mount Coot-tha Road, Toowong, Qld 4066, Australia 8 School of Biological Sciences, Uni- versity of Reading, Whiteknights, Reading, UK Corresponding author: Alan J. Paton ([email protected]) Academic editor: Eberhard Fischer | Received 29 March 2019 | Accepted 18 June 2019 | Published 23 August 2019 Citation: Paton AJ, Mwanyambo M, Govaerts RHA, Smitha K, Suddee S, Phillipson PB, Wilson TC, Forster PI, Culham A (2019) Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera. PhytoKeys 129: 1–158. https://doi.org/10.3897/phytokeys.129.34988 Abstract A synopsis of the genera Coleus Lour, Equilabium A.J.Paton, Mwany. & Culham and Plectranthus L’Hér. (Lamiaceae, Tribe Ocimeae, Subtribe Plecranthinae) is presented. Generic delimitation follows a recently pub- lished molecular phylogeny which identifiedColeus as the sister of the remaining genera of Subtribe Plectran- thinae; Plectranthus as sister to Tetradenia Benth. and Thorncroftia N.E.Br., and a separate phylogenetically distinct genus Equilabium comprising species previously placed in Plectranthus. In this treatment, 294 species of Coleus, 42 of Equilabium, and 72 of Plectranthus are recognized. All but one of the combinations in Equi- labium are new as only the genus and type species have been previously published. Two-hundred and twelve names are changed to combinations in Coleus from Plectranthus, Pycnostachys Hook. and Anisochilus Benth. Copyright Alan J. Paton et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 2 Alan J. Paton et al. / PhytoKeys 129: 1–158 (2019) Keywords Anisochilus, Equilabium, generic delimitation, nomenclature, Ocimeae, Plectranthinae, Pycnostachys, Solenostemon Table of contents Introduction ............................................................................................................. 2 Taxonomic history .................................................................................................... 3 Methods ................................................................................................................... 5 Results and discussion .............................................................................................. 7 Taxonomic account .................................................................................................. 8 Subtribe Plectranthinae ........................................................................................ 9 Key to the genera of the Plectranthinae ................................................................ 9 Synopsis of genera and species of Coleus, Equilabium and Plectranthus.................... 10 Genus Coleus ...................................................................................................... 10 Genus Equilabium ............................................................................................ 111 Genus Plectranthus ........................................................................................... 127 Unplaced names ............................................................................................... 147 Excluded names ............................................................................................... 150 Acknowledgements ............................................................................................... 155 References ............................................................................................................ 156 Introduction Plectranthus L’Hér. is a widely used horticultural and medicinal plant genus (van Jaars- veld 2007, Lukhoba et al. 2006, Rice et al. 2011). It belongs in tribe Ocimeae, subtribe Plectranthinae, an Old World tropical group of around 450 species. Current taxonomic treatments generally follow Harley et al. (2004) by including the formerly recognized Coleus Lour. and Solenostemon Thonn. withinPlectranthus . However, within horticultural literature, Solenostemon is still frequently recognized (e.g. RHS (2017), Shepherd and Maybry (2016)). The commonly cultivated plant shown in Fig. 1 is thus variously known as Plectranthus scutellarioides (L.) R.Br or Solenostemon scutellarioides (L.) Codd., but “co- leus” is frequently used as a common name for this species, increasing the confusion. Recent phylogenetic work has shown that the circumscription of Plectranthus needs to be viewed in the context of related genera in Tribe Ocimeae, subtribe Plec- tranthinae. Paton et al. (2018) demonstrate that a clade containing the type of Coleus (C. amboinicus Lour. ≡ P. amboinicus (Lour.) Spreng.) is sister to the other genera of the Plectranthinae, and the type species of Plectranthus, P. fruticosus L’Hér., belongs in a clade which is sister to Thorncroftia N.E.Br and Tetradenia Benth. A further clade containing species that were previously placed in Plectranthus was recognized by Paton et al. (2018) as Equilabium Mwany., A.J.Paton & Culham (Fig. 2). The aim of this paper is to provide the necessary nomenclatural changes consistent with the recognition of Plectranthus, Coleus and Equilabium as distinct genera follow- ing phylogenetic results provided by Paton et al. (2018). Nomenclatural changes in Coleus and Plectranthus: a tale of more than two genera 3 Figure 1. A Display of cultivated Coleus scutellarioides (L.) Benth. in the Real Jardin Botánico, Madrid. B Close up of the cultivar ‘Campfire’, winner of the Royal Horticultural Society’s peoples choice award at Wisley 2017 (Photo: A.Paton). Taxonomic history Plectranthus was described by L’Héritier in 1788 (L’Héritier 1788) and Coleus by Loureiro in 1790 (de Loureiro 1790). In previous global reviews of the Lamiaceae, e.g. Bentham (1832, 1848, 1876) and Briquet (1897), Coleus was treated as distinct from Plectranthus by having the 4 stamens fused together rather than free to their base. Morton (1962) recognized that the character of the fused stamens was more plastic than previously thought and merged Coleus into Plectranthus, as first suggested by Brown (1810). However, Morton also placed weight on calyx characteristics leading him to recognize several genera based on calyx morphology including Solenostemon, Isodictyophorus A.Chev, Leocus A.Chev., Englerastrum Briq. and Neohyptis J.K.Morton. The merging ofColeus into Plectranthus has been followed by most authors since Morton’s account, for example, Keng (1978), covering Malesian species, Blake (1971, Australia), Codd (1985, southern Africa), Hedge et al. (1998, Madagascar), Suddee et al. (2004, Indochina), Forster (2011, Australia) and Paton et al. (2009, 2013, eastern and southern tropical Africa). Furthermore, although Codd (1985) and Morton (2006) also maintained Solenostemon, Keng (1978), Hedge et al. (1998) and subsequent authors recognised Solenostemon as synonymous with Plectranthus.
Load more

Recommended publications
  • "National List of Vascular Plant Species That Occur in Wetlands: 1996 National Summary."
    Intro 1996 National List of Vascular Plant Species That Occur in Wetlands The Fish and Wildlife Service has prepared a National List of Vascular Plant Species That Occur in Wetlands: 1996 National Summary (1996 National List). The 1996 National List is a draft revision of the National List of Plant Species That Occur in Wetlands: 1988 National Summary (Reed 1988) (1988 National List). The 1996 National List is provided to encourage additional public review and comments on the draft regional wetland indicator assignments. The 1996 National List reflects a significant amount of new information that has become available since 1988 on the wetland affinity of vascular plants. This new information has resulted from the extensive use of the 1988 National List in the field by individuals involved in wetland and other resource inventories, wetland identification and delineation, and wetland research. Interim Regional Interagency Review Panel (Regional Panel) changes in indicator status as well as additions and deletions to the 1988 National List were documented in Regional supplements. The National List was originally developed as an appendix to the Classification of Wetlands and Deepwater Habitats of the United States (Cowardin et al.1979) to aid in the consistent application of this classification system for wetlands in the field.. The 1996 National List also was developed to aid in determining the presence of hydrophytic vegetation in the Clean Water Act Section 404 wetland regulatory program and in the implementation of the swampbuster provisions of the Food Security Act. While not required by law or regulation, the Fish and Wildlife Service is making the 1996 National List available for review and comment.
    [Show full text]
  • Micropropagação E Produção De Monoterpenos E Sesquiterpenos Em Plectranthus Ornatus Codd
    1 UNIVERSIDADE ESTADUAL DE FEIRA DE SANTANA PROGRAMA DE PÓS-GRADUAÇÃO EM BIOTECNOLOGIA HELNA CÉLIA PASSINHO SOARES MICROPROPAGAÇÃO E PRODUÇÃO DE MONOTERPENOS E SESQUITERPENOS EM PLECTRANTHUS ORNATUS CODD Feira de Santana, BA 2010 2 HELNA CÉLIA PASSINHO SOARES MICROPROPAGAÇÃO E PRODUÇÃO DE MONOTERPENOS E SESQUITERPENOS EM PLECTRANTHUS ORNATUS CODD Tese apresentada ao Programa de Pós-graduação em Biotecnologia, da Universidade Estadual de Feira de Santana como requisito parcial para obtenção do título de Doutora em Biotecnologia. Orientadores: Profª. Drª. Juceni Pereira Lima David Prof. Dr José Raniere Ferreira de Santana Feira de Santana, BA 2010 3 Dedico este trabalho primeiramente a Deus, pois sem Ele, nada seria possível. Ao meu saudoso e amado esposo Gilson (in memorian) pelo grande companheiro que foi, pelo exemplo de fé e perseverança, pelo apoio, estímulo e dedicação que sempre me dedicou. Aos meus pais Clóvis e Dulce; pelo esforço, dedicação e compreensão, em todos os momentos desta e de outras caminhadas. Em especial, a minha amada Nathália, presente em tudo na minha vida. Razão de tudo em minha vida. Te amo Filha!! 4 AGRADECIMENTOS A Deus, único e digno de receber honras e glória. Deus eterno, imortal, invisível, mais real. A Ele o meu louvor e gratidão. A Universidade Estadual de Feira de Santana, especialmente ao Programa de pós-graduação em Biotecnologia (UEFS/Fiocruz), pela oportunidade da realização do curso de Doutorado. A FAPESB pela concessão da bolsa de estudos. A Profª Juceni David e ao Prof. Dr. Raniere pela confiança, pela amizade e orientação que auxiliaram na concretização deste estudo. Ao secretário do PPGBiotec, Helton pela disposição, educação e gentiliza que sempre nos acolheu.
    [Show full text]
  • Species ANALYSIS International Journal for Species ISSN 2319 – 5746 EISSN 2319 – 5754
    Species ANALYSIS International Journal for Species ISSN 2319 – 5746 EISSN 2319 – 5754 Diversity and therapeutic potentiality of the family Lamiaceae in Karnataka State, India: An overview Rama Rao V1҉, Shiddamallayya N2, Kavya N3, Kavya B4, Venkateshwarlu G5 1. Research Officer (Botany), Survey of Medicinal Plants Unit, National Ayurveda Dietetics Research Institute (CCRAS), Govt. Central Pharmacy Annexe, Ashoka Pillar, Jayangar, Bangalore-560011, India. 2. Assistant Research Officer (Botany), Survey of Medicinal Plants Unit, National Ayurveda Dietetics Research Institute (CCRAS), Govt. Central Pharmacy Annexe, Ashoka Pillar, Jayangar, Bangalore-560011, India. 3. Senior Research Fellow (Ayurveda), National Ayurveda Dietetics Research Institute (CCRAS), Govt. Central Pharmacy Annexe, Ashoka Pillar, Jayangar, Bangalore-560011, India. 4. Junior Research Fellow (Botany), National Ayurveda Dietetics Research Institute (CCRAS), Govt. Central Pharmacy Annexe, Ashoka Pillar, Jayangar, Bangalore-560011, India. 5. Research Officer (Scientist-3) in-charge, National Ayurveda Dietetics Research Institute (CCRAS), Govt. Central Pharmacy Annexe, Ashoka Pillar, Jayangar, Bangalore-560011, India. ҉Corresponding author: Survey of Medicinal Plants Unit, National Ayurveda Dietetics Research Institute (CCRAS), Govt. Central Pharmacy Annexe, Ashoka Pillar, Jayangar, Bangalore-560011, India, e-mail: [email protected] Publication History Received: 25 November 2014 Accepted: 11 January 2015 Published: 4 March 2015 Citation Rama Rao V, Shiddamallayya N, Kavya N, Kavya B, Venkateshwarlu G. Diversity and therapeutic potentiality of the family Lamiaceae in Karnataka State, India: An overview. Species, 2015, 13(37), 6-14 ABSTRACT The objective of the present study was to review the potential medicinal plants of Lamiaceae distributed throughout the state of Karnataka, India. Lamiaceae, also called as mint family is one of the largest families including herbs or shrubs often with aroma.
    [Show full text]
  • Efeito Do Extrato De Plectranthus Barbatus (Andr.) Benth No Desempenho Reprodutivo De Rattus Novergicus (Berkenhout, 1769)
    Biotemas, 20 (2): 49-58, junho de 2007 Avifauna do município de Araricá, RS 49 ISSN 0103 – 1643 Efeito do extrato de Plectranthus barbatus (Andr.) Benth no desempenho reprodutivo de Rattus novergicus (Berkenhout, 1769) Thelma Duarte Delgado Brandolt1* Cinthia Cardoso Rodrigues1 Sandro Márcio Nunes Ferrão2 Gisélida Maria Baquini da Silva3 ¹Faculdade de Filosofia, Ciências e Letras (FAFIUR), Curso de Ciências Biológicas ²Hospital Veterinário da Faculdade de Zootecnia, Veterinária e Agronomia (FZVA) Biotério e Laboratório de Farmacologia ³Hospital Veterinário da FZVA, Laboratório de Biotecnologia da Reprodução PUCRS, Campus Uruguaiana, BR 472 Km 7, Caixa Postal 249 CEP 97500-970, Uruguaiana – RS *Autora para Correspondência [email protected] Submetido em 10/08/2006 Aceito para publicação em 12/03/2007 Resumo A suposta utilização do extrato de Plectranthus barbatus para fins abortivos motivou esta investigação sobre a ocorrência dos possíveis efeitos tóxicos do Falso-boldo na viabilidade dos embriões e da prole de ratas Wistar. O experimento contou com vinte e sete fêmeas e nove machos, divididos em três grupos. Os chás foram preparados pelo método de infusão, utilizando-se folhas de plantas adultas obtidas em quintais residenciais. A partir do primeiro dia de acasalamento os animais foram submetidos a diferentes tratamentos: o (T1) recebeu água, o (T2) recebeu a dose de chá considerada coloquial e o (T3) recebeu uma dose 200% maior. Para avaliar a qualidade e verificar o estágio de desenvolvimento embrionário, no quarto dia de acasalamento, seis fêmeas de cada grupo foram submetidas a uma laparotomia abdominal mediana para que se procedesse a um lavado uterino que, depois de coletado, foi submetido a análise laboratorial.
    [Show full text]
  • Lamiales Newsletter
    LAMIALES NEWSLETTER LAMIALES Issue number 4 February 1996 ISSN 1358-2305 EDITORIAL CONTENTS R.M. Harley & A. Paton Editorial 1 Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3AE, UK The Lavender Bag 1 Welcome to the fourth Lamiales Universitaria, Coyoacan 04510, Newsletter. As usual, we still Mexico D.F. Mexico. Tel: Lamiaceae research in require articles for inclusion in the +5256224448. Fax: +525616 22 17. Hungary 1 next edition. If you would like to e-mail: [email protected] receive this or future Newsletters and T.P. Ramamoorthy, 412 Heart- Alien Salvia in Ethiopia 3 and are not already on our mailing wood Dr., Austin, TX 78745, USA. list, or wish to contribute an article, They are anxious to hear from any- Pollination ecology of please do not hesitate to contact us. one willing to help organise the con- Labiatae in Mediterranean 4 The editors’ e-mail addresses are: ference or who have ideas for sym- [email protected] or posium content. Studies on the genus Thymus 6 [email protected]. As reported in the last Newsletter the This edition of the Newsletter and Relationships of Subfamily Instituto de Quimica (UNAM, Mexi- the third edition (October 1994) will Pogostemonoideae 8 co City) have agreed to sponsor the shortly be available on the world Controversies over the next Lamiales conference. Due to wide web (http://www.rbgkew.org. Satureja complex 10 the current economic conditions in uk/science/lamiales). Mexico and to allow potential partici- This also gives a summary of what Obituary - Silvia Botta pants to plan ahead, it has been the Lamiales are and some of their de Miconi 11 decided to delay the conference until uses, details of Lamiales research at November 1998.
    [Show full text]
  • Universidade Federal Do Ceará Centro De Ciências Departamento De Química Orgânica E Inorgânica Programa De Pós-Graduação Em Química
    UNIVERSIDADE FEDERAL DO CEARÁ CENTRO DE CIÊNCIAS DEPARTAMENTO DE QUÍMICA ORGÂNICA E INORGÂNICA PROGRAMA DE PÓS-GRADUAÇÃO EM QUÍMICA TAYNARA SIMÃO MATOS INVESTIGAÇÃO DOS CONSTITUINTES QUÍMICOS DO CHÁ DE Plectranthus ornatus Codd FORTALEZA 2019 TAYNARA SIMÃO MATOS INVESTIGAÇÃO DOS CONSTITUINTES QUÍMICOS DO CHÁ DE Plectranthus ornatus Codd Dissertação apresentada ao Programa de Pós- Graduação em Química da Universidade Federal do Ceará, como requisito parcial à obtenção do título de Mestre em Química. Área de concentração: Química Orgânica. Orientador (a): Profa. Dra. Otília Deusdênia Loiola Pessoa. FORTALEZA 2019 TAYNARA SIMÃO MATOS INVESTIGAÇÃO DOS CONSTITUINTES QUÍMICOS DO CHÁ DE Plectranthus ornatus Codd Dissertação apresentada ao Programa de Pós- Graduação em Química da Universidade Federal do Ceará, como requisito parcial à obtenção do título de Mestre em Química. Área de concentração: Química Orgânica. Aprovada em: 26/ 07/ 2019. BANCA EXAMINADORA ________________________________________ Prof. Dra. Otília Deusdênia Loiola Pessoa (Orientador) Universidade Federal do Ceará (UFC) _________________________________________ Prof. Dr. Edilberto Rocha Silveira Universidade Federal do Ceará (UFC) _________________________________________ Prof. Dra. Jane Eire Silva Alencar de Menezes Universidade Estadual do Ceará (UECE) À Deus. À minha mãe, Lourdes. Ao meu pai, Antonio (in memorian). Aos meus irmãos, Thamyres, Thalyta, Tallys. Ao meu sobrinho, Bryan. AGRADECIMENTOS Agradeço primeiramente à Deus por sempre estar ao meu lado me guiando, dando novas possibilidades e mostrando que Nele tudo posso. À Lourdes Francisca Simão, minha mãe, que sempre me incentivou, apoiou e foi minha fonte de paz. Ao Antonio Ferreira Matos, meu pai, que mesmo não estando mais neste plano, sinto que está comigo, sendo minha fortaleza. Aos meus irmãos Thamyres, Thalyta e Tallys, assim como ao meu sobrinho Bryan, por serem fontes de alegria e distração nos momentos de estresse.
    [Show full text]
  • COLEUS 'FAIRWAY RUBY' Solenostemon Scutellarioides 'Fairway Ruby' Characteristics Culture Noteworthy Characteristics Problems Ga
    COLEUS 'FAIRWAY RUBY' Solenostemon scutellarioides 'Fairway Ruby' Characteristics Zone: 10 to 11 Soil: Moist, well-drained Height: 10 to 15 Inches Water: Medium Width: 12 Inches Maintenance: Low Bloom Time: Flowers not showy Flower: Insignificant Bloom Description: Blue to white Leaf: Colorful Sun: Part sun Tolerate: Shade Culture Winter hardy to USDA Zones 10-11. In other zones, grow as garden annuals or in pots that can be overwintered indoors or as houseplants. Outdoors, grow in moist, organically rich, loose soils in part shade. Soils must not be allowed to dry out. ‘Fairway Ruby’ coleus produces foliage with bright red center and wide off-white margin with green edging. Produces tiny blue or white flowers, 1/2" long anytime throughout year. Pinch out to maintain plant health, foliage color, and attractive, compact plants. Does not tolerate water logging. Water moderately when in growth and apply a balanced liquid fertilizer at alternate waterings. When in growth, apply a high nitrogen fertilizer every 2 weeks. Keep just moist. Plants grown in too much shade may become leggy. Noteworthy Characteristics Plectranthus scutellarioides, commonly known as coleus, is a tropical evergreen tender perennial that has been a popular foliage plant since at least Victorian times. It is native to Southeast Asia and Malaysia. It has been assiduously hybridized over the years into a very large number of vegetatively propagated and seed propagated strains with an almost infinite number of leaf color combinations including most colors of the spectrum except true blue. Cultivars range in size from dwarf 6” tall plants to large mounded 36” tall plants.
    [Show full text]
  • Palynological Evolutionary Trends Within the Tribe Mentheae with Special Emphasis on Subtribe Menthinae (Nepetoideae: Lamiaceae)
    Plant Syst Evol (2008) 275:93–108 DOI 10.1007/s00606-008-0042-y ORIGINAL ARTICLE Palynological evolutionary trends within the tribe Mentheae with special emphasis on subtribe Menthinae (Nepetoideae: Lamiaceae) Hye-Kyoung Moon Æ Stefan Vinckier Æ Erik Smets Æ Suzy Huysmans Received: 13 December 2007 / Accepted: 28 March 2008 / Published online: 10 September 2008 Ó Springer-Verlag 2008 Abstract The pollen morphology of subtribe Menthinae Keywords Bireticulum Á Mentheae Á Menthinae Á sensu Harley et al. [In: The families and genera of vascular Nepetoideae Á Palynology Á Phylogeny Á plants VII. Flowering plantsÁdicotyledons: Lamiales (except Exine ornamentation Acanthaceae including Avicenniaceae). Springer, Berlin, pp 167–275, 2004] and two genera of uncertain subtribal affinities (Heterolamium and Melissa) are documented in Introduction order to complete our palynological overview of the tribe Mentheae. Menthinae pollen is small to medium in size The pollen morphology of Lamiaceae has proven to be (13–43 lm), oblate to prolate in shape and mostly hexacol- systematically valuable since Erdtman (1945) used the pate (sometimes pentacolpate). Perforate, microreticulate or number of nuclei and the aperture number to divide the bireticulate exine ornamentation types were observed. The family into two subfamilies (i.e. Lamioideae: bi-nucleate exine ornamentation of Menthinae is systematically highly and tricolpate pollen, Nepetoideae: tri-nucleate and hexa- informative particularly at generic level. The exine stratifi- colpate pollen). While the
    [Show full text]
  • Vascular Plants Diversity and Ethnobotany With
    VASCULAR PLANTS DIVERSITY AND ETHNOBOTANY WITH EMPHASIS TO TRADITIONAL MEDICINAL AND WILD EDIBLE PLANTS IN DUGDA DAWA DISTRICT OF BORANA ZONE, OROMIA REGIONAL STATE, ETHIOPIA Mersha Ashagre Eshete Addis Ababa University Addis Ababa, Ethiopia April 2017 VASCULAR PLANTS DIVERSITY AND ETHNOBOTANY WITH EMPHASIS TO TRADITIONAL MEDICINAL AND WILD EDIBLE PLANTS IN DUGDA DAWA DISTRICT OF BORANA ZONE, OROMIA REGIONAL STATE, ETHIOPIA Mersha Ashagre Eshete A Thesis Submitted to The Department of Plant Biology and Biodiversity Management Presented in Fulfillment of the Requirements for the Degree of Doctor of Philosophy (Plant Biology and Biodiversity Management) Addis Ababa University Addis Ababa, Ethiopia April 2017 i ADDIS ABABA UNIVERSITY GRADUATE PROGRAMMES This is to certify that the thesis prepared by Mersha Ashagre Eshete, entitled: “Vascular Plants Diversity and Ethnobotany with Emphasis to Traditional Medicinal and Wild Edible Plants in Dugda Dawa District of Borana Zone, Oromia Regional State, Ethiopia”, and submitted in fulfillment of the requirements for the Degree of Doctor of Philosophy (Plant Biology and Biodiversity Management) complies with the regulations of the University and meets the accepted standards with respect to originality and quality. Signed by Research Supervisors: Name Signature Date 1. _____________________ _________________ _____________ 2.______________________ _________________ _____________ 3._____________________ _________________ ______________ 4.____________________ __________________ _______________ _____________________
    [Show full text]
  • 398-IJBCS-Article
    Available online at http://ajol.info/index.php/ijbcs Int. J. Biol. Chem. Sci. 3(5): 1100-1113, October 2009 ISSN 1991-8631 Original Paper http://indexmedicus.afro.who.int Morphoagronomical characterization of Solenostemon rotundifolius (Poir. J. K. Morton) (Lamiaceae) germplasm from Burkina Faso Romaric Kiswendsida NANEMA *, Ernest Renan TRAORE, Pauline BATIONO/KANDO and Jean-Didier ZONGO Laboratoire de Génétique et de Biotechnologie Végétales, Unité de Formation et de Recherche en Sciences de la Vie et de la Terre, Université de Ouagadougou, 03 BP 7021 Ouagadougou 03, Burkina Faso. * Corresponding author, E-mail: [email protected], Telephone: 00226 78 84 67 65 ABSTRACT Solenostemon rotundifolius or country potato is a tropical multipurpose tuber crop species. It has been one of the staple crops in West Africa but currently, its genetic resources are in a process of disappearing. Characterization of S. rotundifolius genetic variability is recognized as a prior intervention to support a sustainable conservation and use of its genetic resources. For identifying suitable descriptors for S. rotundifolius , a morphoagronomical characterization was carried out on 155 cultivars from Burkina Faso. A total of 50 morphological traits (16 qualitative and 34 quantitative) related to the foliage, the cycle and the tubers were scored. The results showed variability within cultivars for the foliage, the cycle and the potential yield (number and weight of tubers). However, no significant variability was found for tubers size. Cultivars from different geographical origin discriminated for the cycle and the potential yield. Significant correlations were found between the cycle, the foliage and the potential yield. Most of the qualitative morphological traits were shown to be varietal criteria.
    [Show full text]
  • Atoll Research Bulletin No. 503 the Vascular Plants Of
    ATOLL RESEARCH BULLETIN NO. 503 THE VASCULAR PLANTS OF MAJURO ATOLL, REPUBLIC OF THE MARSHALL ISLANDS BY NANCY VANDER VELDE ISSUED BY NATIONAL MUSEUM OF NATURAL HISTORY SMITHSONIAN INSTITUTION WASHINGTON, D.C., U.S.A. AUGUST 2003 Uliga Figure 1. Majuro Atoll THE VASCULAR PLANTS OF MAJURO ATOLL, REPUBLIC OF THE MARSHALL ISLANDS ABSTRACT Majuro Atoll has been a center of activity for the Marshall Islands since 1944 and is now the major population center and port of entry for the country. Previous to the accompanying study, no thorough documentation has been made of the vascular plants of Majuro Atoll. There were only reports that were either part of much larger discussions on the entire Micronesian region or the Marshall Islands as a whole, and were of a very limited scope. Previous reports by Fosberg, Sachet & Oliver (1979, 1982, 1987) presented only 115 vascular plants on Majuro Atoll. In this study, 563 vascular plants have been recorded on Majuro. INTRODUCTION The accompanying report presents a complete flora of Majuro Atoll, which has never been done before. It includes a listing of all species, notation as to origin (i.e. indigenous, aboriginal introduction, recent introduction), as well as the original range of each. The major synonyms are also listed. For almost all, English common names are presented. Marshallese names are given, where these were found, and spelled according to the current spelling system, aside from limitations in diacritic markings. A brief notation of location is given for many of the species. The entire list of 563 plants is provided to give the people a means of gaining a better understanding of the nature of the plants of Majuro Atoll.
    [Show full text]
  • Vegetation Survey of Mount Gorongosa
    VEGETATION SURVEY OF MOUNT GORONGOSA Tom Müller, Anthony Mapaura, Bart Wursten, Christopher Chapano, Petra Ballings & Robin Wild 2008 (published 2012) Occasional Publications in Biodiversity No. 23 VEGETATION SURVEY OF MOUNT GORONGOSA Tom Müller, Anthony Mapaura, Bart Wursten, Christopher Chapano, Petra Ballings & Robin Wild 2008 (published 2012) Occasional Publications in Biodiversity No. 23 Biodiversity Foundation for Africa P.O. Box FM730, Famona, Bulawayo, Zimbabwe Vegetation Survey of Mt Gorongosa, page 2 SUMMARY Mount Gorongosa is a large inselberg almost 700 sq. km in extent in central Mozambique. With a vertical relief of between 900 and 1400 m above the surrounding plain, the highest point is at 1863 m. The mountain consists of a Lower Zone (mainly below 1100 m altitude) containing settlements and over which the natural vegetation cover has been strongly modified by people, and an Upper Zone in which much of the natural vegetation is still well preserved. Both zones are very important to the hydrology of surrounding areas. Immediately adjacent to the mountain lies Gorongosa National Park, one of Mozambique's main conservation areas. A key issue in recent years has been whether and how to incorporate the upper parts of Mount Gorongosa above 700 m altitude into the existing National Park, which is primarily lowland. [These areas were eventually incorporated into the National Park in 2010.] In recent years the unique biodiversity and scenic beauty of Mount Gorongosa have come under severe threat from the destruction of natural vegetation. This is particularly acute as regards moist evergreen forest, the loss of which has accelerated to alarming proportions.
    [Show full text]